Interesting Facts
Description
Family Amaranthaceae
Herbs, clambering
subshrubs
, shrubs
, or lianas. Leaves alternate or opposite, entire, exstipulate
. Flowers small, bisexual
or unisexual
, or sterile
and reduced, subtended by 1 membranous bract and 2 bracteoles, solitary or aggregated in cymes. Inflorescences elongated or condensed spikes (heads
), racemes
, or thyrsoid
structures of varying complexity. Bracteoles membranous or scarious
. Tepals 3-5, membranous, scarious or subleathery, 1-, 3-, 5-, or 7(-23) -veined. Stamens as many as tepals and opposite these, rarely fewer than tepals; filaments
free
, united
into a cup
at base
or ± entirely into a tube
, filament lobes present or absent, pseudostaminodes present or absent; anthers
(1- or) 2-loculed, dorsifixed
, introrsely dehiscent
. Ovary superior, 1-loculed; ovules 1 to many; style persistent
, short and indistinct or long and slender; stigma capitate, penicillate
, 2-lobed or forming 2 filiform
branches. Fruit a dry utricle or a fleshy
capsule, indehiscent, irregularly bursting, or circumscissile. Seeds lenticular
, reniform
, subglobose, or shortly cylindric
, smooth
or verruculose
.
About 70 genera and 900 species: worldwide; 15 genera (one introduced
) and 44 species (three endemic, 14 introduced) in China.
Morphology of the androecium, perianth (tepals), and the inflorescence has traditionally been used to circumscribe genera and tribes
. Pseudostaminodia are interstaminal appendages
with variously shaped apices. Filament appendages are the lateral
appendages of filaments (one on each side) . The basic structure of the inflorescence is the cyme (branchlets
arising from the bracteole axils, the bracteoles serving as bracts for upper flowers), which can be reduced to one flower with two bracteoles and a bract. Units
of dispersal
vary considerably (capsules opening with lower part persistent, flower and bracteoles falling together, or cymose
partial inflorescences breaking off above bract) and can be characteristic for genera. Several genera possess long trichomes
serving dispersal at the base of the tepals.[1]
Genus Spinacia
Herbs, annual
or biennial, monoecious, glabrous
. Stems erect
, not branched, not jointed
, not armed
, not fleshy
. Leaves basal or alternate, petiolate
; blade
triangular-hastate to ovate
, sometimes with elongated lobes
, margins
entire or dentate
, apex acute. Inflorescences: pistillate
flowers sessile in leaf axils
; staminate flowers
in dense terminal
spikes. Flowers unisexual
, rarely bisexual
; staminate flowers with perianth segments 4-5, stamens 4-5; pistillate flowers enclosed by 2 accrescent
or connate
bracts, perianth absent, stigmas 4-5. Fruiting structures enlarged and hardened bracteoles enclosing achenes; pericarp adherent
. Seeds vertical
, seed coat
dark, spiny
or smooth
; hardened bracteoles serving as seed coat; embryo annular
; perisperm
copious
. x = 6.
Species 3: introduced
; Eurasia
.
The chromosome base number
of six is unusual in the Chenopodiaceae. B
. L. Turner (1994b) suggested that members
with a base
of nine, the most common number in the family
, may be derived.[2]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Caryophyllidae
(
)
- Takhtajan, 1967
- Superorder:
Caryophyllanae
(
)
- Takhtajan, 1967
- Order:
Caryophyllales
(
)
- Perleb, 1826
- Suborder:
Chenopodiineae
(
)
- Family:
Amaranthaceae
(
)
- Adanson, 1763 ex A.L. de Jussieu, 1789, nom. cons.
