Interesting Facts
Common Names
Common Names in English:
Amatillo, Twinleaf Nightshade, Twoleaf Nightshade
Description
Family Solanaceae
Herbs, shrubs
, small trees
, or climbers
. Stems sometimes prickly, rarely thorny; hairs
simple
, branched, or stellate
, sometimes glandular
. Leaves alternate, solitary or paired
, simple or pinnately compound
, without stipules; leaf blade
entire, dentate
, lobed
, or divided
. Inflorescences terminal
, overtopped by continuing axes, appearing axillary
, extra-axillary
, or leaf opposed, often apparently umbellate
, racemose, paniculate
, clustered, or solitary flowers, rarely true cymes, sometimes bracteate
. Flowers mostly bisexual
, usually regular, 5-merous, rarely 4- or 6-9-merous. Calyx mostly lobed. Petals united
. Stamens as many as corolla lobes
and alternate with them, inserted
within corolla, all alike or 1 or more reduced; anthers
dehiscing longitudinally or by apical pores
. Ovary 2-5-locular; placentation mostly axile
; ovules usually numerous
. Style 1. Fruiting calyx often becoming enlarged, mostly persistent
. Fruit a berry or capsule. Seeds with copious
endosperm; embryo mostly curved
.
About 95 genera with 2300 species: best represented in western tropical
America, widespread in temperate
and tropical regions
; 20 genera (ten introduced
) and 101 species in China.
Some species of Solanaceae are known in China only by plants
cultivated in ornamental
or specialty gardens: Atropa belladonna Linnaeus, Cyphomandra betacea (Cavanilles) Sendtner, Brugmansia suaveolens (Willdenow) Berchtold & Presl, Nicotiana alata Link & Otto, and Solanum jasminoides Paxton.[1]
Genus Solanum
Herbs, shrubs
, climbers
, or small trees
, sometimes prickly; hairs
simple
, branched, or stellate
, sometimes glandular
. Leaves solitary or paired
, simple or pinnately compound
, mostly petiolate
; leaf blade
entire, dentate
, lobed
, or parted
. Inflorescences axillary
, extra-axillary
, or leaf opposed, mostly racemose, paniclulate, umbellate
, fasciculate, or solitary flowers, without bracts; peduncle branched or not, sometimes obsolete
. Flowers bisexual
or andromonoecious, mostly actinomorphic
, often 5-merous. Calyx mostly lobed partway and splitting
further at sinuses. Corolla mostly rotate or stellate. Stamens inserted
high in corolla tube
; anthers
often connivent or connate
around style, dehiscing by apical pores
, often later splitting longitudinally. Ovary 2-5-locular, with enlarged placentae; ovules axile
, numerous
. Stigma small. Fruiting calyx persistent, sometimes enlarged and enclosing berry. Berries
mostly juicy. Seeds discoid
or lenticular
; embryo strongly curved
.
About 1200 species: most in tropical
and subtropical
regions, especially in the Americas; 41 species in China, about half introduced
from elsewhere.[2]
Physical Description
Species Solanum diphyllum
Shrubs glabrous . Stems erect , 0.5-2 m tall, young stems angled proximal to nodes from decurrent leaf bases , otherwise terete . Leaves unequal paired ; petiole of major leaf 1-1.5 cm, winged by leaf bases; blade of major leaf elliptic to oblong , sometimes broad, 4-9 × 2-3.5 cm, base attenuate and decurrent on petiole and stem, margin entire, apex rounded ; minor leaves subsessile , blade rounded, 1.5-3 × 1.2-2.2 cm, entire. Inflorescences leaf opposed, short scorpioid racemes , often subumbellate; peduncle unbranched, 3-12 mm, later bearing conspicuous pedicel scars . Pedicel 5-12 mm, erect at anthesis . Calyx constricted at base and just proximal to lobes , 2-2.5 mm; lobes deltate, ca. 1 mm, minutely ciliolate . Corolla white, 3.5-4.5 mm, lobed ca. 3/4 way to base. Filaments short; anthers 1-1.5 mm. Ovary glabrous. Style 3-4 mm. Fruiting pedicel erect, 0.8-1.4 cm. Berry orange, globose , slightly 2-lobed, 7-12 mm in diam. Seeds yellow or tan, reniform-discoid, ca. 3 × 2 mm, margins thickened. Fl. May-Jul, fr. Jun-Sep. [source]
Habit: Subshrub , Shrub
Flowers: Bloom Period: April, May, June, July, August. • Flower Color: near white, pale pink, white
Size/Age/Growth
Size: 4-6' tall.
Habitat
Typically found in the intertidal zone at the water's edge at a mean distance from sea level of 193 meters (634 feet).[3]
Biome: Coastal.
Biology
Reproduction
Duration: Perennial
Growth
Culture: Space 15-18" apart.
Soil: Minimum pH: 6.1 • Maximum pH: 8.5
Sunlight: Sun Exposure: Sun to Partial Shade.
