Interesting Facts
Common Names
Common Names in English:
Lady's-slipper
Description
Family Orchidaceae
Herbs or rarely vines
, perennial
, rarely annual
, strongly mycotrophic, epiphytic, terrestrial
, lithophytic, or rarely aquatic
or subterranean
, usually green and photosynthetic, some without chlorophyll and saprophytic
. Roots
subterranean or aerial
, tuberoid
or stolonoid, usually with spongy
, multilayered velamen. Stems erect
or pendent or modified into creeping
rhizomes, simple
or sympodially or monopodially branched, delicate to stout, or thickened as corms or pseudobulbs
, or greatly reduced, sometimes proliferous (especially diverse
in sympodial orchids) . Leaves solitary, several, or reduced to scales
, basal or cauline, alternate, distichous, or sometimes opposite or whorled
, either convolute or duplicate
, simple, sessile or petiolate
; stipules absent; blade
articulate
or not, plicate
or conduplicate
, cylindric
, triangular, or laterally flattened, margins
entire. Inflorescences terminal
or lateral
, racemes
, spikes, panicles, or rarely cymose
, erect or variously pendent, 1 many-flowered, lax
or dense, flowering successively or simultaneously. Flowers bisexual
[rarely unisexual
], epigynous
, resupinate or not, pedicellate
or sessile, 3-merous, usually bilaterally symmetric
[rarely nearly radially symmetric], with abscission layer between pedicel and peduncle, rarely between ovary and perianth or ovary and pedicel; perianth of 6 tepals in 2 whorls, all petaloid
or sepals sometimes greener and more foliaceous
in texture
; sepals alike or not, lateral sepals often connate
(forming synsepal), or all 3 sepals variously connate and/or adnate
or distinct
and/or free
; petals 3, median
petal modified as lip, commonly larger or differing in form and color, lateral petals commonly but not always similar to sepals; nectaries of various sorts; extrafloral nectaries sometimes present on pedicels, bracts, or leaf sheaths
; stamens usually 1 2( 3, if 3 the 3d modified into sterile
staminode), all on side opposite lip, fully or partially adnate to style, forming column; pollen grains
in monads
or tetrads
, usually in 2 8 pollinia, sometimes subdivided into small packets, rarely granular
, sometimes pollinia with caudicles
and/or stipes; gynoecium 3-carpellate, connate, forming compound
, inferior, 1- or 3-locular ovary; style variously adnate to filaments
; stigmas usually 3-lobed, concave
to convex
, part of median stigma lobe
modified into rostellum
, often separating anther
from fertile
portions of stigma, commonly preventing or in some cases facilitating self-pollination
; ovules numerous
, anatropous
, minute. Fruits capsules, opening (dehiscing) by longitudinal
slits, rarely fleshy
and indehiscent berries
. Seeds numerous (millions in some species), minute; endosperm absent.
Genera ca.
800, species 22,000 35,000 (701 genera, 208 species in the flora
; 1 genus, 6 species introduced) : worldwide except Antarctica, most diverse in tropical
forests
.
The overall count for orchid genera in the flora includes Spathoglottis plicata Blume, which was recently reported from Palm Beach
County, Florida. The plants
, known locally since 1982, are apparently widely naturalized
in old shellpits. The number of species in the flora includes one newly recognized species in Habenaria that is morphologically described, but not fully treated here. Orchidaceae are by far the largest and most diverse monocot family
and rank among the largest families of flowering plants. An accurate account of the number of genera and species has eluded orchid scientists, and species counts published in the last 20 years range
from 15,000 to 35,000. New species are continually being described. In addition, numerous natural and artificial hybrids
exist.
