Zygopetalum maxillare

• Interesting Facts
• Common Names
• Description
• Taxonomy
• Similar Species
• More Info
• Further Reading
• Notes

Interesting Facts

Common Names

Description

Family Orchidaceae

Herbs or rarely vines , perennial , rarely annual , strongly mycotrophic, epiphytic, terrestrial , lithophytic, or rarely aquatic or subterranean , usually green and photosynthetic, some without chlorophyll and saprophytic . Roots subterranean or aerial , tuberoid or stolonoid, usually with spongy , multilayered velamen. Stems erect or pendent or modified into creeping rhizomes, simple or sympodially or monopodially branched, delicate to stout, or thickened as corms or pseudobulbs , or greatly reduced, sometimes proliferous (especially diverse in sympodial orchids) . Leaves solitary, several, or reduced to scales , basal or cauline, alternate, distichous, or sometimes opposite or whorled , either convolute or duplicate , simple, sessile or petiolate ; stipules absent; blade articulate or not, plicate or conduplicate , cylindric , triangular, or laterally flattened, margins entire. Inflorescences terminal or lateral , racemes , spikes, panicles, or rarely cymose , erect or variously pendent, 1 many-flowered, lax or dense, flowering successively or simultaneously. Flowers bisexual [rarely unisexual ], epigynous , resupinate or not, pedicellate or sessile, 3-merous, usually bilaterally symmetric [rarely nearly radially symmetric], with abscission layer between pedicel and peduncle, rarely between ovary and perianth or ovary and pedicel; perianth of 6 tepals in 2 whorls, all petaloid or sepals sometimes greener and more foliaceous in texture ; sepals alike or not, lateral sepals often connate (forming synsepal), or all 3 sepals variously connate and/or adnate or distinct and/or free ; petals 3, median petal modified as lip, commonly larger or differing in form and color, lateral petals commonly but not always similar to sepals; nectaries of various sorts; extrafloral nectaries sometimes present on pedicels, bracts, or leaf sheaths ; stamens usually 1 2( 3, if 3 the 3d modified into sterile staminode), all on side opposite lip, fully or partially adnate to style, forming column; pollen grains in monads or tetrads , usually in 2 8 pollinia, sometimes subdivided into small packets, rarely granular , sometimes pollinia with caudicles and/or stipes; gynoecium 3-carpellate, connate, forming compound , inferior, 1- or 3-locular ovary; style variously adnate to filaments ; stigmas usually 3-lobed, concave to convex , part of median stigma lobe modified into rostellum , often separating anther from fertile portions of stigma, commonly preventing or in some cases facilitating self-pollination ; ovules numerous , anatropous , minute. Fruits capsules, opening (dehiscing) by longitudinal slits, rarely fleshy and indehiscent berries . Seeds numerous (millions in some species), minute; endosperm absent.

Genera ca. 800, species 22,000 35,000 (701 genera, 208 species in the flora ; 1 genus, 6 species introduced) : worldwide except Antarctica, most diverse in tropical forests .

The overall count for orchid genera in the flora includes Spathoglottis plicata Blume, which was recently reported from Palm Beach County, Florida. The plants , known locally since 1982, are apparently widely naturalized in old shellpits. The number of species in the flora includes one newly recognized species in Habenaria that is morphologically described, but not fully treated here. Orchidaceae are by far the largest and most diverse monocot family and rank among the largest families of flowering plants. An accurate account of the number of genera and species has eluded orchid scientists, and species counts published in the last 20 years range from 15,000 to 35,000. New species are continually being described. In addition, numerous natural and artificial hybrids exist.

Although orchids are important in horticulture , most of the plants traded in the national and international market belong to a small number of species and their hybrids in only a few genera; the majority of orchids are not commonly cultivated. Few orchids are economically important outside the horticultural trade: the fruits of several species of Vanilla are the source of the spice vanilla, and the dry roots of some species of Dactylorhiza, Eulophia, and Orchis are made into salep, a flour consumed in northern Africa, the Middle East (especially Turkey), and Asia. Some species are locally used for medicinal purposes; the mucilage from pseudobulbs of several species is sometimes used as glue; and in the Far East the stems of some species of Dendrobium are split into strips used to weave handicrafts. A few orchids have been found to cause contact dermatitis (e.g. , Cypripedium reginae) .

