Overview
Climbing or creeping plant from Mexico to Panama with large, 5-9-segmented leaves. Rarely flowers in Hawaii but provides a good ground cover under low light conditions; Porteus.
Common Names
Common Names in English:
African Evergreen, American Evergreen, Arrow Leaf, Arrowhead Vine, Arrowhead-Vine, Goosefoot Plant, Goosefoot Vine, Nephthytis
Description
Family Araceae
Herbs, perennial
, wetland or terrestrial
, occasionally emergent or floating, [often epiphytic or climbing
], usually with milky
or watery latex, rarely colored
. Rhizomes, corms, or stolons present; rhizomes vertical
or horizontal, creeping
at or near surface
, sometimes branched; corms underground, starchy; stolons at or near surface. Stems absent [sometimes aboveground or aerial
]. Cataphylls usually present. Leaves rarely solitary, alternate or clustered; petiole
rarely absent, with sheathing
base
; blade
simple
or compound
[occasionally perforate
], elliptic
to obovate
or spatulate
, occasionally sagittate-cordate, larger than 1.5 cm; venation
parallel or pinnate- or palmate-netted. Inflorescences spadices, each with 3--900 usually tightly grouped, sessile flowers, subtended by spathe
; spathe rarely absent, persistent
(sometimes only proximally) or deciduous, variously colored; spadix cylindric
or ovoid
, various parts occasionally naked or with sterile
flowers. Flowers bisexual
or unisexual
, staminate
and pistillate
usually on same plants
or functionally on different plants, staminate flowers
distal to pistillate when unisexual; perianth absent or present; stamens 2--12, distinct
or connate
in synandria; ovaryies 1, 1--3(--many) -locular, sessile or embedded
in spadix; styles 1; stigmas hemispheric
, capitate, or discoid
[sometimes strongly lobed
]. Fruits berries
, distinct or connate at maturity. Seeds 1--40(--many) per berry.
Genera 105, species more than 3300 (8 genera, 10 species in the flora
; species in 10 additional genera may persist locally within flora area, see talbe 203.1) : nearly worldwide, primarily tropical regions
.
Araceae are best characterized by the inflorescence, a fleshy
cylindric or ovoid, unbranched spadix subtended or surrounded by a spathe. True spathes are absent in the Nearctic
genus Orontium and in the Australian
genus Gymnostachys. Other plant families with a compressed
spadix-like inflorescence, such as Piperaceae and Cyclanthaceae, either do not have a structure equivalent to a spathe (Piperaceae) or have early-deciduous bracts (Cyclanthaceae) . Plants are usually glabrous
, rarely pubescent
or spiny
(pubescent in Pistia) . Many Araceae exhibit typical monocotyledonous
parallel leaf venation, but some genera have net
leaf venation more typical of dicotyledons.
Infrafamilial classification of the Araceae is under active
study. The only classification of the family
to date to utilize modern phylogenetic
techniques (S. J. Mayo et al.
1997) recognizes seven subfamilies, of which three are represented in native
temperate
North American aroid flora: Orontioideae (Orontium, Symplocarpus, Lysichiton) ; Calloideae (Calla) ; and Aroideae (Peltandra, Arisaema, and Pistia) . Acorus, a genus historically included
in Araceae, is treated as a separate family in theat flora based on extensive morphologic and chemical evidence that supports
its removal from Arales (M.
H. Grayum 1987) .
The number of genera of Araceae occurring in temperate North America is low in comparison with other continents, and primitive taxa are disproportionately represented. Orontioideae and Calloideae, which include four of the seven native genera found in the flora area, are the basal clades within Araceae. Plants in these subfamilies possess the primitive states for many characteristics in Araceae and share few derived characteristics with other aroid genera (M. H. Grayum 1990) . The more advanced
genera native to the flora area include one genus endemic to eastern North America (Peltandra), a pantropical
genus with an uncertain native distribution (Pistia), and a genus clearly Eurasian in origin
(Arisaema) .