Temperature: Cold Hardiness: 9a, 9b, 10a, 10b, 11. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Solananae
(
)
- R. Dahlgren Ex Reveal, 1992
- Order:
Solanales
(
)
- Dumortier, 1829
- Family:
Solanaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- nightshades, solanacées
- Subfamily:
Solanoideae
(
)
- Subfamily:
Solanoideae
(
- Family:
Solanaceae
(
- Order:
Solanales
(
- Superorder:
Solananae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name
Status: Accepted Name
.
Last scrutiny: Jul-2004
Similar Species
Members of the genus Solanum
ZipcodeZoo has pages for 1665 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
S. acaule (Wild Andean Potato) · S. aculeastrum (Sodaapple Nightshade) · S. adscendens (Sonoita Nightshade) · S. aethiopicum (Ethiopian Nightshade) · S. aethiopicum 'Burkina Faso' (Eggplant) · S. aethiopicum 'Gilo Black Stem' (Japanese Golden Eggs) · S. aethiopicum 'Jaxatu Soxna' (African Eggplant) · S. aethiopicum 'Jilo Tingua Verde Claro' (Indian Eggplant) · S. aethiopicum 'Keur M'bir N'dao' (African Eggplant) · S. aethiopicum 'N'drowa' (African Eggplant) · S. aethiopicum 'N'galam' (African Eggplant) · S. aethiopicum 'N'goyo' (African Eggplant) · S. aethiopicum 'Ruffled Red' (Japanese Golden Eggs) · S. aethiopicum 'Turkish Orange' (Japanese Golden Eggs) · S. americanum (Apple of Sodom) · S. anguivi (African Nightshade) · S. annuum 'Maor' (Bell Pepper) · S. atropurpureum (Potato Vine) · S. aviculare (Kangaroo Apple) · S. bahamense (Bahama Nightshade) · S. bahamense var. bahamense (Bahama Nightshade) · S. bahamense var. luxurians (Bahama Nightshade) · S. bahamense var. rugelii (Rugel's Nightshade) · S. betaceum (Tamarillo) · S. bulbocastanum (Ornamental Nightshade) · S. bulbocastanum bulbocastanum (Ornamental Nightshade) · S. burbankii (Wonderberry) · S. cajanumense (Tomate Silvestre) · S. campechiense (Redberry Nightshade) · S. candidum (Fuzzyfruit Nightshade) · S. capsicastrum (False Jerusalem Cherry) · S. capsicastrum 'Variegatum' (Variegated False Jerusalem Cherry) · S. capsicoides (Cockroach Berry) · S. cardiophyllum (Heaftleaf Horsenettle) · S. caripense (Mamoncillo) · S. carolinense (Apple of Sodom) · S. carolinense var. carolinense (Carolina Horse-Nettle) · S. carolinense var. floridanum (Carolina Horsenettle) · S. carolinense var. hirsutum (Carolina Horsenettle) · S. chacoense (Wild Andean Potato) · S. chenopodioides (Goosefoot Nightshade) · S. circinatum ramosa (Tomate De ) · S. citrullifolium (Melon-Leaf Nightshade) · S. citrullifolium var. citrullifolium (Watermelon Nightshade) · S. citrullifolium var. setigerum (Watermelon Nightshade) · S. clokeyi (Clokey's Nightshade) · S. commersonii (Commerson's Nightshade) · S. commersonii commersonii (Commerson's Wild Potato) · S. conocarpum (Marron Bacoba) · S. coronopus (Green False-Nightshade) · S. crispum 'Glasnerium' (Blue Potato Vine) · S. crispum 'Glasnevin' (Chilean Potato Tree) · S. crispum 'Variegata' (Chilean Potato Tree) · S. davisense (Davis Horsenettle) · S. demissum (Nightshade) · S. dimidiatum (Torrey Nightshade) · S. diphyllum (Amatillo) · S. donianum (Mullein Nightshade) · S. douglasii (Douglass Nightshade) · S. drymophilum (Erubia) · S. dulcamara (Bitter Nightshade) · S. dulcamara var. dulcamara (Bittersweet Nightshade) · S. dulcamara var. villosissimum (Bittersweet Nightshade) · S. dulcamara 'Variegatum' (Nightshade) · S. elaeagnifolium (Bull Nettle) · S. ellipticum (Velvet Potato Bush) · S. endopogon guianensis (Uluwukusi) · S. erianthum (Mullein Nightshade) · S. fendleri (Fendler Horsenettle) · S. fendleri var. fendleri (Fendler's Horse-Nettle) · S. fendleri var. texense (Texan Horsenettle) · S. ferox (Nightshade) · S. furcatum (Forked Nightshade) · S. gayanum (Chilean Nightshade) · S. giganteum (African Holly) · S. gilo (Gilo) · S. glaucophyllum (Waxyleaf Nightshade) · S. gracilius (Slender Nightshade) · S. granulo-leprosum (Solanum) · S. heterodoxum (Melon-Leaf Nightshade) · S. heterodoxum var. heterodoxum (Melonleaf Nightshade) · S. heterodoxum var. novomexicanum (New Mexican Nightshade) · S. heterodoxum var. setigeroides (Melonleaf Nightshade) · S. hindsianum (Hinds' Nightshade) · S. hirsutum (Carolina Horsenettle) · S. hyporhodium (Cocona) · S. incanum (Nightshade) · S. incompletum (Popolo Ku Mai) · S. indicum (Sparrow´s Brinjal (India)) · S. interius (Deadly Nightshade) · S. jamaicense (Jamaica Nightshade) · S. jamesii (Wild Potato) · S. jasminoides (Jasmine Nightshade) · S. jasminoides 'Album' (Potato Vine) · S. jasminoides 'Variegata' (Variegated Potato Vine) · S. khasianum (Nightshade) · S. lanceifolium (Lanceleaf Nightshade) · S. lanceolatum (Kangaroo Apple) · S. lasiocarpum (Nightshade) · S. lasiophyllum (Flannel Bush)
More Info
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Further Reading
- A contribution to the botany of the Isle of Pines, Cuba, based upon the specimens of plants from that island contained in the herbarium of the Carnegie Museum under date of October, 1916. By O.E. Jennings. n.p., 1917 url p. 249.
