Common Names in English:
Columbia Ragwort, Gaugeplant, Lamb-Tongue Ragwort, Lambs-Tongue Ragwort, Lambstongue Groundsel, Lambstongue Ragwort, Single-Stem Butterweed, Western Groundsel
The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.
The Senecioneae are a tribe of closely related genera that can be recognized most readily by the nature of the pappus and the involucral bracts or phyllaries. The phyllaries are basically in one well developed, often partially or wholly connate series of equal length that closely envelope the head . Frequently there are a few, very much smaller and mostly randomly distributed, often necrotic-tipped bracts near the base of the main series. The pappus is of fine, soft, often pure white capillary hairs . Heads may be either discoid or radiate . -- Gerald Carr.
, biennials, perennials
, or shrubs
, 5-100(-250+) cm (perennating
taprooted, fibrous-rooted, branched caudices, or suberect to creeping
, seldom branched; herbage
, often glabrescent
at flowering) . Stems single or clustered, erect
or branched) . Leaves basal and/or cauline; alternate; petiolate
or sessile (bases sometimes clasping
) ; blades
subpalmately to pinnately nerved, mostly ovate
or deltate to oblanceolate
, lanceolate, linear
, or filiform
(and most intermediate shapes
), rarely suborbiculate (sometimes palmately or pinnately lobed to 2-3-pinnatifid), ultimate
entire or denticulate
to serrate or toothed
(sometimes with relatively many callous
or teeth), faces
glabrous or hairy (usually arachnose to tomentose
, often glabrescent) . Heads (sometimes nodding
) usually radiate
(rarely quasi-disciform), usually in corymbiform
to cymiform, sometimes paniculiform
, arrays (sometimes from axils of distal leaves), sometimes borne singly. Calyculi usually of 1-8+ bractlets
(bractlets often intergrading with distal peduncular bracts, mostly 1 / 5 - 1 / 2 + times phyllaries), sometimes 0. Involucres mostly cylindric
, 5-15(-40) mm diam. Phyllaries persistent
, usually ± 5, 8, 13, or 21  in (1-) 2 series, distinct
(margins interlocking), erect (often reflexed
in fruit), mostly oblong
to lanceolate or linear, subequal
or equal, margins usually scarious
. Receptacles flat to convex
, foveolate, epaleate. Ray florets usually ± 5, 8, 13, or 21 , pistillate
, sometimes 0; corollas usually yellow, sometimes ochroleucous
or white, rarely reddish to purplish (laminae
sometimes barely surpassing
phyllaries; peripheral pistillate florets usually 0, sometimes 1-8+; corollas usually yellow, sometimes ochroleucous or white) . Disc florets (5-) 13-80+, bisexual
, fertile; corollas usually yellow, rarely ochroleucous, white, reddish, or purplish, tubes
shorter than to equaling campanulate throats, lobes
5, erect to recurved, usually ± deltate; style branches stigmatic
in 2 lines
, apices usually truncate-penicillate. Cypselae cylindric or prismatic
, usually 5-ribbed or -angled, glabrous or hairy (especially on ribs
sometimes myxogenic) ; pappi usually persistent (fragile), sometimes readily falling, of 30-80+, white to stramineous
. x = 10.
Species 1000+: nearly worldwide, mostly in warm-temperate, subtropical , and tropical regions at mid and upper elevations .
The concept of Senecio in traditional North American floristics stems from nineteenth century botanists who saw the genus as a diverse assemblage held together by similar morphologies of the heads and florets . Studies in the past two decades have shown Senecio in the broad sense to be a collection of separate lineages ; a better taxonomy is to be had by treating the lineages as genera. Some of the lineages were recognized in the past as infrageneric assemblages. A treatment of Senecio by T. M. Barkley (1978) reflected the traditional circumscription of the genus; a narrower circumscription is used here. Present concepts, plus a catalogue of genera, were presented by Barkley (1999) .The "species-groups" recognized here are given names purely as a matter of convenience; the groups and their names are intentionally given no formal taxonomic status (T. M. Barkley 1978) . Some of the groups may represent natural evolutionary alliances ; that remains to be clarified.The following taxa are not established members of the flora but are nonetheless noteworthy:Senecio brasiliensis (Sprengel) Lessing var. tripartitus (de Candolle) Baker is a South American weed of disturbed sites, introduced on the Gulf Coast near Pensacola, Florida, in 1893-1894. Its presence was discussed by J. M. Greenman (1917) and by L. J. Uttal (1982), both of whom treated it as Senecio canabinaefolius Hooker & Arnott. It is toxic to livestock; it seems not to have persisted in the flora.Senecio bicolor (Willdenow) Viviani (S. cineraria de Candolle) is one of the plants called "dusty miller" in the horticultural trade. It occasionally persists in the flora after cultivation.Species of the African genus Euryops are commonly cultivated in California and Florida and, to a lesser extent, in other warm areas of the flora. They would key here to Senecio. They are shrubs with leaves dissected or prominently toothed, phyllaries connate for the proximal third of their lengths , and yellow corollas. Apparently none persist for long after cultivation.Relatively recent collections from low-lying (50-100 m ), seasonally wet, disturbed areas in Orange and San Diego counties, California, have included semi-weedy perennial herbs or subshrubs 100-200 cm that are initially arachnose to tomentose, soon glabrescent, and have oblanceolate to linear or filiform leaves (2-7 cm), notably small heads in corymbiform arrays, ± 13 phyllaries 3-4 mm, and 7-8 ray florets with corolla laminae 2-3 mm. G. L. Nesom (pers. comm. ) has suggested that these plants are Senecio linearifolius A. Richard, a native of Australia and Tasmania.References to shapes, sizes, bases, margins, induments , etc. , of "leaves" in keys and descriptions refer to principal (largest, most conspicuous ) leaves at flowering unless otherwise indicated.