- amaranthes, pigweed
- Subfamily:
Chenopodioideae
(
)
- Tribe:
Atripliceae
(
)
- Genus:
Spinacia
(
)
- C. Linnaeus, 1753
- Spinach [Latin spina, spine, in reference to spiny fruit, or Persian ispanakh, spinach]
- Specific epithet:
oleracea
- Subspecies:
occidentalis
- Botanical name: - Spinacia oleracea occidentalis
- Subspecies:
occidentalis
- Specific epithet:
oleracea
- Genus:
Spinacia
(
- Tribe:
Atripliceae
(
- Subfamily:
Chenopodioideae
(
- Family:
Amaranthaceae
(
- Suborder:
Chenopodiineae
(
- Order:
Caryophyllales
(
- Superorder:
Caryophyllanae
(
- Subclass:
Caryophyllidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Similar Species
Members of the genus Spinacia
ZipcodeZoo has pages for 95 species, subspecies, varieties, forms, and cultivars in this genus:
S. oleracea (Spinach) · S. oleracea '7-Green' (Spinach) · S. oleracea 'Amazon' (Spinach 'amazon') · S. oleracea 'America' (Spinach) · S. oleracea 'Andhalu' (Spinach) · S. oleracea 'Avenger' (Spinach) · S. oleracea 'Avon' (Spinach) · S. oleracea 'Bahamas' (Spinach 'bahamas') · S. oleracea 'Barbados' (Spinach 'barbados') · S. oleracea 'Bizet' (Spinach) · S. oleracea 'Blackbird' (Spinach) · S. oleracea 'Bloomsdale' (Spinach) · S. oleracea 'Bolero' (Spinach) · S. oleracea 'Bordeaux' (Spinach) · S. oleracea 'Bossanova' (Spinach) · S. oleracea 'Catalina' (Spinach) · S. oleracea 'Cherokee' (Spinach) · S. oleracea 'Correnta' (Spinach) · S. oleracea 'Cxf 94684' (Spinach) · S. oleracea 'Cypress' (Spinach) · S. oleracea 'Dolphin Rz' (Spinach) · S. oleracea 'Emilia' (Spinach) · S. oleracea 'Esa661070' (Spinach 'esa661070') · S. oleracea 'Esmeralda' (Spinach) · S. oleracea 'Falcon' (Spinach) · S. oleracea 'Fidalgo' (Spinach) · S. oleracea 'Fiorano' (Spinach 'fiorano') · S. oleracea 'Four Season' (Spinach) · S. oleracea 'Galaxy' (Spinach) · S. oleracea 'Giant Nobel' (Spinach) · S. oleracea 'Giant Winter' (Spinach) · S. oleracea 'Green Magic' (Spinach) · S. oleracea 'Hector' (Spinach) · S. oleracea 'Hellcat' (Spinach) · S. oleracea 'Industra' (Spinach 'industra') · S. oleracea 'Interceptor' (Spinach) · S. oleracea 'Lorelay' (Spinach) · S. oleracea 'Matador' (Spinach) · S. oleracea 'Melody' (Spinach) · S. oleracea 'Merlo Nero' (Spinach) · S. oleracea 'Mig' (Spinach) · S. oleracea 'Mikado' (Spinach 'mikado') · S. oleracea 'Missouri' (Spinach 'missouri') · S. oleracea 'Monstrueux De Viroflay' (Spinach) · S. oleracea 'Nadong' (Spinach) · S. oleracea 'Okame' (Spinach) · S. oleracea 'Olympia' (Spinach) · S. oleracea 'Orai - Alrite' (Spinach) · S. oleracea 'Oriental Giant' (Spinach) · S. oleracea 'Padre' (Spinach) · S. oleracea 'Palco' (Spinach) · S. oleracea 'Polka' (Spinach) · S. oleracea 'Polydane' (Spinach) · S. oleracea 'Razzle Dazzle' (Spinach) · S. oleracea 'Regiment' (Spinach) · S. oleracea 'Rhino' (Spinach 'rhino') · S. oleracea 'Ricco Di Asti' (Spinach) · S. oleracea 'Sais #7' (Spinach) · S. oleracea 'Samish' (Spinach) · S. oleracea 'Scarlet' (Spinach) · S. oleracea 'Scenic' (Spinach) · S. oleracea 'Seb661076' (Spinach 'seb661076') · S. oleracea 'Seychelles' (Spinach 'seychelles') · S. oleracea 'Skookum' (Spinach) · S. oleracea 'Sp870' (Spinach) · S. oleracea 'Space' (Spinach) · S. oleracea 'Spain' (Spinach) · S. oleracea 'Spinnaker' (Spinach) · S. oleracea 'Spinner' (Spinach) · S. oleracea 'Spokane' (Spinach) · S. oleracea 'Sporter' (Spinach) · S. oleracea 'Spring-Summer' (Spinach) · S. oleracea 'Springer' (Spinach) · S. oleracea 'Springfield' (Spinach) · S. oleracea 'Summer Perfection' (Spinach) · S. oleracea 'Tetona' (Spinach) · S. oleracea 'Tigercat' (Spinach) · S. oleracea 'Tirza' (Spinach 'tirza') · S. oleracea 'Tornado' (Spinach 'tornado') · S. oleracea 'Tyee' (Spinach) · S. oleracea 'Unipack 12' (Spinach) · S. oleracea 'Unipack 131' (Spinach) · S. oleracea 'Unipack 144' (Spinach) · S. oleracea 'Unipack 151' (Spinach) · S. oleracea 'Unipack 277' (Spinach) · S. oleracea 'Vancouver' (Spinach) · S. oleracea 'Van Gogh' (Spinach) · S. oleracea 'Verdi' (Spinach) · S. oleracea 'Viking' (Spinach) · S. oleracea 'Viroflay' (Spinach) · S. oleracea 'Vivos' (Spinach) · S. oleracea 'Whale' (Spinach) · S. oleracea 'Winter Bloomsdale' (Spinach) · S. sativus 'Seven R' (Spinach) · S. turkestanica (Wild Spinach)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Kuan Ke-chien. 1979. Amaranthaceae. In: Kung Hsien-wu & Tsien Cho-po, eds., Fl. Reipubl. Popularis Sin. 25(2): 194241.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 15, 2012.
Identifiers
- Zipcode Zoo Species Identifier: 2219529
Footnotes
- Bojian Bao, Thomas Borsch & Steven E. Clemants "Amaranthaceae". in Flora of China Vol. 5 Page 415. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Leila M. Shultz "Spinacia". in Flora of North America Vol. 4 Page 260, 302. Oxford University Press. Online at EFloras.org. [back]
