Interesting Facts
Common Names
Common Names in English:
Sweet Black-Eyed Susan, Sweet Coneflower
Description
Family Compositae
The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.
Tribe Heliantheae
The Heliantheae are a tribe of closely related genera of the sunflower family that can be readily recognized due to the association of a receptacular bract or chaff scale with each disk floret in the head . The heads usually include bisexual , actinomorphic disk florets with tubular corollas that have 4 or 5 distal lobes and also peripheral zygomorphic female or sometimes sterile florets with strap-shaped corollas that have 3 or fewer distal teeth. However, the ray flowers are sometimes absent and the heads are then discoid , containing only bisexual florets with tubular corollas. The pappus is absent or more commonly ranges from scales to stiff bristles . -- Gerald Carr.
Genus Rudbeckia
Annuals
, biennials, or perennials
, mostly 50-300 cm (mostly fibrous
rooted or rhizomatous
, sometimes taprooted). Stems 1-15+, erect
, branched distally, glabrous
or hairy
, sometimes glaucous. Leaves basal and cauline; alternate; petiolate
or sessile; blades
elliptic
, lanceolate, linear
, oblanceolate
, ovate
, or spatulate
, often pinnately lobed
to 1-2-pinnatifid, ultimate
margins
entire, dentate
, serrate, or coarsely toothed
, faces
glabrous or hairy, sometimes glaucous, sometimes gland-dotted. Heads radiate
or discoid
, borne singly or in ± corymbiform
or paniculiform
arrays. Involucres (early flowering) hemispheric
to rotate, 15-30+ mm diam. Phyllaries persistent
, 5-20 in 1-2(-3) series (narrowly triangular to lanceolate or narrowly elliptic, subequal
, herbaceous, distally or throughout). Receptacles subspheric to ovoid
, or conic to columnar
, paleate (paleae mostly tan proximally, green to maroon distally, obovate
, concave
, each ± clasping
a floret, apices acute to cuspidate
or truncate
to rounded
, abaxial
tips
glabrous or hairy, sometimes gland-dotted, resin ducts
2-3, maroon, 1 medial
and 1 near each margin; receptacles plus paleae and florets
= discs, 8-80 × 5-30 mm). Ray florets 0 or 5-25+, neuter
; corollas (spreading
to drooping
or reflexed
) usually yellow to yellow-orange or bicolor (laminae
often proximally maroon or each with a maroon splotch, distally yellow), sometimes wholly maroon (orangish red to maroon in R. graminifolia. Disc florets 50-800+, bisexual
, fertile
; corollas yellow, yellowish green, or brown-purple (often bicolor), tubes
shorter than cylindric
to funnelform
throats
, lobes
5, triangular. Cypselae (black) ± obpyramidal
and 4-angled (often minutely cross
rugose
), faces glabrous, angles
sometimes hairy; pappi 0, ± coroniform
, or of 2-8+ unequal scales
. x = 16, 18, 19.
Species 23: North America; introduced
in Europe.
The species of Rudbeckia are distributed among three major clades or lineages
. Although relationships
among the lineages are not robustly resolved, the lineages are treated here as sections
(as they have been traditionally). Rudbeckia hirta and sometimes other species of the genus are used in experimental studies relating to initiation of flowering and hairy root culture
. Most species are rich sources of phytochemicals that may offer
potential for pharmaceutical or other uses.[1]
Physical Description
Species Rudbeckia subtomentosa
Perennials
, to 200 cm (rhizomatous
, rhizomes stout). Stems
densely hirsute
(hairs
mostly antrorse
, to 0.5 mm). Leaves:
blades
ovate
to elliptic
(not lobed
), margins
denticulate
to serrate,
apices acute to obtuse
or acuminate, faces
densely hirsute and gland-dotted
(glands
fewer adaxially) ; basal 15-30 × 3-10 cm, bases
attenuate;
cauline petiolate
, ovate to elliptic, proximal
3-25 × 1-15
cm, usually 3-5-lobed, bases truncate
to cuneate or rounded
. Heads
(8-25) in loose
, corymbiform
to paniculiform
arrays. Phyllaries
to 1.5 cm (faces hairy
and ± gland-dotted). Receptacles
conic to hemispheric
; paleae 4-6 mm, apices acute, abaxial
tips
hirsute
and gland-dotted. Ray florets 10-16; laminae
(yellow to yellow-orange)
linear
to oblanceolate
, 20-40 × 5-8 mm, abaxially sparsely
hairy, abundantly gland-dotted. Discs 10-17 × 5-15 mm.