Species Senecio integerrimus
(possibly biennials), (10-) 20-70 cm (caudices
fleshy-fibrous). Herbage arachnose,
, or villous
Stems single. Leaves progressively reduced distally;
, lanceolate, linear
rounded-deltate, or suborbiculate, 6-25 × 1-6 cm, bases
tapered or truncate
to cordate, margins
entire or ± dentate
(distal leaves sessile, bractlike). Heads 6-20(-40+) in corymbiform
arrays (peduncle of terminal
often shorter than others). Calyculi
of 1-5+ linear to filiform
(seldom more than 2 mm). Phyllaries
usually ± 13 or ± 21, rarely ± 8, (4-) 5-12(-15)
usually black, sometimes green. Ray florets usually
± 8 or ± 13, sometimes 0; corolla laminae
mm (usually yellow, ochroleucous
to white in one variety). Cypselae
, sometimes hirtellous (mostly on angles
= 40, 80. [source]
The varieties of Senecio integerrimus are distinguished by morphology and geography. [source]
Flowers: Bloom Period: April, May. • Flower Color: yellow
Size: 12-18" tall.
Typically found at an altitude of 0 to 3,632 meters (0 to 11,916 feet).
Duration: Biennial , Perennial
Sunlight: Sun Exposure: Full Sun .
- Whittaker & Margulis,1978
- Haeckel, 1866
- Cavalier-Smith, 1981
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Takhtajan, 1967
- Takhtajan Ex Reveal, 1992
- Lindley, 1833
- Giseke, 1792, nom. cons., nom. alt.
- C. Linnaeus, 1753
- Groundsel, ragwort, butterweed [reputedly from Latin senex, old man or woman, alluding to the white pappus bristles resembling the white hair of an elderly person]
- Specific epithet:
- Botanical name: - Senecio integerrimus Nutt.
- Specific epithet: integerrimus - Nutt.
- Genus: Senecio () - C. Linnaeus, 1753 - Groundsel, ragwort, butterweed [reputedly from Latin senex, old man or woman, alluding to the white pappus bristles resembling the white hair of an elderly person]
- Subtribe: Senecioninae ()
- Tribe: Senecioneae ()
- Subfamily: Asteroideae ()
- Family: Compositae () - Giseke, 1792, nom. cons., nom. alt.
- Order: Asterales () - Lindley, 1833
- Superorder: Campanulanae () - Takhtajan Ex Reveal, 1992
- Subclass: Asteridae () - Takhtajan, 1967
- Subphylum: Euphyllophytina ()
- Phylum: Tracheophyta () - Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae () - Cavalier-Smith, 1981
- Kingdom: Plantae () - Haeckel, 1866 - Plants
Status: Accepted Name
Comment: Data Providers: IPNI, Tropicos. GCC LSID: urn :lsid:compositae.org:names:86AF04E3-2355-4807-AD8F-0EAFFADB5A2A
Last scrutiny: 21-Aug-09
Members of the genus Senecio
ZipcodeZoo has pages for 174 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
Myrica pennsylvanica (Northern Bayberry) · Myrica pensylvanica 'Morton' (Silver Sprite Bayberry) · S. acaulis (Senecio) · S. actinella (Flagstaff Ragwort) · S. amplectens (Alpine Groundsel) · S. amplectens holmii (Holm's Groundsel) · S. amplectens var. amplectens (Showy Alpine Ragwort) · S. amplectens var. holmii (Holm's Groundsel) · S. ampullaceus (Texas Groundsel) · S. angulatus (Climbing Groundsel) · S. anteuphorbium (Senecio) · S. aphanactis (Chaparral Ragwort) · S. aquaticus (Water Ragwort) · S. arborescens (Estrella) · S. arizonicus (Arizona Ragwort) · S. aronicoides (Rayless Ragwort) · S. articulatus (Candle Plant) · S. aschenborianus (Senecio) · S. astephanus (San Gabriel Ragwort) · S. atratus (Tall Blacktip Ragwort) · S. aurea (Golden Groundsel) · S. ballyi (Senecio) · S. bellidioides (Alpine Groundsel) · S. bicolor (Silver Ragwort) · S. bicolor cineraria (Dusty Miller) · S. bigelovii (Nodding Groundsel) · S. bigelovii Gray var. hallii Gray (Nodding Ragwort) · S. bigelovii var. bigelovii (Biglow's Ragwort) · S. bigelovii var. hallii (Hall's Ragwort) · S. blochmaniae (Dune Ragwort) · S. bolanderi var. bolanderi (Bolander's