Although orchids are important in horticulture
, most of the plants traded in the national and international market belong to a small number of species and their hybrids in only a few genera; the majority of orchids are not commonly cultivated. Few orchids are economically important outside the horticultural trade: the fruits of several species of Vanilla are the source of the spice vanilla, and the dry roots of some species of Dactylorhiza, Eulophia, and Orchis are made into salep, a flour
consumed in northern Africa, the Middle
East (especially Turkey), and Asia. Some species are locally used for medicinal purposes; the mucilage from pseudobulbs of several species is sometimes used as glue; and in the Far East the stems of some species of Dendrobium are split into strips used to weave handicrafts. A few orchids have been found to cause contact dermatitis
(e.g.
, Cypripedium reginae) .
Orchids range vegetatively from Lilliputian plants a few millimeters long (Bulbophyllum Thouars and Platystele Schlechter) to gigantic clusters
weighing several hundred
kilograms
(Grammatophyllum Blume) to some as much as 13.4 meters in height
(Sobralia altissima D. E. Bennett & Christenson, a recently described species from Peru) . Likewise, flowers vary in size from less than 1 mm and barely visible to the naked eye (Platystele Garay), to 15 20 cm diameter (some Paphiopedilum Pfitzer, Phragmipedium Rolfe, and Cattleya Lindley spp.
), and ultimately to 76 cm [Phragmipedium caudatum (Lindley) Rolfe]. Weight
can vary from a fraction of a gram
(many Pleurothallus R. Brown spp.) to nearly 100 grams (Coryanthes Hooker spp.) . Their fragrances vary from delightful (Cattleya Lindley) to repulsive and unbearable (in some species of Bulbophyllum Thouars) . The plants colonize habitats
ranging from some of the driest and hottest places on earth to the wettest and coolest, literally occurring from polar
regions to the equator. Within the monocots, the most important diagnostic features of Orchidaceae are reduction of adaxial
stamens, fusion of the remaining stamens to the gynoecium forming the column, aggregation of pollen into compact
pollinia (present elsewhere only in the dicots
, in Asclepiadaceae), differentiation
of the median petal into the lip, a sometimes complex
organ, and the exceedingly small size of the seed, which lacks endosperm. Among other distinguishing characteristics: pollen in the pollinia is usually not available as a nutrient-source (Cleistes Richard ex
Lindley being a notable exception), and the often complex interaction with pollinators culminates in the phenomenon of pseudocopulation in several genera (e.g., Ophrys Linnaeus, Caladenia R. Brown sect. Calonema, Drakaea Lindley) . In the latter process
, the flower mimics the appearance
, the smell, and often the movements of a female wasp, attracting a male of a suitable species that tries to copulate with the flower. It usually only succeeds in becoming attached to a pollinium
, which will then be transferred if the male tries to copulate with another flower.
Roots of orchids may be covered with velamen, spongy layers derived from the epidermis
; fleshy thickenings of roots are tuberoids (tubers being restricted
to stems) . Stems may be swollen or thickened, underground corms or aerial pseudobulbs. Flowers are often resupinate: the lip (modified median petal) is lowermost, usually as a result of the pedicel being twisted or bent in its development by 180°. Pedicellate ovary, usually used in reference to length
, refers to the combined pedicel and ovary. Flowers are not always borne on pedicels; when they are, it is sometimes difficult to distinguish between a slender ovary and the pedicel. Consequently, because of their slender ovaries, flowers of a racemose spike appear to be pedicellate even though they are sessile, while a spicate raceme has pedicels so short that they appear to be absent. Orchid flowers often have a modified median sepal, the dorsal sepal. Sepals coalescing at their tips