Orchids range vegetatively from Lilliputian plants a few millimeters long (Bulbophyllum Thouars and Platystele Schlechter) to gigantic clusters weighing several hundred kilograms (Grammatophyllum Blume) to some as much as 13.4 meters in height (Sobralia altissima D. E. Bennett & Christenson, a recently described species from Peru) . Likewise, flowers vary in size from less than 1 mm and barely visible to the naked eye (Platystele Garay), to 15 20 cm diameter (some Paphiopedilum Pfitzer, Phragmipedium Rolfe, and Cattleya Lindley spp. ), and ultimately to 76 cm [Phragmipedium caudatum (Lindley) Rolfe]. Weight can vary from a fraction of a gram (many Pleurothallus R. Brown spp.) to nearly 100 grams (Coryanthes Hooker spp.) . Their fragrances vary from delightful (Cattleya Lindley) to repulsive and unbearable (in some species of Bulbophyllum Thouars) . The plants colonize habitats ranging from some of the driest and hottest places on earth to the wettest and coolest, literally occurring from polar regions to the equator. Within the monocots, the most important diagnostic features of Orchidaceae are reduction of adaxial stamens, fusion of the remaining stamens to the gynoecium forming the column, aggregation of pollen into compact pollinia (present elsewhere only in the dicots , in Asclepiadaceae), differentiation of the median petal into the lip, a sometimes complex organ, and the exceedingly small size of the seed, which lacks endosperm. Among other distinguishing characteristics: pollen in the pollinia is usually not available as a nutrient-source (Cleistes Richard ex Lindley being a notable exception), and the often complex interaction with pollinators culminates in the phenomenon of pseudocopulation in several genera (e.g., Ophrys Linnaeus, Caladenia R. Brown sect. Calonema, Drakaea Lindley) . In the latter process , the flower mimics the appearance , the smell, and often the movements of a female wasp, attracting a male of a suitable species that tries to copulate with the flower. It usually only succeeds in becoming attached to a pollinium , which will then be transferred if the male tries to copulate with another flower.

Roots of orchids may be covered with velamen, spongy layers derived from the epidermis ; fleshy thickenings of roots are tuberoids (tubers being restricted to stems) . Stems may be swollen or thickened, underground corms or aerial pseudobulbs. Flowers are often resupinate: the lip (modified median petal) is  lowermost,  usually as a result of the pedicel being twisted or bent in its development by 180°. Pedicellate ovary, usually used in reference to length , refers to the combined pedicel and ovary. Flowers are not always borne on pedicels; when they are, it is sometimes difficult to distinguish between a slender ovary and the pedicel. Consequently, because of their slender ovaries, flowers of a  racemose spike  appear to be pedicellate even though they are sessile, while a  spicate raceme  has pedicels so short that they appear to be absent. Orchid flowers often have a modified median sepal, the dorsal sepal. Sepals coalescing at their tips form a synsepal. The middle portion of the upper (adaxial) face of the lip is the disc: it may be a thickened callus and may bear hairs , papillae, or other ornamentation. In orchids the style, stigmas, filaments, and one or more anthers are united to form a column; appendages projecting laterally from the stigma are column wings; the lip may be attached to the protrusion at the base of the column to form a column foot ; lateral sepals that are also attached to the foot form a mentum (chin) . In most orchids the column bears a single anther at its apex; the clinandrium is the cavity within which the anther is borne or embedded . Pollen is borne in discrete masses (pollinia) . Genera with mealy (sectile) pollinia may have pollinia within the anther tapering into a caudicle (stalk ), which is attached to a sticky viscidium . Those with waxy pollinia have pollinia attached to one or two stipes (of stigmatic origin and formed outside the anther), which in turn are attached to a viscidium. The various aggregations of pollinia, caudicles, stipes, and viscidium form a pollinarium , the pollination unit carried by pollinators. The median stigma lobe may have a slender extension or little beak (rostellum), which aids in gluing the pollinarium to the pollinator.[1]

Taxonomy

Notes

Name Status: Accepted Name .