Ragwort) · S. californicus (California Ragwort) · S. cannabifolius (Aleutian Ragwort) · S. cannabinifolius (Hempleaf Ragwort) · S. cineraria 'Cirrus' (Cirrus Dusty Miller) · S. cineraria 'Silver Dust' (Dusty Miller) · S. clarkianus (Clark's Ragwort) · S. clivorum (Summer Ragwort) · S. confusus 'Sao Paulo' (Mexican Flame Vine) · S. congestus (Clustered Marsh Ragwort) · S. crassifolius (Senecio) · S. crassissimus (Vertical Leaf Senecio) · S. crassulus (Meadow Groundsel) · S. cristobalensis (Senecio) · S. cylindricus (Senecio) · S. dangarensis (Largescaled Rasbora) · S. deflersii (Pickle Plant) · S. dimorphophyllus var. dimorphophyllus (Splitleaf Groundsel) · S. dryadens (Longnose Tapirfish) · S. elegans (Purple Groundsel) · S. elmeri (Elmer's Ragwort) · S. eremophilus (Desert Groundsel) · S. eremophilus var. eremophilus (Desert Ragwort) · S. eremophilus var. kingii (Ragwood Groundsel) · S. eremophilus var. macdougalii (Macdougal's Groundsel) · S. ertterae (Ertter's Ragwort) · S. erucifolius (Hoary Ragwort) · S. erucifolius erucifolius (Hoary Groundsel) · S. eurycephalus Torr. & A.Gray var. eurycephalus Torr. & A.Gray ex A.Gray (Widehead Groundsel) · S. eurycephalus var. eurycephalus (Siskiyou Ragwort) · S. ficoides (Senecio) · S. flaccidus (Douglas Senecio) · S. flaccidus var. douglasii (Douglas' Groundsel) · S. flaccidus var. flaccidus (Threadleaf Groundsel) · S. flaccidus var. monoensis (Green Groundsel) · S. fremontii (Dwarf Mountain Ragwort) · S. fremontii var. blitoides (Dwarf Mountain Ragwort) · S. fremontii var. fremontii (Dwarf Mountain Ragwort) · S. fremontii var. inexpectatus (Dwarf Mountain Ragwort) · S. fremontii var. occidentalis (Dwarf Mountain Ragwort) · S. glastifolius (Holly-Leaved Senecio) · S. gunnii (Mountain Fireweed) · S. haworthii (Senecio) · S. herreanus (String of Beads Cactus) · S. herreianus (Gooseberry) · S. hispidulus (Hispid Fireweed) · S. hybridus (Common Ragwort) · S. hydrophiloides (Tall Groundsel) · S. hydrophilus (Alkali Marsh Groundsel) · S. hypoleucus (Pale Groundsel) · S. inaequidens (Narrow-Leaved Ragwort) · S. integerrimus (Columbia Ragwort) · S. integerrimus Nutt. var. exaltatus (Nutt.) Cronq. (Columbia Groundsel) · S. integerrimus var. exaltatus (Columbia Groundsel) · S. integerrimus var. integerrimus (Lambstongue Ragwort) · S. integerrimus var. major (Lambstongue Groundsel) · S. integerrimus var. ochroleucus (Paleyellow Ragwort) · S. integerrimus var. scribneri (Scribner's Ragwort) · S. jacobaea (Ragwort) · S. jacobaea dunensis (Ragwort) · S. jacobaea jacobaea (Ragwort) · S. jacobsenii (Trailing Jade) · S. kirkii (Tree Daisy) · S. kleinia (Mountain Grass) · S. kleiniaeformis (Spear Head) · S. kleiniiformis (Spearhead) · S. lamarckianus (Bois De Ch) · S. lemmonii (Lemmon Groundsel) · S. linearifolius (Fireweed) · S. littoralis (Woolly Ragwort)
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Accessed through GBIF Data Portal November 19, 2007:
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- Utah State University, USU-UTC Specimen Database
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- , Utah Valley State College Herbarium
- Biodiversity Heritage Library NamebankID: 2657686
- Catalogue of Life Accepted Name Code: Ast-7247
- Global Biodiversity Information Facility Taxonkey: 13748863
- Globally Unique Identifier: urn:lsid:ipni.org:names:245679-1
- GRIN Nomen Number: 409987
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 36148
- U.S.D.A. Plant Symbol: SELUO SEIN
- Zipcode Zoo Species Identifier: 62625
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- Mean = 1,211.170 meters (3,973.655 feet), Standard Deviation = 625.040 based on 938 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]