form a synsepal. The middle portion of the upper (adaxial) face
of the lip is the disc: it may be a thickened callus and may bear hairs
, papillae, or other ornamentation. In orchids the style, stigmas, filaments, and one or more anthers are united
to form a column; appendages
projecting
laterally from the stigma are column wings; the lip may be attached to the protrusion at the base
of the column to form a column foot
; lateral sepals that are also attached to the foot
form a mentum
(chin) . In most orchids the column bears a single anther at its apex; the clinandrium
is the cavity
within which the anther is borne or embedded
. Pollen is borne in discrete masses (pollinia) . Genera with mealy
(sectile) pollinia may have pollinia within the anther tapering into a caudicle (stalk
), which is attached to a sticky viscidium
. Those with waxy pollinia have pollinia attached to one or two stipes (of stigmatic
origin
and formed outside the anther), which in turn
are attached to a viscidium. The various aggregations of pollinia, caudicles, stipes, and viscidium form a pollinarium
, the pollination unit
carried by pollinators. The median stigma lobe may have a slender extension
or little beak
(rostellum), which aids in gluing the pollinarium to the pollinator.[1]
Genus Cypripedium
Herbs, perennial
. Roots
closely to widely spaced along rhizome, slender, fleshy
; rhizomes short to elongate
. Stems leafy or scapose
. Leaves alternate, in single radical
pair, or subopposite near midpoint of stem, ascending
to spreading
, plicate
, bases
sheathing
stem. Inflorescences terminal
, solitary; flowers solitary or 2-several in lax
racemose spike; bracts large, foliaceous
. Flowers resupinate, showy; sepals distinct
or lateral
sepals usually connate
proximal
to lip forming synsepal; petals entire; lip inflated, slipper- or sac-shaped, with adaxial
orifice
; pollinaria absent; loose
granular
pollen in 2 lateral anthers, dorsal anther
a large subapical
staminode; stigma free
, 2-3-lobed. Fruits capsules, ellipsoid
to oblong-ellipsoid.
Species ca.
45: primarily temperate
Eurasia
and North America, ranging from arctic
to subtropics in East Asia and Cordilleran North America and Central America.[2]
Physical Description
Habit: Deciduous.
Flowers: Bloom Period: April, May. • Flower Color: maroon
Size/Age/Growth
Size: 12-18" tall.
Biology
Growth
Culture: Space 12-15" apart.
Soil: Minimum pH: 6.1 • Maximum pH: 7.8
Sunlight: Sun Exposure: Full sun to partial shade.
Temperature: Cold Hardiness: 3a, 3b, 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Liliidae
(
)
- Takhtajan, 1967
- Superorder:
Lilianae
(
)
- Takhtajan, 1967
- Order:
Asparagales
(
)
- Bromhead, 1838
- Series:
href="http://www.biodiversitylibrary.org/title/493
(
)
- Family:
Orchidaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- Orchid Family
- Subfamily:
Cypripedioideae
(
)
- Genus:
Cypripedium
(
)
- C. Linnaeus, 1753
- Lady's-slipper, moccasin-flower [Greek Kypris, Aphrodite, and Latin pes, foot, perhaps an orthographic error for Greek pedilon, slipper]
- Form:
l literature
up to the publication of Icones
- Botanical name: - Cypripedium
- Form:
l literature
up to the publication of Icones
- Genus:
Cypripedium
(
- Subfamily:
Cypripedioideae
(
- Family:
Orchidaceae
(
- Series:
href="http://www.biodiversitylibrary.org/title/493
(
- Order:
Asparagales
(
- Superorder:
Lilianae
(
- Subclass:
Liliidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Cypripedium macranthos {var} speciosum • Cypripedium rebunense
Notes
Registrant name
: RHS
Originator name: O/U
Date of registration
: 31/03/2004
Similar Species
Members of the genus Cypripedium