Araceae contain crystals of calcium oxalate
, which are often cited as causing the intense irritation experienced when handling
or consuming the raw plant tissue of many genera in the family. This supposition is contradicted by the fact that although irritation generally is not produced
by properly cooked plants, the crystals remain after heating. Other compounds must therefore be involved with causing this reaction. Studies of Dieffenbachia demonstrated that a proteolytic enzyme
, as well as other compounds, are responsible for the severe irritation caused by this plant and that raphides
of calcium oxalate do not play a major role (J. Arditti and E. Rodriguez 1982) . Whether irritation is caused by enzymes or crystals, that aspect
of Araceae has resulted in aroid genera being included in many lists
of poisonous plants (e.g.
, K
. F. Lampe and M. A. McCann 1985; G. A. Mulligan and D. B
. Munro 1990; K. D. Perkins and W. W. Payne 1978) .
Despite the toxic
effects of Araceae, species of several genera are cultivated as food plants, mainly as subsistence crops
in tropical
areas. The major edible Araceae are Colocasia esculenta and several species of Xanthosoma, grown primarily for their corms and sometimes for their leaves. Most North American species of Araceae were historically used by Native Americans, as both food and medicine (T. Plowman 1969) . The family, is currently more valued for its many ornamental species
, and is the most important family in North America for indoor foliage
plants (T. B. Croat 1994) . Araceae commonly grown as ornamentals
in American homes
include species of Aglaonema (Chinese-evergreen), Anthurium, Caladium, Dieffenbachia (dumbcane), Epipremnum (golden pothos), Philodendron, Spathiphyllum, Syngonium, and Zantedeschia (calla-lily) .
Plants of some cultivated species of Araceae escape and may persist or naturalize
, especially in warmer climates. One of these species, Colocasia esculenta, is widespread enough to warrant full inclusion in the flora, but other introduced species
of Araceae are very local in occurrence. Uncommon species represented by herbarium
specimens or literature reports as escaped or persisting from cultivation are listed (table
203.1) with distinguishing characteristics and areas of occurrence.[1]
Physical Description
Habit: Vine
Flowers: Bloom Period: blooms repeatedly • Flower Color: near white, white
Size/Age/Growth
Size: 12-18" tall.
Habitat
Typically found at an altitude of 0 to 3,750 meters (0 to 12,303 feet).[2]
Biology
Reproduction
Duration: Perennial
Growth
Soil: Minimum pH: 6.1 • Maximum pH: 7.8
Sunlight: Sun Exposure: Sun to Partial Shade.
Temperature: Cold Hardiness: 9a, 9b, 10a, 10b, 11. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Liliopsida
(
)
- Scopoli, 1760
- Class:
Liliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Nephthytis triphylla Nash
- Syngonium albolineatum W. Bull
- Syngonium gracile Matuda
- Syngonium oerstedianum Schott
- Syngonium podophyllum var. albolineatum (W. Bull) Engl.
- Syngonium podophyllum var. oerstedianum (Schott) Engl.
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: Govaerts R., 11-Nov-2003.
Place of publication
: Bot. Zeitung (Berlin) 9:85. 1851 (Syn.
Aroid. 68. 1856).
Name verified on 23-Feb-2006 by ARS Systematic Botanists. Last updated: 23-Feb-2006.