Disc florets 200-400+; corollas yellowish green on basal 1/2,
otherwise brown-purple, 3-4.2 mm; style branches ca.
1 mm, apices
acute. Cypselae 2-3.5 mm; pappi coroniform
, to ca.
0.2 mm. 2n = 38. [source]
Rudbeckia subtomentosa is often cultivated as an ornamental
.
[source]
Habit: Forb/herb
Flowers: Bloom Period: July, August, September, October. • Flower Color: gold, yellow-orange
Size/Age/Growth
Size: 36-48" tall.
Habitat
Mesic to wet prairies, stream banks, and woodland openings; 20-300 m [2].
Typically found at an altitude of 0 to 3,028 meters (0 to 9,934 feet).[3]
Biology
Reproduction
Duration: Perennial
Growth
Culture: Space 3-6" apart.
Sunlight: Sun Exposure: Full Sun .
Moisture: Drought Tolerance: High
Temperature: Cold Hardiness: 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Campanulanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Asterales
(
)
- Lindley, 1833
- Family:
Compositae
(
)
- Giseke, 1792, nom. cons., nom. alt.
- Subfamily:
Asteroideae
(
)
- Tribe:
Heliantheae
(
)
- Subtribe:
Rudbeckiinae
(
)
- Genus:
Rudbeckia
(
)
- C. Linnaeus, 1753
- Coneflower [For Olaus (Olof) Johannes Rudbeck, 1630-1702, and Olaus (Olof) Olai Rudbeck, 1660-1740, father and son, professors at Uppsala University, predecessors of Linnaeus]
- Specific epithet:
subtomentosa
- Pursh
- Botanical name: - Rudbeckia subtomentosa Pursh
- Specific epithet:
subtomentosa
- Pursh
- Genus:
Rudbeckia
(
- Subtribe:
Rudbeckiinae
(
- Tribe:
Heliantheae
(
- Subfamily:
Asteroideae
(
- Family:
Compositae
(
- Order:
Asterales
(
- Superorder:
Campanulanae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name
Status: Accepted Name
.
Comment: Data Providers: New Zealand Plant Name Database, Govaerts
World Compositae Checklist
A-G, IPNI, Tropicos. GCC LSID: urn
:lsid:compositae.org:names:249582E5-C77C-442F-A863-F4AA70260EA9
Last scrutiny: 15-Aug-09
Similar Species
Members of the genus Rudbeckia
ZipcodeZoo has pages for 84 species, subspecies, varieties, forms, and cultivars in this genus:
R. alpicola (Showy Coneflower) · R. auriculata (Eared Coneflower) · R. bicolor (Pinewoods Coneflower) · R. californica (California Coneflower) · R. californica var. californica (California Coneflower) · R. fulgida (Black-Eyed Susan) · R. fulgida var. fulgida (Eastern Coneflower) · R. fulgida var. deamii (Deam's Coneflower) · R. fulgida var. fulgida (Rachel's Eyes Black Eyed Susan) · R. fulgida var. palustris (Prairie Coneflower) · R. fulgida var. spathulata (Orange Coneflower) · R. fulgida var. speciosa (Orange Coneflower) · R. fulgida var. sullivantii (Sullivant's Coneflower) · R. fulgida var. sullivantii 'Goldsturm' (Black Eyed Susan) · R. fulgida var. umbrosa (Orange Coneflower) · R. fulgida 'Irish Eyes' (Black-Eyed Susan) · R. fulgida 'Marmalade' (Black-Eyed Susan) · R. glaucescens (Waxy Coneflower) · R. graminifolia (Grassleaf Coneflower) · R. grandiflora (Rough Coneflower) · R. grandiflora var. alismifolia (Rough Coneflower) · R. grandiflora var. grandiflora (Rough Coneflower) · R. heliopsidis (Sunfacing Coneflower) · R. hirta (Black-Eyed Susan) · R. hirta L. 