ZipcodeZoo has pages for 187 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
C. (Lady's-Slipper) · C. acaule (Lady's-Slipper Orchid) · C. Achim (Lady's-Slipper) · C. Aki (Lady's-Slipper) · C. alaskanum (Lady's-Slipper) · C. andrewsii (Andrew's Lady's Slipper) · C. andrewsii {nm} favillianum (Lady's-Slipper) · C. andrewsii {nm} landonii (Lady's-Slipper) · C. Annette (Lady's-Slipper) · C. arietinum (Ram's Head Lady's Slipper) · C. Axel (Lady's-Slipper) · C. Barbara Imfeld-Pinkepank (Lady's-Slipper) · C. Bernd (Lady's-Slipper) · C. Bernie (Lady's-Slipper) · C. Betty Maier (Lady's-Slipper) · C. Bill (Lady's-Slipper) · C. Birgit (Lady's-Slipper) · C. Blondi (Lady's-Slipper) · C. Boots (Lady's-Slipper) · C. bulbosum (False Lady's-Slipper Orchid) · C. calceolus (Fraueschub) · C. calcicola (Lady's-Slipper) · C. californicum (California Lady's Slipper) · C. Camiel (Lady's-Slipper) · C. candidum (Small White Lady's-Slipper) · C. Carolin (Lady's-Slipper) · C. Carol Ilene (Lady's-Slipper) · C. Carson (Lady's-Slipper) · C. catherinae (Lady's-Slipper) · C. Chauncey (Lady's-Slipper) · C. Cleo Pinkepank (Lady's-Slipper) · C. columbianum (Lady's-Slipper) · C. cordigerum (Lady's-Slipper) · C. corrugatum (Lady's-Slipper) · C. debile (Lady's-Slipper) · C. Dietrich (Lady's-Slipper) · C. Dominique (Lady's-Slipper) · C. Emil (Lady's-Slipper) · C. Erich Maier (Lady's-Slipper) · C. Erika (Lady's-Slipper) · C. Eurasia (Lady's-Slipper) · C. Fantasy (Lady's-Slipper) · C. fargesii (Lady's-Slipper) · C. farreri (Lady's-Slipper) · C. fasciculatum (Brownie Lady's-Slipper) · C. fasciolatum (Lady's-Slipper) · C. Favillianum (Lady's Slipper) · C. flavum (Lady's-Slipper) · C. Florence (Lady's-Slipper) · C. formosanum (Ladys Slipper) · C. franchetii (Lady's-Slipper) · C. Francis (Lady's-Slipper) · C. froschii (Lady's-Slipper) · C. Gabriela (Lady's-Slipper) · C. Genesis (Lady's-Slipper) · C. Gerhard (Lady's-Slipper) · C. Gidget (Lady's-Slipper) · C. Gisela (Lady's-Slipper) · C. GPH Crimson Promise (Lady's-Slipper) · C. guttatum (Lady Slipper Orchid) · C. guttatum var. guttatum (Spotted Lady's Slipper) · C. guttatum var. yatabeanum (Spotted Lady's Slipper) · C. Günter (Lady's-Slipper) · C. Hank Small (Lady's-Slipper) · C. Hans Arpagaus (Lady's-Slipper) · C. Hans Erni (Lady's-Slipper) · C. Hedi (Lady's-Slipper) · C. Heike (Lady's-Slipper) · C. Henning Pinkepank (Lady's-Slipper) · C. henryi (Henrys Cypripedium) · C. herae (Lady's-Slipper) · C. Hideki Okuyama (Lady's-Slipper) · C. Hilda (Lady's-Slipper) · C. Hildegard (Lady's-Slipper) · C. himalaicum (Lady's-Slipper) · C. Holger (Lady's-Slipper) · C. hybrid A (Lady's-Slipper) · C. hybrid b (Lady's-Slipper) · C. hybrid c (Lady's-Slipper) · C. Inge (Lady's-Slipper) · C. Ingrid (Lady's-Slipper) · C. irapeanum (Flor De Pelicano) · C. Irene (Lady's-Slipper) · C. January Sunshine (Lady's-Slipper) · C. japonicum (Lady's-Slipper) · C. Jimmy (Lady's-Slipper) · C. Johanna (Lady's-Slipper) · C. John (Lady's-Slipper) · C. Johnny Petersen (Lady's-Slipper) · C. Joseph Henry (Lady's-Slipper) · C. José (Lady's-Slipper) · C. Judith (Lady's-Slipper) · C. Julie Ann (Lady's-Slipper) · C. Karel Polivka (Lady's-Slipper) · C. Karl Heinz (Lady's-Slipper) · C. Kathleen Anne Green (Lady's-Slipper) · C. Katrin (Lady's-Slipper) · C. kentuckiense (Kentucky Lady's Slipper) · C. Kristi Lyn (Lady's-Slipper) · C. Laila (Lady's-Slipper)
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Further Reading
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- A biochemic basis for the study of problems of taxonomy, heredity, evolution, etc., with especial reference to the starches and tissues of parent-stock and hybrid-stocks and the starches and hemoglobi Washington, D.C.Carnegie institution of Washington1919 url , p. 13, p. 282, p. 338, p. 339, p. 340, p. 341, p. 352, p. 354, p. 357, p. 817, p. 826.
