Interesting Facts
Description
Family Juglandaceae
Trees
or rarely shrubs
, deciduous, semievergreen, or evergreen
, monoecious or rarely dioecious; bark
tight (or exfoliating) . Branchlets
with solid or chambered
pith
. Terminal
buds subglobose or ovoid
to oblong
, naked or with scales
. Stipules absent. Leaves alternate (or opposite), odd- or even-pinnate, sometimes trifoliolate
, rarely simple
; leaflets
with glandular
, peltate scales, often resinous
and aromatic
, particularly conspicuous
abaxially on young leaves and twigs
, margin
serrate or rarely entire. Inflorescences pendulous or sometimes erect
, lateral
or terminal, on reduced shoots
arising on branchlets of previous year (old growth) or on current
year’s growth (new growth), of several types: androgynous panicle with male, lateral spikes and female, central spike; androgynous panicle with male, mainly lateral spikes and female, central spike male at apex; cluster
of male spikes and solitary female spike; or solitary male and female spikes. Flowers unisexual
, anemophilous
, rarely entomophilous. Male flowers subtended by an entire or 3-lobed bract; bracteoles 2 or absent; sepals 0-4, adnate
to receptacle when present; stamens 3-40(-100), inserted
on receptacle; filaments
short to nearly absent, free
or united
at base
; anthers
glabrous
or pubescent
, 2-loculed, dehiscing longitudinally. Female flowers with an entire or 3-lobed bracts; bracteoles 2 or 3 (or absent) ; sepals 0-4, adnate to ovary, free at apex; gynoecium of 2 carpels united into an inferior ovary
, 1-loculed, but at base 2-4(-8) -loculed; style 1, short or elongate
, rarely absent; stigmas 2, carinal or commissural
, sometimes 4-lobed, plumose
or fleshy
; ovule 1, orthotropous
. Fruiting spike elongate, and pendulous or short and erect, rarely conelike. Fruit a drupelike nut, 2-4(-8) -chambered at base, with a dehiscent
or indehiscent husk
, or a 2- or 3-winged or disc-winged nutlet
. Seed solitary, without endosperm. Cotyledons 4-lobed, much contorted. Germination hypogeal or epigeal. 2n = (28), 32, (64) .
Nine genera and 60 or more species: mostly in temperate
and subtropical
regions of the N hemisphere; seven genera (one endemic) and 20 species (seven endemic, one introduced
) in China.[1]
Genus Juglans
Shrubs
or trees
, 3-50 m.
Bark
light to dark gray or gray-brown, smooth
or split into ridges
or plates
. Twigs
purplish brown, terete
, stout, sparsely to densely covered with glands
and capitate-glandular hairs
, sometimes also with scales
or fasciculate hairs, early in season
with multiradiate
hairs; leaf scars
triangular or 3-lobed, large; pith
chambered
. Bud scales valvate
, densely hirsute
. Leaves usually odd-, sometimes even-pinnate; petiole
and rachis with indument
as twigs. Leaflets
5-25, sessile or subsessile
, often aromatic
, uniform
in size or median
leaflets largest, (2.5-) 4.3-15(-17.5) × 0.8-6.5 cm; surfaces usually with nonglandular hairs (simple
and/or fasciculate), glandular
hairs, sessile glands, and/or scales, sometimes glabrous
. Staminate
catkins solitary from 2d-year twigs, sessile; stamens 7-50 per flower, glabrous or pilose
. Pistillate
flowers solitary or in terminal
racemes
. Fruits nuts enclosed in husks
, not compressed
; husks thick, indehiscent; nuts tan, neither compressed nor angled
, grooved
, ridged
, rugulose
, or smooth; shells
thick. Seeds sweet. x
= 16.
Species 21: North America, Mexico, West Indies, Central America, South America, Eurasia
.
Juglans is a very important source of edible nuts, dyes, and wood
for cabinet work, furniture, and construction. Juglans regia Linnaeus, the walnut of commerce, is widely cultivated in California; it is easily distinguished from native species
by its leaves with 5-11 broad, entire leaflets and nuts with thin rugulose shells, not grooved or ridged. Because of its sensitivity to native
pathogens
, J. regia is usually grown as stem-grafts on roots
of native or hybrid walnuts (see discussion under J. hindsii). Occasional seedlings of J. regia have been reported from the vicinity of cultivated plants
, but these seldom, if ever, live to maturity.
The growth form
, bark, and fruit are important taxonomically in Juglans, but these usually are not available on herbarium
specimens. As with many woody plants
, the first one or two leaves of the season (i.e.
, the lowermost leaves on the twig
) are sometimes atypical
in structure, having broader, blunter leaflets. The fasciculate hairs on the veins normally have more rays than those on the blade
. In addition to the types of hairs described below, small multiradiate hairs are found on the immature
twigs, petioles, rachises, and midribs
. They are usually gone when the leaf is fully expanded, but they may persist for a short time afterwards.[2]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Juglandanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Fagales
(
)
- Engler, 1892
- Family:
Juglandaceae
(
)
- DC. ex Perleb, 1818, nom. cons.
- walnuts
- Subfamily:
Juglandoideae
(
)
- Tribe:
Juglandeae
(
)
- Genus:
Juglans
(
)
- C. Linnaeus, 1753
- Walnut
- Specific epithet:
nigra
- Marshall
- Variety:
oblonga
- Botanical name: - Juglans nigra var. oblonga Marshall
- Variety:
oblonga
- Specific epithet:
nigra
- Marshall
- Genus:
Juglans
(
- Tribe:
Juglandeae
(
- Subfamily:
Juglandoideae
(
- Family:
Juglandaceae
(
- Order:
Fagales
(
- Superorder:
Juglandanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Juglans nigra L. f. oblonga (Marshall) Fernald
Notes
Publishing author : Marshall .
