Overview
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Interesting Facts
Common Names
Click on the language to view common names.
Common Names in English:
Walnut, West Indian Walnut
Common Names in Spanish:
Palo De Nuez, Nogal
Description
Family Juglandaceae
Trees
or rarely shrubs
, deciduous, semievergreen, or evergreen
, monoecious or rarely dioecious; bark
tight (or exfoliating) . Branchlets
with solid or chambered
pith
. Terminal
buds subglobose or ovoid
to oblong
, naked or with scales
. Stipules absent. Leaves alternate (or opposite), odd- or even-pinnate, sometimes trifoliolate
, rarely simple
; leaflets
with glandular
, peltate scales, often resinous
and aromatic
, particularly conspicuous
abaxially on young leaves and twigs
, margin
serrate or rarely entire. Inflorescences pendulous or sometimes erect
, lateral
or terminal, on reduced shoots
arising on branchlets of previous year (old growth) or on current
year’s growth (new growth), of several types: androgynous panicle with male, lateral spikes and female, central spike; androgynous panicle with male, mainly lateral spikes and female, central spike male at apex; cluster
of male spikes and solitary female spike; or solitary male and female spikes. Flowers unisexual
, anemophilous
, rarely entomophilous. Male flowers subtended by an entire or 3-lobed bract; bracteoles 2 or absent; sepals 0-4, adnate
to receptacle when present; stamens 3-40(-100), inserted
on receptacle; filaments
short to nearly absent, free
or united
at base
; anthers
glabrous
or pubescent
, 2-loculed, dehiscing longitudinally. Female flowers with an entire or 3-lobed bracts; bracteoles 2 or 3 (or absent) ; sepals 0-4, adnate to ovary, free at apex; gynoecium of 2 carpels united into an inferior ovary
, 1-loculed, but at base 2-4(-8) -loculed; style 1, short or elongate
, rarely absent; stigmas 2, carinal or commissural
, sometimes 4-lobed, plumose
or fleshy
; ovule 1, orthotropous
. Fruiting spike elongate, and pendulous or short and erect, rarely conelike. Fruit a drupelike nut, 2-4(-8) -chambered at base, with a dehiscent
or indehiscent husk
, or a 2- or 3-winged or disc-winged nutlet
. Seed solitary, without endosperm. Cotyledons 4-lobed, much contorted. Germination hypogeal or epigeal. 2n = (28), 32, (64) .
Nine genera and 60 or more species: mostly in temperate
and subtropical
regions of the N hemisphere; seven genera (one endemic) and 20 species (seven endemic, one introduced
) in China.[1]
Genus Juglans
Shrubs
or trees
, 3-50 m.
Bark
light to dark gray or gray-brown, smooth
or split into ridges
or plates
. Twigs
purplish brown, terete
, stout, sparsely to densely covered with glands
and capitate-glandular hairs
, sometimes also with scales
or fasciculate hairs, early in season
with multiradiate
hairs; leaf scars
triangular or 3-lobed, large; pith
chambered
. Bud scales valvate
, densely hirsute
. Leaves usually odd-, sometimes even-pinnate; petiole
and rachis with indument
as twigs. Leaflets
5-25, sessile or subsessile
, often aromatic
, uniform
in size or median
leaflets largest, (2.5-) 4.3-15(-17.5) × 0.8-6.5 cm; surfaces usually with nonglandular hairs (simple
and/or fasciculate), glandular
hairs, sessile glands, and/or scales, sometimes glabrous
. Staminate
catkins solitary from 2d-year twigs, sessile; stamens 7-50 per flower, glabrous or pilose
. Pistillate
flowers solitary or in terminal
racemes
. Fruits nuts enclosed in husks
, not compressed
; husks thick, indehiscent; nuts tan, neither compressed nor angled
, grooved
, ridged
, rugulose
, or smooth; shells
thick. Seeds sweet. x
= 16.
Species 21: North America, Mexico, West Indies, Central America, South America, Eurasia
.
Juglans is a very important source of edible nuts, dyes, and wood
for cabinet work, furniture, and construction. Juglans regia Linnaeus, the walnut of commerce, is widely cultivated in California; it is easily distinguished from native species
by its leaves with 5-11 broad, entire leaflets and nuts with thin rugulose shells, not grooved or ridged. Because of its sensitivity to native
pathogens
, J. regia is usually grown as stem-grafts on roots
of native or hybrid walnuts (see discussion under J. hindsii). Occasional seedlings of J. regia have been reported from the vicinity of cultivated plants
, but these seldom, if ever, live to maturity.
