Overview
Interesting Facts
Common Names
Common Names in English:
Addersmeat, Easter-Bell, Greater Stichwort, Greater Stitchwort
Description
Family Caryophyllaceae
Herbs annual
or perennial
, rarely subshrubs
or shrubs
. Stems and branches usually swollen at nodes. Leaves opposite, decussate, rarely alternate or verticillate
, simple
, entire, usually connate
at base
; stipules scarious
, bristly
, or often absent. Inflorescence of cymes or cymose
panicles, rarely flowers solitary or few in racemes
, capitula, pseudoverticillasters, or umbels. Flowers actinomorphic
, bisexual
, rarely unisexual
, occasionally cleistogamous
. Sepals (4 or) 5, free
, imbricate, or connate into a tube
, leaflike or scarious, persistent
, sometimes bracteate
below calyx. Petals (4 or) 5, rarely absent, free, often comprising claw
and limb; limb entire or split, usually with coronal scales
at juncture of claw and limb. Stamens (2--) 5--10, in 1 or 2 series. Pistil 1; carpels 2--5, united
into a compound
ovary. Ovary superior, 1-loculed or basally imperfectly 2--5-loculed. Gynophore
present or absent. Placentation free, central, rarely basal; ovules (1 or) few or numerous
, campylotropous. Styles (1 or) 2--5, sometimes united at base. Fruit usually a capsule, with pericarp crustaceous
, scarious, or papery
, dehiscing by teeth or valves
1 or 2 Ã as many as styles, rarely berrylike with irregular dehiscence or an achene. Seeds 1 to numerous, reniform
, ovoid
, or rarely dorsiventrally compressed
, abaxially grooved
, blunt
, or sharply pointed
, rarely fimbriate-pectinate; testa granular
, striate
or tuberculate
, rarely smooth
or spongy
; embryo strongly curved
and surrounding perisperm
or straight but eccentric
; perisperm mealy.
Between 75 and 80 genera and ca.
2000 species: widespread but mainly of temperate
or warm-temperate occurrence in the N hemisphere, with principal centers of distribution in the Mediterranean region and W Asia to W China and the Himalayas, fewer species in Africa S of the Sahara, America, and Oceania; 30 genera (two endemic) and 390 species (193 endemic) in China.
Arenaria, Silene, and Stellaria contain over half the species in the family
in China. They are mostly concentrated in the Qinghai-Xizang plateau
, and are especially rich from the Hengduan Mountains to the Himalayas. The main uses of this family are medicinal and ornamental
. Dianthus superbus, Pseudostellaria heterophylla, Stellaria dichotoma var. lanceolata, and Vaccaria hispanica are commonly used in traditional Chinese medicine
. Some species of Arenaria, Dianthus, Gypsophila, Psammosilene, and Silene are used as medicinal herbs among the people or are habitually used in local Chinese medicine. Many species of Dianthus, Gypsophila, Lychnis, Saponaria, and Silene are grown as ornamentals. Atocion armeria (Linnaeus) Rafinesque ( Silene armeria Linnaeus), native
to Russia and Europe, is also cultivated in China. It differs from Silene in having a corymbose
inflorescence and obscure
calyx veins. Wu Cheng-yih, Ke Ping, Zhou Li-hua, Tang
Chang-lin & Lu De-quan. 1996. Caryophyllaceae. In: Tang Chang-lin, ed., Fl.
Reipubl. Popularis Sin. 26: 47â449.[1]
Genus Stellaria
Plants annual
, winter annual
, or perennial
. Taproots usually slender, perennial taxa often rhizomatous
, rooting
at nodes. Stems prostrate
to ascending
or erect
, simple
or branched, terete
or 4-angled. Leaves sometimes connate
basally into sheath
, often sessile; blade
1-veined, linear
or lanceolate to ovate
or deltate, succulent (S. crassifolia [gemmae], S. fontinalis, S. humifusa, and S. irrigua) or not, apex acute or obtuse
. Inflorescences terminal
, open cymes, rarely axillary
(S. alsine, S. americana) or umbellate
(S. umbellata), or terminal or axillary solitary flowers; bracts paired
(1 in S. dicranoides), foliaceous
, scarious
and reduced, or absent. Pedicels erect, sometimes reflexed
in fruit, glabrous
or pubescent
, not glandular
. Flowers usually bisexual
(S. dicranoides unisexual
) ; perianth and androecium hypogynous or weakly perigynous; hypanthium cup- or disc-shaped; sepals (4-) 5, distinct
, green, occasionally purple tinged (S. irrigua) or red proximally (S. pallida), lanceolate to ovate-triangular, 2-12 mm, herbaceous (rarely coriaceous
), margins
often white, scarious, apex acute, acuminate, or obtuse, not hooded
; petals (1-) 5 or absent, white (sometimes translucent
in S. borealis), not clawed, blade apex 2-fid usually for 3- 5 its length
(S. holostea occasionally laciniate
) ; nectaries at base
of filaments
opposite sepals usually present, disc sometimes prominent
; stamens (1-) 5 or 10 or absent, arising from nectariferous
disc (prominent in S. dicranoides and S. irrigua) at ovary base; filaments distinct; staminodes absent; styles [2-]3(-5), capitate to clavate
, 0.2-7 mm, glabrous proximally; stigmas [2-]3(-5), terminal or subterminal
, papillate
(30×). Capsules globose
to conic, opening by 3 or 6, occasionally 4, 8, or 10 ascending to recurved valves
; carpophore present or absent. Seeds (1-) 3-20+, yellow-brown to dark brown, globose to ellipsoid
, laterally compressed
, rarely shiny, papillate or rugose
, rarely smooth
, marginal
wing absent, appendage
absent. x = 10, 11, 12, 13, 15.