Similar Species
Members of the genus Syngonium
ZipcodeZoo has pages for 79 species, subspecies, varieties, forms, and cultivars in this genus:
S. amazonicum · S. angustatum (Fivefingers) · S. angustifolium · S. armigerum · S. atrovirens · S. auritum · S. birdseyanum · S. castroi · S. chiapense · S. chocoanum · S. 'Christmas' (Syngonium) · S. crassifolium · S. dennell-smithii · S. dodsonianum · S. donnell-smithii · S. erythrophyllum (Syngonium) · S. foreroanum · S. gentryanum · S. 'Glogo' · S. gracilis · S. harlingianum · S. hastiferum · S. hastifolium · S. hoffmannii · S. 'Jenny' · S. laterinervium · S. llanoense · S. macrocarpum · S. macrophyllum (Arrowhead Vine) · S. mauroanum · S. 'Maya Red' · S. meridense · S. neglectum · S. oduberi · S. 'Pink Allusion' · S. 'Pink Candy' · S. podophyllum (American Evergreen) · S. podophyllum 'Albo-Variegatum' (American Evergreen) · S. podophyllum Schott 'Albo-Virens' · S. podophyllum 'Albolineatum' · S. podophyllum 'Albo Maculatum' · S. podophyllum 'Berry Allusion' · S. podophyllum 'Christmas' · S. podophyllum 'Dali' (American Evergreen) · S. podophyllum 'Emerald Gem' (American Evergreen) · S. podophyllum 'Emerald Gem Variegated' · S. podophyllum 'Golden' · S. podophyllum 'Gold Illusion' (American Evergreen) · S. podophyllum 'Holly M' (American Evergreen) · S. podophyllum 'Imperial White' (White-Veined Arrowhead Vine) · S. podophyllum 'Mango Allusion' (American Evergreen) · S. podophyllum 'Maria Allusion' · S. podophyllum Schott 'Mini Form' · S. podophyllum 'Mottled Arrowhead' (American Evergreen) · S. podophyllum 'Mouse Ears' (American Evergreen) · S. podophyllum 'Neon' (American Evergreen) · S. podophyllum 'Neon Pink' (American Evergreen) · S. podophyllum peliocladum · S. podophyllum 'Regina Red' (American Evergreen) · S. podophyllum 'Silver Knight' · S. podophyllum var. peliocladum · S. podophyllum var. podophyllum · S. podophyllum 'Variegatum' · S. rayi · S. sagittatum · S. salvadorense · S. schottianum · S. sparreorum · S. standleyanum · S. steyermarkii · S. ternatum · S. triphyllum · S. vellozianum · S. vellozianum Schott 'Variegata' · S. venulosum · S. wendlandii · S. 'White Butterfly' (Nephthytis) · S. xanthophidium · S. yurimaguense
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Addisonia: colored illustrations and popular descriptions of plants. New York: New York Botanical Garden, 1916-[1964]. ENG url p. 7.
- Annales de l'Institut colonial de Marseille. Marseille: Institut colonial de Marseille, -1906. FRE url p. 483, p. 653.
- Birdsey, M. R. 1951. The cultivated aroids. 124 –125.
- Croat, T. B. and N. Lambert. 1986. The Araceae of Venezuela. Aroideana 9:211.
- Encke, F. et al. 1984. Zander: Handwörterbuch der Pflanzennamen, 13. Auflage. (Zander ed13)
- Flora of Costa Rica by Paul C. Standley Chicago, 1937. ENG url p. 144.
- Flora of Guatemala / by Paul C. Standley and Julian A. Steyermark. Chicago: Chicago Museum of Natural History, 1958. ENG url p. 354, p. 355, p. 356.
- Flora of Yucatan, by Paul C. Standley. Chicago, Field Museum of Natural History, 1930. ENG url p. 224.
- Flora of the Lancetilla Valley, Honduras, by Paul C. Standley. Chicago, Field Museum of Natural History, 1931. ENG url p. 123.
- Govaerts, R. and D. G. Frodin. 2002. World checklist and bibliography of Araceae (and Acoraceae). (L Araceae)
- Howard, R. 1974 –1989. Flora of the lesser Antilles. (F LAnt)
- Judd, W.S., Campbell, C.S., Kellog, E.A. and Donoghue, M.J. (2002): Plant Systematics: a phylogenetic approach, Sinauer, Sunderland, Mass.
- Just's botanischer jahresbericht. Systematisch geordnetes repertorium der botanischen literatur aller länder. Berlin, Gebr. Borntraeger, 1874-98; GER url p. 325, p. 565.