Last scrutiny: 11-Nov-2003

Similar Species

Members of the genus Zygopetalum

ZipcodeZoo has pages for 152 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:

Z. (Zygopetalum Orchid) · Z. Advance Australia (Zygopetalum Advance Australia Orchid) · Z. Alan Greatwood (Zygopetalum Alan Greatwood Orchid) · Z. Alexis Linder (Zygopetalum Alexis Linder Orchid) · Z. Artemis (Zygopetalum Artemis Orchid) · Z. Artur Elle (Zygopetalum Artur Elle Orchid) · Z. Bayswater (Zygopetalum Bayswater Orchid) · Z. Betty (Zygopetalum Betty Orchid) · Z. Big Country (Zygopetalum Big Country Orchid) · Z. Blackii (Zygopetalum Blackii Orchid) · Z. Black Opal (Zygopetalum Black Opal Orchid) · Z. Black Pearl (Zygopetalum Black Pearl Orchid) · Z. Black Plague (Zygopetalum Black Plague Orchid) · Z. Blanchetown (Zygopetalum Blanchetown Orchid) · Z. Blinky (Zygopetalum Blinky Orchid) · Z. Blue Banks (Zygopetalum Blue Banks Orchid) · Z. Blue Bear (Zygopetalum Blue Bear Orchid) · Z. Blue Blazes (Zygopetalum Blue Blazes Orchid) · Z. Blue Blood (Zygopetalum Blue Blood Orchid) · Z. Blue Eyes (Zygopetalum Blue Eyes Orchid) · Z. Blue Lake (Zygopetalum Blue Lake Orchid) · Z. Bolero (Zygopetalum Bolero Orchid) · Z. Bon Voyage (Zygopetalum Bon Voyage Orchid) · Z. brachypetalum (Zygopetalum Brachypetalum Orchid) · Z. Bright Eyes (Zygopetalum Bright Eyes Orchid) · Z. Bright Imagination (Zygopetalum Bright Imagination Orchid) · Z. Brookside (Zygopetalum Brookside Orchid) · Z. B. G. White (Zygopetalum B. G. White Orchid) · Z. Carolyn Southwood Karges (Zygopetalum Carolyn Southwood Karges Orchid) · Z. Centenary (Zygopetalum Centenary Orchid) · Z. Clarksonii (Zygopetalum Clarksonii Orchid) · Z. Clayi (Zygopetalum Clayi Orchid) · Z. Conway's Choice (Zygopetalum Conway's Choice Orchid) · Z. Cosmo-Amethyst (Zygopetalum Cosmo-Amethyst Orchid) · Z. Crinito-gautieri (Zygopetalum Crinito-Gautieri Orchid) · Z. crinitum (Zygopetalum Crinitum Orchid) · Z. Dark Star (Zygopetalum Dark Star Orchid) · Z. Days Gone By (Zygopetalum Days Gone By Orchid) · Z. Debbie (Zygopetalum Debbie Orchid) · Z. Dinkum Aussie (Zygopetalum Dinkum Aussie Orchid) · Z. Doctor Ben (Zygopetalum Doctor Ben Orchid) · Z. Eden (Zygopetalum Eden Orchid) · Z. Elaine (Zygopetalum Elaine Orchid) · Z. Everspring (Zygopetalum Everspring Orchid) · Z. Freestyle (Zygopetalum Freestyle Orchid) · Z. Freestyle Hawaii (Zygopetalum Freestyle Hawaii Orchid) · Z. Gary Baker (Zygopetalum Gary Baker Orchid) · Z. gautieri (Zygopetalum Gautieri Orchid) · Z. Gidget (Zygopetalum Gidget