Description
Family Verbenaceae
Shrubs
or trees
, sometimes climbing
shrubs, rarely herbs. Indumentum of simple
, stellate
, and/or other complex
hairs
. Leaves opposite or rarely whorled
, without stipules, simple or 3-foliolate, less often palmately [or pinnately] compound
. Inflorescences terminal
or axillary
, racemose, cymose
, spicate
, or thyrses
. Flowers bisexual
or polygamous by abortion
, zygomorphic or rarely actinomorphic
. Calyx persistent
. Corolla 4- or 5- or more lobed
; lobes
usually spreading
, aestivation
overlapping. Fertile
stamens inserted
on corolla tube
, alternate with lobes; filaments
free
; anthers
dorsifixed
, 1- or 2-locular, dehiscing by longitudinal
slits or sometimes a circular pore
. Ovary entire
or 4-grooved, 2-8-locular; ovules 1 or 2 per locule, erect
or pendulous. Style
terminal, simple, entire or 2-cleft. Fruit a drupe or indehiscent capsule, sometimes breaking up into nutlets
. Seeds (1 or) 2-4, endosperm usually absent, seed coat
thin; embryo straight, as long as seed; radicle short, inferior.
Some 91 genera and ca.
2000 species: primarily tropical
and subtropical
, 20 genera and 182 species in China.
The classification of Verbenaceae is in a state of flux, especially regarding its relationship
to Lamiaceae. There is evidence to suggest a significant division between members
of subfamily
Verbenoideae, genera 1-5 in this account, and the remaining genera, including genera 6-20, which for convenience are here referred to as subfamily Viticoideae s.l. The latter are more closely allied to each other and to genera traditionally kept within Lamiaceae (including genera 1-8 in this Flora
) . Avicennia is often placed in a family
of its own, but its affinities are clearly with Viticoideae, especially genera 17-19 in this account which have traditionally been placed in a separate subfamily, Symphorematoideae.Shou-liang Chen & Michael G. Gilbert "Verbenaceae". in Flora of China Vol. 17 Page 1. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
Tribe Senecioneae
The Senecioneae are a tribe of closely related genera that can be recognized most readily by the nature of the pappus and the involucral bracts or phyllaries. The phyllaries are basically in one well developed, often partially or wholly connate series of equal length that closely envelope the head . Frequently there are a few, very much smaller and mostly randomly distributed, often necrotic-tipped bracts near the base of the main series. The pappus is of fine, soft, often pure white capillary hairs . Heads may be either discoid or radiate . -- Gerald Carr.
Genus Vitex
Trees
or shrubs
. Branches glabrous
or sparsely pubescent
. Leaves opposite, palmately (1-) 3-8-foliolate; leaflets
petiolulate
, margin
entire
, dentate
, serrate, or incised. Inflorescences terminal
or axillary
cymes, thyrses
, or panicles; bracts usually small, often early deciduous. Calyx campanulate
, tubular
, or funnelform
, sometimes 2-lipped, usually truncate
or shortly 5-dentate. Corolla blue, white, or yellow, 2-lipped, lower lip 3-lobed with middle
lobe
greatly elongated, upper lip usually 2-lobed. Stamens 4, didynamous
, sometimes exserted; anther
locules attached only at tip
, becoming divaricate
. Ovary 2-4-locular; ovules 1 or 2 per locule. Style
filiform
; stigma 2-cleft. Drupes subtended by enlarged calyx, globose
, ovoid
, or obovoid
, normally 4-locular and 4-seeded but often some locules suppressed
and base
of pyrene forming a hollowed cavity
, endocarp a bony pyrene, mesocarp
generally fleshy
. Seeds obovoid or oblong
, endosperm absent; cotyledons usually fleshy.
About 250 species: chiefly tropical
, few in temperate regions
of both hemispheres; 14 species in China."Vitex". in Flora of China Vol. 17 Page 28. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Lamiidae
(
)
- Takhtajan Ex Reveal, 1992
- Superorder:
Lamianae
(
)
- Takhtajan, 1967
- Order:
Lamiales
(
)
- Bromhead, 1838
- Family:
Verbenaceae
(
)
- Jaume Saint-Hilaire, 1805, Nom. Cons.