- Liberty Hyde Bailey Hortorium. 1976. Hortus third. (Hortus 3)
- Liogier, H. A. and L. F. Martorell. 1982. Flora of Puerto Rico and adjacent islands: a systematic synopsis. (F PR)
- Standley, P. C. 1920. Trees and shrubs of Mexico. Contr. U.S. Natl. Herb. 23:1-1721. (TreesS Mex)
- Standley, P. C. and J. A. Steyermark. 1946 –1976. Flora of Guatemala. (F Guat)
- The carnivorous plants, by Francis Ernest Lloyd Waltham, Mass., Chronica Botanica Company, 1942. ENG url p. 349.
- The forests and flora of British Honduras / by Paul C. Standley and Samuel J. Record; in cooperation with the Conservator of Forests and the Agricultural Officer of the Colony. Chicago: [Field Museum of Natural History], 1936. ENG url p. 89.
- Walters, S. M. et al., eds. 1986 –. European garden flora. (Eur Gard F)
- Woodson, R. E. and R. W. Schery, eds. 1943 –1980. Flora of Panama. (F Panama)
- Bown, D. 1988. Aroids: Plants of the Arum Family. Portland.
- Grayum, M. H. 1990. Evolution and phylogeny of the Araceae. Ann. Missouri Bot. Gard. 77: 628--697.
- Lampe, K. F. and M. A. McCann. 1985. AMA Handbook of Poisonous and Injurious Plants. Chicago.
- Mayo, S. J., J. Bogner, and P. C. Boyce. 1997. The Genera of Araceae. 1 vol. + laser disc. [London.]
- Mulligan, G. A. and D. B. Munro. 1990. Poisonous Plants of Canada. Ottawa, Canada.
- Perkins, K. D. and W. W. Payne. 1978. Guide to the Poisonous and Irritant Plants of Florida. Gainesville, Florida.
- Plowman, T. 1969. Folk uses of New World aroids. Econ. Bot. 23: 97--122.
- Thompson, S. A. 1995. Systematics and Biology of the Araceae and Acoraceae of Temperate North America. Ph.D. dissertation. University of Illinois. Add Urbana-Champaign.
- Wilson, K. A. 1960. The genera of the Arales in the southeastern United States. J. Arnold Arbor. 41: 47--72.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 and ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Global Biodiversity Information Facility. Accessed June 07, 2007. http://www.gbif.org Mediated distribution data from 3 providers.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (May 01, 2008)
- World Checklist of Selected Plant FamiliesFeb 2, 2006.
Data Sources
Accessed through GBIF Data Portal November 16, 2007:
- Australian National Herbarium
- , Australian National Herbarium
- Bernice Pauahi Bishop Museum, Bishop Museum Natural History Specimen Data
- Botanical Research Institute of Texas, Andes to Amazon Biodiversity Program
- Comisión nacional para el conocimiento y uso de la biodiversidad, Herbario del Instituto de EcologÃa, A.C., México
- Fairchild Tropical Botanic Garden, Fairchild Tropical Botanic Garden Virtual Herbarium Darwin Core format
- Herbarium of the University of Aarhus, The AAU Herbarium Database
- Herbier de la Guyane, Herbier de la Guyane
- Instituto Nacional de Biodiversidad
- , Biodiversidad de Costa Rica
- Missouri Botanical Garden, Missouri Botanical Garden
- National Herbarium of New South Wales, NSW herbarium collection
- Royal Botanic Gardens, Kew, Royal Botanic Gardens, Kew
- SysTax, Herbarium Universitat Ulm
- SysTax, SysTax
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2662267
- Catalogue of Life Accepted Name Code: Kew-199060
- Global Biodiversity Information Facility Taxonkey: 14252981
- Globally Unique Identifier: urn:lsid:ipni.org:names:89152-1
- GRIN Nomen Number: 36053
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 42553
- U.S.D.A. Plant Symbol: SYPOA
- Zipcode Zoo Species Identifier: 65151
Footnotes
- Sue A. Thompson "Araceae". in Flora of North America Vol. 22. Oxford University Press. Online at EFloras.org. [back]
- Mean = 391.730 meters (1,285.203 feet), Standard Deviation = 765.790 based on 686 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