Orchid) · Z. Glenelg (Zygopetalum Glenelg Orchid) · Z. Gottianum (Zygopetalum Gottianum Orchid) · Z. graminifolium (Zygopetalum Graminifolium Orchid) · Z. Granite Island (Zygopetalum Granite Island Orchid) · Z. Great Eisen (Zygopetalum Great Eisen Orchid) · Z. Hawker (Zygopetalum Hawker Orchid) · Z. Helen-Ku (Zygopetalum Helen-Ku Orchid) · Z. Hove (Zygopetalum Hove Orchid) · Z. Hybrid ignota (Zygopetalum Hybrid Ignota Orchid) · Z. Imagination (Zygopetalum Imagination Orchid) · Z. Impulse (Zygopetalum Impulse Orchid) · Z. Inky Dink (Zygopetalum Inky Dink Orchid) · Z. intermedium (Zygopetalum Intermedium Orchid) · Z. Intuition (Zygopetalum Intuition Orchid) · Z. Iron Baron (Zygopetalum Iron Baron Orchid) · Z. Issy (Zygopetalum Issy Orchid) · Z. James Strauss (Zygopetalum James Strauss Orchid) · Z. Jean Inouye (Zygopetalum Jean Inouye Orchid) · Z. John Banks (Zygopetalum John Banks Orchid) · Z. Jumpin Jack (Zygopetalum Jumpin Jack Orchid) · Z. June Bloodworth (Zygopetalum June Bloodworth Orchid) · Z. Kiwi (Zygopetalum Kiwi Orchid) · Z. Kiwi Auckland (Zygopetalum Kiwi Auckland Orchid) · Z. Kiwi Bayswater (Zygopetalum Kiwi Bayswater Orchid) · Z. Kiwi Black (Zygopetalum Kiwi Black Orchid) · Z. Kiwi Blue (Zygopetalum Kiwi Blue Orchid) · Z. Kiwi Choice (Zygopetalum Kiwi Choice Orchid) · Z. Kiwi Dazzler (Zygopetalum Kiwi Dazzler Orchid) · Z. Kiwi Dusk (Zygopetalum Kiwi Dusk Orchid) · Z. Kiwi Fifty One (Zygopetalum Kiwi Fifty One Orchid) · Z. Kiwi Geyser (Zygopetalum Kiwi Geyser Orchid) · Z. Kiwi Hit (Zygopetalum Kiwi Hit Orchid) · Z. Kiwi Klassic (Zygopetalum Kiwi Klassic Orchid) · Z. Kiwi Korker (Zygopetalum Kiwi Korker Orchid) · Z. Kiwi Land (Zygopetalum Kiwi Land Orchid) · Z. Kiwi Magic (Zygopetalum Kiwi Magic Orchid) · Z. Kiwi Mud Pool (Zygopetalum Kiwi Mud Pool Orchid) · Z. Kiwi Redvale (Zygopetalum Kiwi Redvale Orchid) · Z. Kosciusko (Zygopetalum Kosciusko Orchid) · Z. Leopardinum (Zygopetalum Leopardinum Orchid) · Z. Leopard Prince (Zygopetalum Leopard Prince Orchid) · Z. Lois (Zygopetalum Lois Orchid) · Z. mackayi (Caper Bean) · Z. maculatum (Zygopetalum Maculatum Orchid) · Z. Margaret (Zygopetalum Margaret Orchid) · Z. Marilyn Southwood Stemaly (Zygopetalum Marilyn Southwood Stemaly Orchid) · Z. Marino Rocks (Zygopetalum Marino Rocks Orchid) · Z. maxillare (Zygopetalum Maxillare Orchid) · Z. Memoria Bob Paice (Zygopetalum Memoria Bob Paice Orchid) · Z. Memoria Captain Alan Thoma (Zygopetalum Memoria Captain Alan Thoma Orchid) · Z. Memoria Grace Drewry (Zygopetalum Memoria Grace Drewry Orchid)