- Verbena Family
- Subfamily:
Asteroideae
(
)
- Tribe:
Senecioneae
(
)
- Genus:
Vitex
(
)
- Linnaeus, Sp. Pl. 2: 638. 1753.
- Chastetree
- Specific epithet:
agelaeifolia
- Variety:
rufula
- Botanical name: - Vitex agelaeifolia var. rufula Moldenke
- Variety:
rufula
- Specific epithet:
agelaeifolia
- Genus:
Vitex
(
- Tribe:
Senecioneae
(
- Subfamily:
Asteroideae
(
- Family:
Verbenaceae
(
- Order:
Lamiales
(
- Superorder:
Lamianae
(
- Subclass:
Lamiidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Similar Species
Members of the genus Vitex
There are approximately 569 species in this genus. Here are just 100 of them:
V. acuminata · V. acunae · V. adulterina · V. aesculifolia · V. agelaeifolia · V. agelaeifolia var. rufula · V. agnus · V. agnus-castus (Chaste Tree) · V. agnus-castus 'Abbeville Blue' (Chaste Tree) · V. agnus-castus 'Alba' (Chaste Tree) · V. agnus-castus 'Blue Spire' · V. agnus-castus 'Blushing Bride' · V. agnus-castus 'Blushing Spires' (Chaste Tree) · V. agnus-castus 'Lavender Lady' (Chaste Tree) · V. agnus-castus 'Montrose Purple' (Chaste Tree) · V. agnus-castus 'Rosea' · V. agnus-castus 'Shoal Creek' (Chaste Tree) · V. agnus-castus 'Silver Spire' (Silver Spire Chaste Tree) · V. agnus-castus 'Silver Spires' (Chaste Tree) · V. agnus-castus var. agnus-castus · V. agnus-castus var. caerulea · V. agnus-castus var. latifolia (Chaste Tree) · V. agnus-castus var. subtrisecta · V. agnuscastus · V. agraria · V. aherniana · V. ajugaeflora · V. alata · V. altissima · V. altissima f. alata · V. altmanni · V. amaniensis · V. amazonica · V. amboniensis · V. andongensis · V. angolensis · V. angus-castus · V. annamense · V. annamensis · V. appendiculata · V. appuni · V. arborea · V. articulata · V. aurea · V. avicennioides · V. babula · V. bahiensis · V. bakeri · V. balbi · V. balbisii · V. bankae · V. barbata · V. befotakensis · V. benthamiana · V. benuensis · V. bequaertii · V. beraviensis · V. beraviensis f. pilosa · V. beraviensis f. villosa · V. berteroana · V. betsiliensis · V. betsiliensis subsp. barorum · V. bicolor · V. bignonioides · V. bipindensis · V. bogalensis · V. bogoriensis · V. bojeri · V. bojeri var. suborbicularis · V. bombacifolia · V. bracteata · V. bracteosa · V. brasiliensis · V. brevilabiata · V. brevipetiolata · V. brittoniana · V. buchananii · V. buchneri · V. buddingii · V. bulusanensis · V. bunguensis · V. burmensis · V. caespitosa · V. californica · V. calothyrsa · V. camporum · V. canescens · V. cannabifolia (Chinese Chaste Tree) · V. capitata · V. capitata f. albiflora · V. capulin · V. carbunculorum · V. caribaea · V. carvalhi · V. cauliflora · V. cauliflora var. longifolia · V. cauliflora var. villosissima · V. celebica · V. cestroides · V. chapelieri
Bibliography
- P'ei Chien & Chen Shou-liang, eds. 1982. Verbenaceae. Fl. Reipubl. Popularis Sin. 65(1): 1-229.
- P'ei Chien & Chen Shou-liang, eds. 1982. Verbenaceae. Fl. Reipubl. Popularis Sin. 65(1): 1-229.
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Notes
Identifiers
- Zipcode Zoo Species Identifier: 1115377
