, or less often trees
; indumentum usually with peltate scales
. Leaves alternate, stipulate
; leaf blade
usually palmately veined, entire or various lobed
. Flowers solitary, less often in small cymes or clusters
, often aggregated into terminal
or panicles, usually conspicuous
, usually bisexual
in Kydia) . Epicalyx
often present, forming an involucre around calyx, 3- to many lobed. Sepals 5, valvate
. Petals 5, free, contorted, or imbricate, basally adnate
. Stamens usually very many, filaments connate into tube; anthers
1-celled. Pollen spiny
. Ovary superior, with 2-25 carpels, often separating from one another and from axis; ovules 1 to many per locule; style as many or 2 Ã— as many as pistils, apex branched or capitate. Fruit a loculicidal capsule or a schizocarp, separating into individual mericarps, rarely berrylike when mature
(Malvaviscus) ; carpels sometimes with an endoglossum (a crosswise projection from back wall of carpel to make it almost completely septate
. Seeds often reniform
, sometimes conspicuously so.
About 100 genera and ca. 1000 species: tropical and temperate regions of N and S Hemisphere; 19 genera (four introduced ) and 81 species (24 endemic, 16 introduced) in China.
Molecular studies have shown that the members of the Bombacaceae, Malvaceae, Sterculiaceae, and Tiliaceae form a very well-defined monophyletic group that is divided into ten also rather well-defined clades, only two of which correspond to the traditional families Bombacaceae and Malvaceae. Some of the remaining groups are included entirely within either of the remaining families but others cut across the traditional divide between the Sterculiaceae and Tiliaceae. A majority of authors , most notably Bayer and Kubitzki (Fam. Gen. Vasc. Pl. 5: 225-311. 2003), has favored including everything within a greatly enlarged Malvaceae, and treating the individual clades as subfamilies. The alternative view is that the individual clades should be treated as a series of ten families: Bombacaceae (Bombacoideae), Brownlowiaceae (Brownlowioideae), Byttneriaceae (Byttnerioideae), Durionaceae (Durionoideae), Helicteraceae (Helicteroideae), Malvaceae (Malvoideae), Pentapetaceae (Dombeyoideae), Sparrmanniaceae (Grewioideae), Sterculiaceae (Sterculioideae), and Tiliaceae (Tilioideae) (Cheek in Heywood et al. , Fl. Pl. Fam. World. 201-202. 2007) . For the present treatment, we prefer to retain the familiar, traditional four families, so as to maintain continuity with the treatments in FRPS, and to await a consensus on the two alternative strategies for dealing with the very widely accepted clades.
The traditional Malvaceae coincides exactly with one of the major clades. The only possible problem is the relationship with the Bombacaceae, which also has primarily 1-loculed anthers, and some authorities have suggested that the Bombacaceae should be included within the Malvaceae.
Members of the Malvaceae are important as fiber crops (particularly cotton, Gossypium) . Young leaves of many species can be used as vegetables, and species of Abelmoschus and Hibiscus are grown as minor food crops. Many species have attractive flowers and an ever-increasing selection is grown as ornamentals . Several have been cultivated for a very long time, particularly species of Hibiscus, and some of these are not known in the wild.
. Leaves simple
, entire or palmately lobed
, rarely palmately compound
entire or dentate
. Inflorescence usually axillary
, usually paniculate
, rarely racemose. Flowers unisexual
. Calyx 5-lobed or 5-partite. Petals absent. Male flowers: anthers
clustered at top of androgynophore
, enclosing undeveloped
carpels. Female flowers: androgynophore very short, staminodes at top of androgynophore in whorl around base
of carpels. Carpels 5; ovules 2 to many per carpel; styles connate
at base; stigmas as many as carpels, free
. Fruit a group of follicles, usually leathery, less often woody, dehiscent
. Seeds 1 to many per follicle, usually with endosperm.
Between 100 and 150 species: tropics and subtropics of both hemispheres, most abundant in Asian tropics; 26 species (14 endemic, one introduced ) in China.
The fruits of nine species and male and/or female flowers of a further four species have not been recorded, indicating that the Chinese members of this genus require much more study in the field .
Biome: Terrestrial .
Ecology: Occurred in subtropical forest between 1,000 and 1,500 m. .
- Whittaker & Margulis,1978
- Haeckel, 1866
- Cavalier-Smith, 1981
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Kenrick & Crane, 1997
- Brongniart, 1843
- Takhtajan, 1967
- Superorder: Malvanae () - Takhtajan, 1967
- Subclass: Rosidae () - Takhtajan, 1967
- Class: Spermatopsida () - Brongniart, 1843
- Infraphylum: Radiatopses () - Kenrick & Crane, 1997
- Subphylum: Euphyllophytina ()
- Phylum: Tracheophyta () - Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae () - Cavalier-Smith, 1981
- Kingdom: Plantae () - Haeckel, 1866 - Plants
Publishing author : Debb. Publication : U. N. & P. C. Kanj. & Das, Fl. Assam i. Pt . 1, 154 (1934); Debbarm. exBiswas in Assam Forest Rec., Bot. i. 5 (1934)
Members of the genus Sterculia
ZipcodeZoo has pages for 12 species, subspecies, varieties, forms, and cultivars in this genus:
S. africana (African Star Chestnut) · S. apetala (Panama Tree) · S. ceramica (Fairchild's Sterculia) · S. colorata (Bonfire Tree) · S. foetida (Hazel Sterculia) · S. oblonga (Yellow Sterculia) · S. rhinopetala (Brown Sterculia) · S. rogersii (Ulumbu Tree) · S. setigera (Sterculia) · S. tragacantha (Gum Tragacanth) · S. urens (Sterculia) · S. verticillata (Cola)
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- 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 577.
- The World List of Threatened Trees WCMC, IUCN url p. 526.
- Feng Kuo-mei. 1984. Malvaceae. In: Feng Kuo-mei, ed., Fl. Reipubl. Popularis Sin. 49(2): 1-102.
- Hsue Hsiang-hao. 1984. Sterculiaceae. In: Feng Kuo-mei, ed., Fl. Reipubl. Popularis Sin. 49(2): 112-189.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 11, 2012.
- IUCN 2012. IUCN Red List of Threatened Species. Version 2011.2. . Downloaded on January 28, 2012.
- World Conservation Monitoring Centre 1998. Sterculia khasiana. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloadedon 05February2012.
- Biodiversity Heritage Library NamebankID: 7143280
- Global Biodiversity Information Facility Taxonkey: 15870970
- Globally Unique Identifier: urn:lsid:ipni.org:names:825296-1
- International Plant Names Index (IPNI) ID: 825296-1
- IUCN ID: 243821
- Zipcode Zoo Species Identifier: 1021171
- Ya Tang, Michael G. Gilbert & Laurence J. Dorr "Malvaceae". in Flora of China Vol. 12 Page 240, 264,299, 302. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Sterculia". in Flora of China Vol. 12 Page 302, 303,327. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- World Conservation Monitoring Centre 1998. Sterculia khasiana. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloaded on 05 February 2012. [back]