Similar Species
Members of the genus Juglans
ZipcodeZoo has pages for 82 species, subspecies, varieties, forms, and cultivars in this genus:
J. ailanthifolia (Japanese Walnut) · J. ailanthifolia var. cordiformis (Heartnut) · J. ailantifolia (Japanese Walnut) · J. australis (Nogal Criollo) · J. bixbyi (Bixby Walnut) · J. boliviana (Bolivian Walnut) · J. californica (Northern California Black Walnut) · J. californica S.Watson var. californica S.Watson (Southern California Black Walnut) · J. californica var. californica (California Walnut) · J. cinerea (Butternut) · J. cinerea 'Bountiful' (Butternut) · J. cinerea 'Loumis' (Butternut) · J. cordiformis 'Campbell Cw1' (Heartnut) · J. cordiformis 'Campbell Cw3' (Heartnut) · J. cordiformis 'Campbell Cww' (Heartnut) · J. cordiformis 'Frank' (Heartnut) · J. cordiformis 'Imshu' (Heartnut) · J. cordiformis 'Locket' (Heartnut) · J. cordiformis 'Marvel' (Heartnut) · J. cordiformis 'Mitchell Hybrid' (Heartnut) · J. cordiformis 'Rhodes' (Heartnut) · J. cordiformis 'Simcoe' (Heartnut) · J. cordiformis 'Stealth' (Heartnut) · J. cordiformis 'Wright' (Heartnut) · J. hindsii (Northern California Black Walnut) · J. illinoinensis (Pecan) · J. intermedia (Intermediate Walnut) · J. jamaicensis (Walnut) · J. major (Arizona Black Walnut) · J. mandshurica (Manchurian Walnut) · J. microcarpa (Little Walnut) · J. microcarpa var. microcarpa (Little Walnut) · J. microcarpa var. stewartii (Stewarts Little Walnut) · J. microcarpa var. stewartii (I.M.Johnst.) W.Manning (Stewart's Little Walnut) · J. microcarpa subsp. major (Little Walnut) · J. neotropica (Andean Walnut) · J. nigra (Black Walnut) · J. nigra 'Beineke 11' (Black Walnut 'beineke 11') · J. nigra 'Black Gem' (Black Walnut) · J. nigra 'Daniels' (Black Walnut) · J. nigra 'Emma Kay' (Black Walnut) · J. nigra 'Football 2' (Black Walnut) · J. nigra 'Krause' (Black Walnut) · J. nigra 'Kwik Krop' (Black Walnut) · J. nigra 'Laciniata' (Black Walnut) · J. nigra 'Lamb's Curly' (Black Walnut) · J. nigra 'Ridgeway' (Black Walnut) · J. nigra 'Rowher' (Black Walnut) · J. nigra 'Sauber 1' (Black Walnut) · J. nigra 'Schrieber' (Black Walnut) · J. nigra 'Sparrow' (Black Walnut) · J. nigra 'Surprise' (Black Walnut) · J. nigra 'Thomas Black' (Black Walnut) · J. nigra 'Thomas Myers' (Black Walnut) · J. regia (Carpathian Walnut) · J. regia var. Carpathian (Carpathian Walnut) · J. regia 'Allegheny' (Carpathian Walnut) · J. regia 'Bedco 1' (Carpathian Walnut) · J. regia 'Broadview' (Dwarf English Walnut) · J. regia 'Cascade' (Carpathian Walnut) · J. regia 'Champion' (Carpathian Walnut) · J. regia 'China-B' (Carpathian Walnut) · J. regia 'Coble #2' (Carpathian Walnut) · J. regia 'Colby' (Carpathian Walnut) · J. regia 'Greenhaven' (Carpathian Walnut) · J. regia 'Hansen' (Carpathian Walnut) · J. regia 'Kaiser' (Carpathian Walnut) · J. regia 'Ky Giant' (Carpathian Walnut) · J. regia 'Lake' (Carpathian Walnut) · J. regia 'Looking Glass' (Carpathian Walnut) · J. regia 'Mckinster' (Carpathian Walnut) · J. regia 'Northern Prize' (Carpathian Walnut) · J. regia 'Perry' (Carpathian Walnut) · J. regia 'Purpurea' (Carpathian Walnut) · J. regia 'Reda' (Carpathian Walnut) · J. regia 'S-1' (Carpathian Walnut) · J. regia 'Shiawassee' (Carpathian Walnut) · J. regia 'Somers' (Carpathian Walnut) · J. regia 'Utah Giant' (Carpathian Walnut) · J. × bixbyi (Bixby Walnut) · J. x intermedia (Intermediate Walnut) · J. 'Royal' (Walnut)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Kuang Ko-zen & Lu An-ming. 1979. Juglandaceae. In: Kuang Ko-zen & Li Pei-chun, eds., Fl. Reipubl. Popularis Sin. 21: 6-44.
- McGranahan, G. H. and P. B. Catlin. 1987. Juglans rootstocks. In: R. C. Rom and R. F. Carlson, eds. 1987. Rootstocks for Fruit Crops. New York. Pp. 411-450.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Identifiers
- Zipcode Zoo Species Identifier: 4597406
Footnotes
- Anmin Lu, Donald E. Stone & L. J. Grauke "Juglandaceae". in Flora of China Vol. 4 Page 277. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Alan T. Whittemore & Donald E. Stone "Juglans". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