The growth form
, bark, and fruit are important taxonomically in Juglans, but these usually are not available on herbarium
specimens. As with many woody plants
, the first one or two leaves of the season (i.e.
, the lowermost leaves on the twig
) are sometimes atypical
in structure, having broader, blunter leaflets. The fasciculate hairs on the veins normally have more rays than those on the blade
. In addition to the types of hairs described below, small multiradiate hairs are found on the immature
twigs, petioles, rachises, and midribs
. They are usually gone when the leaf is fully expanded, but they may persist for a short time afterwards.[2]
Physical Description
Habit: Tree
Habitat
Biome: Terrestrial [3].
Ecology: A large tree occurring in wet montane forest .[3].
Biology
Reproduction
Duration: Perennial
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Juglandanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Fagales
(
)
- Engler, 1892
- Family:
Juglandaceae
(
)
- DC. ex Perleb, 1818, nom. cons.
- walnuts
- Subfamily:
Juglandoideae
(
)
- Tribe:
Juglandeae
(
)
- Tribe:
Juglandeae
(
- Subfamily:
Juglandoideae
(
- Family:
Juglandaceae
(
- Order:
Fagales
(
- Superorder:
Juglandanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author
: C.DC. Publication
: Prodr. (DC.) 16(2.1): 138 1864
[late Dec 1864]
Name
Status: Accepted Name
.
Last scrutiny: 15-Mar-2000
Similar Species
Members of the genus Juglans
ZipcodeZoo has pages for 82 species, subspecies, varieties, forms, and cultivars in this genus:
J. ailanthifolia (Japanese Walnut) · J. ailanthifolia var. cordiformis (Heartnut) · J. ailantifolia (Japanese Walnut) · J. australis (Nogal Criollo) · J. bixbyi (Bixby Walnut) · J. boliviana (Bolivian Walnut) · J. californica (Northern California Black Walnut) · J. californica S.Watson var. californica S.Watson (Southern California Black Walnut) · J. californica var. californica (California Walnut) · J. cinerea (Butternut) · J. cinerea 'Bountiful' (Butternut) · J. cinerea 'Loumis' (Butternut) · J. cordiformis 'Campbell Cw1' (Heartnut) · J. cordiformis 'Campbell Cw3' (Heartnut) · J. cordiformis 'Campbell Cww' (Heartnut) · J. cordiformis 'Frank' (Heartnut) · J. cordiformis 'Imshu' (Heartnut) · J. cordiformis 'Locket' (Heartnut) · J. cordiformis 'Marvel' (Heartnut) · J. cordiformis 'Mitchell Hybrid' (Heartnut) · J. cordiformis 'Rhodes' (Heartnut) · J. cordiformis 'Simcoe' (Heartnut) · J. cordiformis 'Stealth' (Heartnut) · J. cordiformis 'Wright' (Heartnut) · J. hindsii (Northern California Black Walnut) · J. illinoinensis (Pecan) · J. intermedia (Intermediate Walnut) · J. jamaicensis (Walnut) · J. major (Arizona Black Walnut) · J. mandshurica (Manchurian Walnut) · J. microcarpa (Little Walnut) · J. microcarpa var. microcarpa (Little Walnut) · J. microcarpa var. stewartii (Stewarts Little Walnut) · J. microcarpa var. stewartii (I.M.Johnst.) W.Manning (Stewart's Little Walnut) · J. microcarpa subsp. major (Little Walnut) · J. neotropica (Andean Walnut) · J. nigra (Black Walnut) · J. nigra 'Beineke 11' (Black Walnut 'beineke 11') · J. nigra 'Black Gem' (Black Walnut) · J. nigra 'Daniels' (Black Walnut) · J. nigra 'Emma Kay' (Black Walnut) · J. nigra 'Football 2' (Black Walnut) · J. nigra 'Krause' (Black Walnut) · J. nigra 'Kwik Krop' (Black Walnut) · J. nigra 'Laciniata' (Black Walnut) · J. nigra 'Lamb's Curly' (Black Walnut) · J. nigra 'Ridgeway' (Black Walnut) · J. nigra 'Rowher' (Black Walnut) · J. nigra 'Sauber 1' (Black Walnut) · J. nigra 'Schrieber' (Black Walnut) · J. nigra 'Sparrow' (Black Walnut) · J. nigra 'Surprise' (Black Walnut) · J. nigra 'Thomas Black' (Black Walnut) · J. nigra 'Thomas Myers' (Black Walnut) · J. regia (Carpathian Walnut) · J. regia var. Carpathian (Carpathian Walnut) · J. regia 'Allegheny' (Carpathian Walnut) · J. regia 'Bedco 