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Further Reading

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• Botanical Museum leaflets, Harvard University. 21 1966 Cambridge, Mass.: Botanical Museum, Harvard University, 1932- url p. 186, p. 187.
• Bulletin of miscellaneous information /Royal Botanic Gardens, Kew. 1913 London: H.M. Stationery Office, 1900-1941. url p. 78.
• Cool orchids, and how to grow them: with a descriptive list of all the best species in cultivation / by F.W. Burbidge London: R. Hardwicke, 1874 url p. 132.
• Cyclopedia of American horticulture, comprising suggestions for cultivation of horticultural plants, descriptions of the species of fruits, vegetables, flowers and ornamental plants sold in the United States and Canada, together with geo by L.H. Bailey assisted by Wilhelm Miller, PH. D., associate editor, and many expert cultivators and botanists. Illustrated with nearly three thousand engravings and one hundred and forty-five full-pa New York, Doubleday, Page & Company, 1906. url p. 2015.
• Garden and forest; a journal of horticulture, landscape art and forestry. New York: The Garden and forest publishing co., 1888-97. url p. 510.
• Handbook of flower pollination based upon Hermann Müller's work 'The fertilisation of flowers by insects'; tr. by J.R. Ainsworth Davis. Oxford, Clarendon Press, 1906. url p. 421.
• Hortus Veitchii: a history of the rise and progress of the nurseries of Messrs. James Veitch and sons, together with an account of the botanical collectors and hybridists employed by them and a list of the most remark by James H. Veitch; with fifty illustrations. London: J. Veitch & sons, 1906. url p. 217, p. 218, p. 474, p. 476.
• Nicholson, G. The illustrated dictionary of gardening: a practical and scientific encyclopædia of horticulture for gardeners and botanists /edited by George Nicholson; assisted by J.W.H. Trail. .. and J. Garrett. .. 9 1884 London: L.U. Gill, [1884]-88. url p. 430.
• Orchid album: comprising coloured figures and descriptions of new, rare and beautiful orchidaceous plants /Conducted by Robert Warner and Benjamin Samuel Williams; the botanical descriptions by Thomas Moore; the coloured figures by John Nugent Fitch. 1883 London: B. S. Williams, 1882-97. url , .
• Orchids / by James O'Brien. London: T. C. & E. C. Jack, [1911]. url p. 107, p. 107.
• Orchids for everyone, London, E.P. Dutton, 1910. url .
• Orchids for everyone. London: Dent, 1910. url p. 324.
• Orchids, New York, Frederick A. Stokes co.[1911] url .
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• Orchids: their culture and management. London, C. Scribner's sons, 1903. url , .
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• Orchids; being the report on the Orchid Conference held at South Kensington. .. London, Macmillan & Co., 1886. url p. 86.
• Orchids; their culture and management. With descriptions of all the kinds in general cultivation / By W. Watson assisted by W. Bean. London, L. U. Gill, 1890. url p. 513, p. 537.
• Proceedings of the South London Entomological & Natural History Society. London [Eng]: South London Entomological and Natural history Society, [1897-1933] url p. 64.
• The American florist: a weekly journal for the trade. Chicago: American Florist Company, [1885-1931] url p. 1048, p. 4.
• The Book of gardening; a handbook of horticulture. London, C. Scribner's Sons, 1900. url .
• The Floral world and garden guide. London, Groombridge and Sons, 1858-1880. url p. 353.