'Sputnik' (Black-Eyed Susan) · R. hirta var. angustifolia (Blackeyed Susan) · R. hirta var. angustifolia (T.V. Moore) Perdue (Blackeyed Susan) · R. hirta var. floridana (Blackeyed Susan) · R. hirta var. hirta (Blackeyed Susan) · R. hirta var. pulcherrima (Blackeyed Susan) · R. hirta 'Autumn Colors' (Black-Eyed Susan) · R. hirta 'Becky Mix' (Black-Eyed Susan) · R. hirta 'Cherokee Sunset' (Black-Eyed Susan) · R. hirta 'Chim Chiminee' (Black-Eyed Susan) · R. hirta 'Chocolate Orange' (Black-Eyed Susan) · R. hirta 'Cordoba' (Cordoba Gloriosa Daisy) · R. hirta 'Goldilocks' (Black-Eyed Susan) · R. hirta 'Hot Chocolate' (Black-Eyed Susan) · R. hirta 'Indian Summer' (Black-Eyed Susan) · R. hirta 'Irish Eyes' (Irish Eyes Black-Eyed Susan) · R. hirta 'Marmalade' (Black-Eyed Susan) · R. hirta 'Maya' (Black-Eyed Susan) · R. hirta 'Moreno' (Black-Eyed Susan) · R. hirta 'Prairie Sun' (Black-Eyed Susan) · R. hirta 'Rustic Dwarfs Mix' (Black-Eyed Susan) · R. hirta 'Sonora' (Black-Eyed Susan) · R. hirta 'Toto' (Black-Eyed Susan) · R. hirta 'Toto Lemon' (Black-Eyed Susan) · R. hirta 'Toto Rustic' (Black-Eyed Susan) · R. klamathensis (Klamath Coneflower) · R. laciniata (Cut-Leaf Coneflower) · R. laciniata var. ampla (Green-Head Coneflower) · R. laciniata var. bipinnata (Cutleaf Coneflower) · R. laciniata var. digitata (Cutleaf Coneflower) · R. laciniata var. gaspereauensis (Greenhead Coneflower) · R. laciniata var. humilis (Greenhead Coneflower) · R. laciniata var. laciniata (Cutleaf Coneflower) · R. laciniata 'Goldquelle' (Cutleaf Cone Flower) · R. laciniata 'Gold Drop' (Cut-Leaf Coneflower) · R. laciniata 'Herbstsonne' (Herbstsonne Cutleaf Coneflower) · R. laciniata 'Hortensia' (Cut-Leaf Coneflower) · R. maxima (Cabbage Leaf Coneflower) · R. missouriensis (Missouri Coneflower) · R. mohrii (Mohr's Coneflower) · R. mollis (Softhair Coneflower) · R. montana (Montane Coneflower) · R. nitida (Shining Coneflower) · R. nitida var. nitida (Shiny Coneflower) · R. nitida'Herbstonne' (Autumn Glory) · R. occidentalis (Western Coneflower) · R. occidentalis var. occidentalis (Western Coneflower) · R. occidentalis 'Black Beauty' (Western Coneflower) · R. occidentalis 'Green Wizard' (Black Eyed Susan) · R. purpurea (Eastern Purple Coneflower) · R. scabrifolia (Roughleaf Coneflower) · R. speciosa 'Viette's Little Suzy' (Black-Eyed Susan) · R. subtomentosa (Sweet Black-Eyed Susan) · R. subtomentosa 'Henry Eilers' (Sweet Black-Eyed Susan) · R. texana (Rudbeckia Texana) · R. triloba (Brown-Eyed Susan) · R. triloba var. pinnatiloba (Browneyed Susan) · R. triloba var. rupestris (Browneyed Susan) · R. triloba var. triloba (Browneyed Susan) · R. 'Takao' (Coneflower)
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Further Reading
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- Aquatic and wetland plants of southwestern United States, by Donovan S. Correll and Helen B. Correll. [Washington]Environmental Protection Agency; [For sale by the Supt. of Docs., U.S. Govt. Print. Off.]1972. url p. 1651, p. 1652.