Species ca.
120: worldwide, mainly north-temperate regions.[2]
Physical Description
Species Stellaria holostea
Plants perennial , scrambling to ascending , from slender, creeping rhizomes. Stems branched dis-tally, 4-angled, 15-60 cm, glabrous or hispid-puberulent distally. Leaves sessile; blade narrowly lanceolate, widest near base , 4-8 cm × 2-10 mm, somewhat coriaceous , base round and clasping , margins and abaxial midrib very rough, apex narrowly and sharply acuminate, scabrid , otherwise glabrous, slightly glaucous. Inflorescences terminal , loose , 3-31-flowered cymes; bracts foliaceous , 5-50 mm, margins and abaxial midrib scabrid. Pedicels ascending, 1-60 mm, slender, pubescent . Flowers 20-30 mm diam.; sepals 5, inconspicuously 3-veined, ovate-lanceolate, 6-8 mm, margins narrow, scarious , apex acute, glabrous; petals 5 (rarely absent), 8-14 mm, longer than sepals, blade apex 2-fid to middle ; stamens 10, sometimes fewer by degeneration ; styles 3, ascending, ca. 4 mm. Capsules green, subglobose, 5-6 mm, ± equaling sepals, apex obtuse , opening by 3 valves , tardily splitting into 6; carpophore absent. Seeds reddish brown, reniform , 2-3 mm diam., papillose . 2n = 26 (Europe). [source]
Habit: Forb/herb
Flowers: Bloom Period: February, March, April. • Flower Color: near white, white
Size/Age/Growth
Size: 12-18" tall.
Habitat
Woodlands, hedgerows; 0-500 m [3].
Typically found at an altitude of 0 to 1,303 meters (0 to 4,275 feet).[4]
Biology
Reproduction
Duration: Perennial
Growth
Culture: Space 15-18" apart.
Soil: Minimum pH: 6.1 • Maximum pH: 7.8
Sunlight: Sun Exposure: Full Sun .
Temperature: Cold Hardiness: 3a, 3b, 4a, 4b, 5a, 5b, 6a, 6b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Caryophyllidae
(
)
- Takhtajan, 1967
- Superorder:
Caryophyllanae
(
)
- Takhtajan, 1967
- Order:
Caryophyllales
(
)
- Perleb, 1826
- Suborder:
Caryophyllineae
(
)
- Family:
Caryophyllaceae
(
)
- Durande, 1782 ex A.L. de Jussieu, 1789, nom. cons.
- cariophyllaces, pinks
- Subfamily:
Alsinoideae
(
)
- Subfamily:
Alsinoideae
(
- Family:
Caryophyllaceae
(
- Suborder:
Caryophyllineae
(
- Order:
Caryophyllales
(
- Superorder:
Caryophyllanae
(
- Subclass:
Caryophyllidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Alsine graminea (L.) Britt. • Alsine holostea (Linnaeus) Britton • Alsine holstea (L.) Britton • Stellaria graminea var. latifolia Peterm.
Notes
Name
Status: Accepted Name
.