- An enumeration of Philippine flowering plants, Manila, Bureau of Printing, 1922-26. url p. 425.
- An enumeration of the vascular plants known from Surinam: together with their distribution and synonymy / by A. Pulle. Leiden: E. J. Brill, 1906. url p. 409, p. 549.
- Annals of the Carnegie Museum. [Pittsburgh]: Published by authority of the Board of Trustees of the Carnegie Institute, 1901- url p. 249.
- Biologia centrali-americana; or, Contributions to the knowledge of the fauna and flora of Mexico and Central America. London, Pub. for the editors by R. H. Porter and Dulau & co., 1879-88. url .
- Contributions from the United States National Herbarium 23 1920-1926 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 1298.
- Elements of botany: or, Outlines of the natural history of vegetables / by Benjamin Smith Barton. Philadelphia: Robert Desilver, 1827. url p. 102.
- Flora of Guatemala / Johnnie L. Gentry, Paul C. Standley. 24 1974 Chicago, Ill.: Field Museum of Natural History, 1974. url p. 115.
- Journal of ethnobiology. 11 1991 Flagstaff, Ariz.: Center for Western Studies, 1981- url p. 93.
- Novon a journal of botanical nomenclature from the Missouri Botanical Garden. 8 1998 St. Louis, MO: Missouri Botanical Garden, url p. 131.
- Occasional papers of the California Academy of Sciences. San Francisco: California Academy of Sciences, url p. 48.
- Phytologia. Bronx Park, New York, H.A. Gleason and H.N. Moldenke, url p. 37, p. 377.
- The botany of the voyage of H. M. S. Sulphur, under the command of Captain Sir Edward Belcher, during the years 1836-42. Published under the authority of the Lords Commissioners of the Admiralty /Edited and superintended by Richard Brinsley Hinds, attached to the expedition. The botanical descriptions by George Bentham. 1844 London: Smith, Elder, 1844-[46] url p. 194.
- Trees and shrubs of Mexico / By Paul C. Standley. Washington, Govt. Print. Off., 1920-1926. url p. 1298.
- Kuang Ko-zen & Lu An-ming, eds. 1978. Solanaceae. Fl. Reipubl. Popularis Sin. 67(1): 1-175.
- Kuang Ko-zen & Lu An-ming, eds. 1978. Solanaceae. Fl. Reipubl. Popularis Sin. 67(1): 1-175.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Data Sources
Accessed through GBIF Data Portal November 22, 2007:
- Fairchild Tropical Botanic Garden, Fairchild Tropical Botanic Garden Virtual Herbarium Darwin Core format
- Harvard University Herbaria, Harvard University Herbaria
- Herbarium of the University of Aarhus, The AAU Herbarium Database
- Missouri Botanical Garden, Missouri Botanical Garden
- SysTax, Herbarium Universitat Ulm
- SysTax, SysTax
- UNIBIO, IBUNAM, MEXU/Plantas Vasculares
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2653524
- Catalogue of Life Accepted Name Code: Sol-2212
- Global Biodiversity Information Facility Taxonkey: 13745045
- Globally Unique Identifier: urn:lsid:ipni.org:names:819023-1
- GRIN Nomen Number: 101505
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 30426
- U.S.D.A. Plant Symbol: SODI2
- Zipcode Zoo Species Identifier: 63549
Footnotes
- Zhi-Yun Zhang, Anmin Lu & William G. D'Arcy "Solanaceae". in Flora of China Vol. 17 Page 300. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Solanum". in Flora of China Vol. 17 Page 314. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Standard Deviation = 1,003.920 based on 50 observations. Terrestrial altitude and ocean depth information for each observation from British Oceanographic Data Centre. [back]