1' (Carpathian Walnut) · J. regia 'Broadview' (Dwarf English Walnut) · J. regia 'Cascade' (Carpathian Walnut) · J. regia 'Champion' (Carpathian Walnut) · J. regia 'China-B' (Carpathian Walnut) · J. regia 'Coble #2' (Carpathian Walnut) · J. regia 'Colby' (Carpathian Walnut) · J. regia 'Greenhaven' (Carpathian Walnut) · J. regia 'Hansen' (Carpathian Walnut) · J. regia 'Kaiser' (Carpathian Walnut) · J. regia 'Ky Giant' (Carpathian Walnut) · J. regia 'Lake' (Carpathian Walnut) · J. regia 'Looking Glass' (Carpathian Walnut) · J. regia 'Mckinster' (Carpathian Walnut) · J. regia 'Northern Prize' (Carpathian Walnut) · J. regia 'Perry' (Carpathian Walnut) · J. regia 'Purpurea' (Carpathian Walnut) · J. regia 'Reda' (Carpathian Walnut) · J. regia 'S-1' (Carpathian Walnut) · J. regia 'Shiawassee' (Carpathian Walnut) · J. regia 'Somers' (Carpathian Walnut) · J. regia 'Utah Giant' (Carpathian Walnut) · J. × bixbyi (Bixby Walnut) · J. x intermedia (Intermediate Walnut) · J. 'Royal' (Walnut)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 307.
- Bulletin of miscellaneous information /Royal Botanic Gardens, Kew. 1906 London: H.M. Stationery Office, 1900-1941. url p. 150.
- Bulletin of the U.S. Department of Agriculture. [Washington, D.C.?]: Supt. of Docs., G.P.O., 1913-1923. url p. 61, p. 66.
- Contributions from the United States National Herbarium 51 2005 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 17.
- Flora of Jamaica, containing descriptions of the flowering plants known from the island, by William Fawcett and Alfred Barton Rendle. London.Printed by order of the Trustees of the British Museum, 1910- url p. 34, p. 34.
- Forest of Port Rico; Washington, Govt. print. off., 1916. url p. 61.
- Journal of the New York Botanical Garden. 27 1926 Lancaster, Pa.: Published for the Garden by the New Era Printing Co., 1900- url p. 19, p. 225.
- Land use and landscape change along the Dominican-Haitian borderlands / by Ernest Charles Palmer. 1976. url p. 33.
- Science progress. London: Scientific Press, [1894-1898] url p. 29.
- The Bradley bibliography; a guide to the literature of the woody plants of the world published before the beginning of the twentieth century; Cambridge, Riverside Press, 1911-18. url p. 113.
- The World List of Threatened Trees WCMC, IUCN url p. 299.
- Kuang Ko-zen & Lu An-ming. 1979. Juglandaceae. In: Kuang Ko-zen & Li Pei-chun, eds., Fl. Reipubl. Popularis Sin. 21: 6-44.
- McGranahan, G. H. and P. B. Catlin. 1987. Juglans rootstocks. In: R. C. Rom and R. F. Carlson, eds. 1987. Rootstocks for Fruit Crops. New York. Pp. 411-450.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
- IUCN 2012. IUCN Red List of Threatened Species. Version 2011.2. . Downloaded on January 28, 2012.
- World Conservation Monitoring Centre 1998. Juglans jamaicensis. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloadedon 01February2012.
Identifiers
- Biodiversity Heritage Library NamebankID: 2645918
- Catalogue of Life Accepted Name Code: ITS-19255
- Global Biodiversity Information Facility Taxonkey: 13729880
- Globally Unique Identifier: urn:lsid:ipni.org:names:442376-1
- GRIN Nomen Number: 20753
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19255
- International Plant Names Index (IPNI) ID: 442376-1
- IUCN ID: 217051
- U.S. Fish and Wildlife Service Species Identifier: Q34N
- U.S.D.A. Plant Symbol: JUJA
- Zipcode Zoo Species Identifier: 44791
Footnotes
- Anmin Lu, Donald E. Stone & L. J. Grauke "Juglandaceae". in Flora of China Vol. 4 Page 277. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Alan T. Whittemore & Donald E. Stone "Juglans". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- World Conservation Monitoring Centre 1998. Juglans jamaicensis. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloaded on 01 February 2012. [back]