• The Garden: an illustrated weekly journal of gardening in all its branches. London: [s.n., url , p. 106, p. 119, p. 169, p. 193, p. 195, p. 204, p. 214, p. 220, p. 272, p. 283, p. 329, p. 329, p. 336, p. 338, p. 339, p. 351, p. 36, p. 365, p. 366, p. 366, p. 380, p. 406, p. 433, p. 436, p. 441, p. 464, p. 465, p. 480, p. 498, p. 50, p. 520, p. 53, p. 535, p. 54, p. 55, p. 554, p. 570, p. 609, p. 612.
• The Gardener. Edinburgh;W. Blackwood and sons, 1867-1882. url p. 535.
• The Gardeners' chronicle and agricultural gazette. London: published for the proprietors, 1844-1873. url p. 1297, p. 1361, p. 1477, p. 1562.
• The Gardeners' chronicle: a weekly illustrated journal of horticulture and allied subjects. London: [Gardeners Chronicle], 1874-1955. url , p. 100, p. 130, p. 147, p. 158, p. 160, p. 162, p. 165, p. 179, p. 18, p. 193, p. 200, p. 214, p. 23, p. 230, p. 235, p. 246, p. 263, p. 267, p. 279, p. 290, p. 291, p. 30, p. 307, p. 308, p. 310, p. 335, p. 347, p. 354, p. 38, p. 381, p. 388, p. 389, p. 39, p. 395, p. 395, p. 401, p. 420, p. 428, p. 432, p. 443, p. 458, p. 461, p. 463, p. 464, p. 485, p. 49, p. 495, p. 514, p. 519, p. 519, p. 524, p. 531, p. 532, p. 542, p. 542, p. 550, p. 564, p. 571, p. 578, p. 582, p. 582, p. 585, p. 591, p. 630, p. 632, p. 68, p. 684, p. 690, p. 7, p. 72, p. 742, p. 749, p. 750, p. 754, p. 79, p. 814, p. 818.
• The Journal of horticulture, cottage gardener and country gentlemen. London: George W. Johnson and Robert Hogg, 1861-1877. url , , p. 170, p. 207, p. 31, p. 348, p. 349, p. 461, p. 96.
• The Orchid review. [Kingsteignton, Eng., etc.: Orchid Review ltd.] url p. 214, p. 298, p. 300, p. 307, p. 311, p. 311, p. 315, p. 319, p. 319, p. 322, p. 352, p. 364, p. 374, p. 64, p. 7.
• The Woodlands orchids described and illus. with stories of orchid collecting, by Frederick Boyle. Coloured plates by F.L. MacFarlane. London: Macmillan, 1901. url p. 131.
• The culture of greenhouse orchids: old system and new / by Frederick Boyle; supervised by Joseph Godseff; with three plates in colour and fifty illustrations from photographs by Col. F.C. Taylor. London: Chapman & Hall, 1902. url p. 87.
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• The standard cyclopedia of horticulture; a discussion, for the amateur, and the professional and commercial grower, of the kinds, characteristics and methods of cultivation of the species of plants grown in the regions of the United States a Illustrated with colored plates, four thousand engravings in the text, and ninety-six full-page cuts. New York, Macmillan, 1919 [c1914] url p. 3550.
• Transactions of the Massachusetts Horticultural Society. [S.l.: s.n.], 1843-1920. url p. 99.
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• Bentley, S. L. Native Orchids of the Southern Appalachian Mountains. Chapel Hill.
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Notes

Contributors

• Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 30, 2012.
• The Royal Horticultural Society Horticultural Database, available here.

Data Sources

Accessed through GBIF Data Portal March 15, 2008:

• Missouri Botanical Garden

Identifiers

• Biodiversity Heritage Library NamebankID: 5981814
• Catalogue of Life Accepted Name Code: Kew-216093
• Global Biodiversity Information Facility Taxonkey: 15721362
• Globally Unique Identifier: urn:lsid:ipni.org:names:661747-1
• Zipcode Zoo Species Identifier: 1093793

Footnotes

1. Gustavo A. Romero-González, Germán Carnevali Fernández-Concha, Robert L. Dressler, Lawrence K. Magrath & George W. Argus "Orchidaceae". in Flora of North America Vol. 26 Page 15, 16, 17, 26, 27, 490, 491, 617. Oxford University Press. Online at EFloras.org. [back]

©2004-2012 David Stang. All rights reserved.