- Book of garden plans, Garden City, N. Y., Doubleday, Page & company, 1916. url .
- Bulletin of miscellaneous information /Royal Botanic Gardens, Kew. 1911 London: H.M. Stationery Office, 1900-1941. url p. 11.
- Bulletin of the Illinois State Laboratory of Natural History. Bloomington, Ill.: The Laboratory, 1876-1918. url p. 367, p. 367.
- Bulletin of the New York Botanical Garden. 1 1898 Lancaster, Pa.: Published for the Garden by the New Era Printing Co., url p. 157.
- Bulletin of the Torrey Botanical Club. 39 1912 New York: Torrey Botanical Club, 1870-1996 url p. 453.
- Catalog of hymenoptera in America north of Mexico / prepared cooperatively by specialists on the various groups of Hymenoptera under the direction of Karl V. Krombein. .. [et al.]. Washington: Smithsonian Institution Press, 1979- url , p. 2134.
- Catalogue of the flowering plants and ferns of Connecticut growing without cultivation / by Charles Burr Graves. .. [et al.]; Committee of the Connecticut Botanical Society. Hartford: Printed for the State Geological and Natural History Survey, 1910. url p. 393, p. 545.
- Erigenia: journal of the Illinois Native Plant Society. Carbondale, Ill.: The Society, 1982- url p. 15, p. 19.
- Fieldbook of Illinois wild flowers; six hundred fifty of the more common flowering plants in the state. Urbana, 1936. url p. 363.
- Flora Peoriana; the vegetation in the climate of middle Illinois. Peoria, Ill., J. W. Franks & Sons, 1887. url , , p. 51.
- Flora of Illinois, containing keys for identification of flowering plants and ferns. Notre Dame, Ind., University of Notre Dame Press, 1963. url p. 257.
- Flora of Indiana, by Charles C. Deam. Indianapolis, Wm. B. Burford printing co., contractor for state printing and binding, 1940. url p. 964, p. 967.
- Flora of the southeastern United States; being descriptions of the seed-plants, ferns and fern-allies growing naturally in North Carolina, South Carolina, Georgia, Florida, Tennessee, Alabama, Mississippi, Arkansas, Louisiana, and in Oklahom by John Kunkel Small. New York, The author, 1913. url p. 1254.
- Flower grouping in English, Scotch & Irish gardens; notes & 56 sketches in colour, by Margaret Waterfield. .. with contributions by E. V. B., S. Arnott [and others] London: E. P. Dutton & co., 1907. url , .
- Flowers and insects; lists of visitors of four hundred and fifty-three flowers, by Charles Robertson. Carlinville, Ill., n.p.1928. url p. 85.
- Fritz Bahr's commercial floriculture; a practical manual for the retail grower, by Fritz Bahr. .. with two hundred and eighty-eight illustrations. New York, The A. T. De la Mare company, inc.[c1922] url p. 514.
- Garden and forest; a journal of horticulture, landscape art and forestry. New York: The Garden and forest publishing co., 1888-97. url p. 394, p. 440.
- Hand-list of herbaceous plants cultivated in the Royal Botanic Gardens. London, Printed for H. M. Stationery Off. by Darling, 1902. url p. 969.
- Horticulture. Boston, Mass.: Horticulture Pub. Co., c1904- url p. 546.
- Illinois River Bluffs area assessment / Illinois Department of Natural Resources, Office of Scientific Research and Analysis, [and the] State Geological Survey Division. Springfield, Ill.: Illinois Dept. of Natural Resources, 1998- url p. 171, p. 184.
- International catalogue of scientific literature. LondonPublished for the International Council by the Royal Society of London1901-1920 url p. 794.
- Iowa parks. Des Moines[1920] url p. 228.
- Journal of the Horticultural Society of New York. New York City: The Society, 1906-1924. url .
- Journal of the Kentucky Academy of Science. Lexington, KY: The Academy, 1998- url p. 122, p. 122, p. 189, p. 189.