- A commodity subsector analysis of the U.S. cut flower industry / by Marvin Neal Miller. 1983. url p. 191.
- A cultural table of orchidaceous plants. [Editing by P.A. Gilbert.] Sydney, Shepherd Press, 1946. url , , , p. 105, p. 110, p. 21, p. 24, p. 278, p. 315, p. 93, p. 94, p. 95.
- A dictionary of American plant names / compiled by Willard N. Clute. Joliet, Ill.: W.N. Clute, 1923. url p. 185, p. 47.
- A dictionary of the flowering plants and ferns, by J. C. Willis. CambridgeThe University Press, 1919 url p. 202, p. 271, p. 469, p. 470, p. 471, p. 487.
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- A guide to the poisonous plants and weed seeds of Canada and the northern United States / by R.B. Thomson and H.B. Sifton. Toronto: University of Toronto Press, 1922. url p. 121, p. 13, p. 163.
- A guide to the wild flowers east of the Mississippi and north of Virginia / by Norman Taylor. New York: Greenberg, 1928. url p. 49.
- A list of the fishes of Hawaii, with notes and descriptions of new species, by David Starr Jordan and Eric Knight Jordan. Pittsburgh, Board of trustees of the Carnegie Institute, 1922. url .
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- A manual of organic materia medica and pharmacognosy; an introduction to the study of the vegetable kingdom and the vegetable and animal drugs (with syllabus of inorganic remedial agents) comprising the botanical and physical characteristics, source, con Philadelphia, Blakiston[1917] url p. 128, p. 129, p. 479, p. 53, p. 63, p. xiii, p. xiv.
- A manual of the flora of northern Idaho /Carl Epling and Joe Ewan. 4 1941 1941. url p. 1062, p. 1063, p. 1068, p. 1105.
- A manual of the penicillia, by Kenneth B. Raper and Charles Thom; with the technical assistance and illus. by Dorothy I. Fennel. Baltimore, Williams & Wilkins Co., [c1949] url p. 169.
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- A new British flora; British wild flowers in their natural haunts, described by A. R. Horwood, with 64 plates in col. representing 350 different plants, from drawings by J. N. Fitch and many illus. from photo. London, Gresham Pub. Co., 1919. url p. 217, p. 72.
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Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 30, 2012.
- The Royal Horticultural Society Horticultural Database, available here.
Identifiers
- Global Biodiversity Information Facility Taxonkey: 4490920
- Zipcode Zoo Species Identifier: 4213271
Footnotes
- Gustavo A. Romero-González, Germán Carnevali Fernández-Concha, Robert L. Dressler, Lawrence K. Magrath & George W. Argus "Orchidaceae". in Flora of North America Vol. 26 Page 15, 16, 17, 26, 27, 490, 491, 617. Oxford University Press. Online at EFloras.org. [back]
- Charles J. Sheviak "Cypripedium". in Flora of North America Vol. 26 Page 494, 495, 496, 497, 498, 499. Oxford University Press. Online at EFloras.org. [back]