Last scrutiny: 15-Mar-2000
Similar Species
Members of the genus Stellaria
ZipcodeZoo has pages for 51 species, subspecies, varieties, forms, and cultivars in this genus:
S. alaskana (Alaska Starwort) · S. alsine (Bog Chickweed) · S. americana (American Chickweed) · S. antillana (West Indian Starwort) · S. borealis (Boreal Starwort) · S. borealis borealis (Boreal Starwort) · S. borealis subsp. sitchana (Sitka Starwort) · S. calycantha (Northern Chickweed) · S. ciliatosepala (Edwards' Starwort) · S. corei (Tennessee Chickweed) · S. crassifolia (Fleshy Starwort) · S. crassifolia var. crassifolia (Fleshy Starwort) · S. crassifolia var. eriocalycina (Fleshy Starwort) · S. crassifolia var. linearis (Fleshy Starwort) · S. crassipes (Tundra Starwort) · S. crispa (Crisp Starwort) · S. cuspidata (Mexican Starwort) · S. dicranoides (Chamisso's Starwort) · S. fontinalis (American Water Starwort) · S. graminea (Common Starwort) · S. graminea var. graminea (Grass-Leaf Starwort) · S. holostea (Addersmeat) · S. humifusa (Salt-Marsh Starwort) · S. irrigua (Colorado Starwort) · S. littoralis (Beach Starwort) · S. longifolia (Long-Leaf Starwort) · S. longifolia var. atrata (Longleaf Starwort) · S. longifolia var. eciliata (Longleaf Starwort) · S. longifolia var. longifolia (Longleaf Chickweed) · S. longipes (Long-Stalk Starwort) · S. longipes longipes (Long-Stalk Starwort) · S. longipes longipes var. longipes (Meadow Starwort) · S. longipes subsp. arenicola (Longstalk Starwort) · S. media (Chickweed) · S. media media (Chickweed) · S. media neglecta (Chickweed) · S. media subsp. neglecta (Common Chickweed) · S. nemorum (Wood Stitchwort) · S. nitens (Shiny Chickweed) · S. obtusa (Blunt-Sepaled Starwort) · S. oxyphylla (Robust Starwort) · S. pallida pallida (Common Chickweed) · S. palustris (Marsh Stitchwort) · S. parva (Pygmy Starwort) · S. porsildii (Porsild's Starwort) · S. prostrata (Prostrate Starwort) · S. pubera (Great Chickweed) · S. ruscifolia (Circumpolar Starwort) · S. ruscifolia subsp. aleutica (Circumpolar Starwort) · S. sitchana sitchana (Sitka Starwort) · S. umbellata (Umbellate Chickweed)
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Further Reading
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Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Data Sources
Accessed through GBIF Data Portal November 11, 2007:
- Biologiezentrum der Oberoesterreichischen Landesmuseen, Biologiezentrum Linz
- Conservatoire botanique national du Bassin parisien, Conservatoire botanique national du Bassin parisien
- GBIF-Spain, Departamento de Biolog. Veg. II, Facultad de Farmacia, Universidad Complutense, Madrid: MAF
- Jyväskylä University Museum - The Section of Natural Sciences, Vascular plant collection of Jyvaskyla University Museum
- Marine Science Institute, UCSB, Paleobiology Database
- Missouri Botanical Garden, Missouri Botanical Garden
- Museum of Natural History, Wroclaw University, Museum of Natural History, Wroclaw University, Flora of the Stołowe Mts.
- NLBIF, Limnodata
- Natural History Museum, University of Oslo, Vascular Plant Herbarium, Oslo
- Natural History Museum, University of Oslo, Vascular Plants, Field notes, Oslo
- SysTax, Herbarium Universitat Ulm
- The Danish Biodiversity Information Facility, Botany registration database by Danish botanists
- The Swedish Museum of Natural History
- , Botany
- The Swedish Museum of Natural History
- , Herbarium of Oskarshamn
- The Swedish Museum of Natural History
- , Lund Botanical Museum
- The Swedish Museum of Natural History
- , Plants
- UK National Biodiversity Network, Environment and Heritage Service - EHS Species Datasets
Identifiers
- Biodiversity Heritage Library NamebankID: 1
- Catalogue of Life Accepted Name Code: ITS-20182
- Global Biodiversity Information Facility Taxonkey: 4395043
- Globally Unique Identifier: urn:lsid:ipni.org:names:159248-1
- GRIN Nomen Number: 35496
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 20182
- U.S.D.A. Plant Symbol: STHO2
- Zipcode Zoo Species Identifier: 20655
Footnotes
- Dequan Lu, Zhengyi Wu, Lihua Zhou, Shilong Chen, Michael G. Gilbert, Magnus Lidn, John McNeill, John K. Morton, Bengt Oxelman, Richard K. Rabeler, Mats Thulin, Nicholas J. Turland & Warren L. Wagner "Caryophyllaceae". in Flora of China Vol. 6 Page 1. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- John K. Morton "Stellaria". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org. [back]
- "Stellaria holostea". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org. [back]
- Mean = 107.990 meters (354.298 feet), Standard Deviation = 173.410 based on 855 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