- List of Pteridophyta and Spermatophyta growing without cultivation in northeastern North America. Prepared by a Committee of the Botanical Club, American Association for the Advancement of Science. New York, 1894. url p. 333.
- List of herbaceous perennials tested in the arboretum and botanic garden: central experimental farm, Ottawa, Canada, with descriptions of flowers, and other notes by W. T. Macoun. Ottawa: Government Printing Bureau, 1908. url p. 93.
- Lists of plant types for landscape planting; the materials of planting for ornament listed according to their various uses, by Stephen F. Hamblin. Cambridge, Harvard University Press, 1923. url , , , , , p. 107, p. 142, p. 144, p. 149, p. 80, p. 96.
- Manual of the Flora of Jackson County, Missouri, By Kenneth K. MacKenzie, assisted by B. F. Bush and others. Kansas City, Mo., s.n., 1902 url p. vii.
- Manual of the flora of the northern states and Canada / by Nathaniel Lord Britton. New York: Holt, 1905. url p. 986.
- Memoirs of the Torrey Botanical Club. 5 1893 - 18 Durham, N.C.: Published for the Club by the Seeman Printery, 1889- url p. 333.
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- The Metaspermae of the Minnesota valley; a list of the higher seed-producing plants indigenous to the drainage-basin of the Minnesota river. Minneapolis[Harrison & Smith, State Printers]1892 url p. 538, p. 710, p. 711.
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- The garden month by month; describing the appearance, color, dates of bloom, height and cultivation of all desirable, hardy herbaceous perennials for the formal or wild garden with additional lists of aquatics, vines, ferns, et by Mabel Cabot Sedgwick assisted by Robert Cameron with over two hundred half-tone engravings from photographs of growing plants, and a chart in colors. New York, F.A. Stokes Co., [1907] url .
- The standard cyclopedia of horticulture; a discussion, for the amateur, and the professional and commercial grower, of the kinds, characteristics and methods of cultivation of the species of plants grown in the regions of the United States a Illustrated with colored plates, four thousand engravings in the text, and ninety-six full-page cuts. New York, Macmillan, 1919 [c1914] url p. 1470, p. 1471.
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- Transactions of the Kentucky Academy of Science. [Lexington, Ky.]Kentucky Academy of Science, 1923-1997. url p. 170, p. 77, p. 88, p. 90.
- Transactions of the. .. annual meetings of the Kansas Academy of Science. Topeka, Kan.: Kansas Pub. House, 1883-1901. url p. 84.
- Vascular plants of the Sangamon River basin; annotated checklist and ecological summary [by] Almut G. Jones and David T. Bell. [Urbana], University of Illinois at Urbana-Champaign, College of Agriculture, [1974] url p. 27, p. 6.
- Cox, P. B. and L. E. Urbatsch. 1994. A taxonomic revision of Rudbeckia subg. Macrocline (Asteraceae: Heliantheae: Rudbeckiinae). Castanea 59: 300-318.
- Perdue, R. E. Jr. 1957. Synopsis of Rudbeckia subgenus Rudbeckia. Rhodora 59: 293-299.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed March 27, 2012.
Data Sources
Accessed through GBIF Data Portal November 22, 2007:
- Missouri Botanical Garden, Missouri Botanical Garden
- USDA PLANTS, USDA PLANTS Database
- University of Alabama Biodiversity and Systematics, Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2658120
- Catalogue of Life Accepted Name Code: Ast-12045
- Global Biodiversity Information Facility Taxonkey: 13749218
- Globally Unique Identifier: urn:lsid:ipni.org:names:241771-1
- GRIN Nomen Number: 416345
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 36783
- U.S.D.A. Plant Symbol: RUSU
- Zipcode Zoo Species Identifier: 60784
Footnotes
- Lowell E. Urbatsch, Patricia B. Cox "Rudbeckia". in Flora of North America Vol. 21 Page 43, 44, 45. Oxford University Press. Online at EFloras.org. [back]
- "Rudbeckia subtomentosa". in Flora of North America Vol. 21 Page 53, 59. Oxford University Press. Online at EFloras.org. [back]
- Mean = 196.240 meters (643.832 feet), Standard Deviation = 231.030 based on 8,227 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
