Interesting Facts
Description
Family Rhamnaceae
Deciduous or evergreen
, often thorny trees
, shrubs
, woody climbers
, or lianas, rarely herbs. Leaves simple
, petiolate
, alternate or opposite, pinnately veined or 3-5-veined, entire to serrate, sometimes much reduced; stipules small, caducous
or persistent
, sometimes transformed into spines. Flowers yellowish to greenish, rarely brightly colored
, small, bisexual
or unisexual
, rarely polygamous, (4 or) 5-merous, hypogynous to epigynous
, in mostly axillary
, sessile or pedunculate
cymes, or reduced to few in fascicles. Calyx tube
patelliform
or hemispherical to tubular
, sometimes absent, at rim
with calyx, corolla, and stamens; sepals 4 or 5, valvate
in bud, triangular, erect
or ± recurved during anthesis
, adaxially often distinctly keeled
, alternate with petals. Petals 4 or 5, rarely absent, usually smaller than sepals, concave
or hooded
, rarely nearly flat, often shortly clawed. Stamens 4 or 5, antepetalous
and often ± enclosed by petals; filaments
thin, adnate
to bases
of petals; anthers
minute, versatile or not, 2(or 4) -celled, dehiscing by longitudinal
slits, usually introrse
. Disk intrastaminal
, nectariferous
, thin to ± fleshy
, entire or lobed
, glabrous
or rarely pubescent
, free
from ovary or tightly surrounding it, or adnate to calyx tube. Ovary superior to inferior, (1 or) 2-4-loculed, with 1(or 2) ovules per locule; ovules anatropous
, basal and erect; styles simple or ± deeply 3-lobed or 3-cleft. Fruit either an indehiscent, rarely explosively dehiscent
, sometimes winged
, schizocarpic capsule, or a ± fleshy drupe with 1-4 indehiscent, rarely dehiscent, pyrenes (stones
) . Seeds with thin, oily albumen, sometimes exalbuminous
; embryo large, oily, straight or rarely bent.
About 50 genera and more than 900 species: almost cosmopolitan
, mainly in subtropical
to tropical
areas; 13 genera and 137 species (82 endemic, one introduced
) in China.
Former classifications usually placed Rhamnaceae in the Rhamnales, together with Vitaceae and Leeaceae (Suessenguth in Engler & Prantl, Nat. Pflanzenfam., ed. 2, 20d. 1953), or together with Elaeagnaceae (Thorne, Bot. Rev. 58: 225-348. 1992) . Orders
such as Celastrales, Urticales, and Euphorbiales have often been considered as closely related groups. Recent analyses of DNA sequences strongly supported including the family
in the Rosales, beside the closest relatives Barbeyaceae and Dirachmaceae (see Angiosperm Phylogeny Group II, Bot. J. Linn. Soc. 141: 399-436. 2003) . Suessenguth (loc. cit.
) grouped the family into five tribes
, mainly characterized by fruit characters. Richardson et al.
(Kew Bull
. 55: 311-340. 2000; Amer. J. Bot. 87: 1309-1324. 2000) revised this tribal classification on the basis of a phylogenetic
analysis using rbcL and trnL-F sequences of the plastid genome. Now 11 tribes are recognized, of which four are represented in the Flora
area.
The bark
, leaves, and fruit of several species of Rhamnus have been used as laxatives
, notably R. cathartica and R. frangula. Diverse
Old World species of Rhamnus provide yellow and green dyes as well as drugs. Timber of Alphitonia, Colubrina, Hovenia, and Ziziphus species is used for construction, fine furniture, carving, lathework, and musical instruments. Many Ziziphus species yield edible fruit; among them, Z. jujuba (Chinese jujube) and Z. mauritiana (Indian jujube) are cultivated on a commercial
scale. Hovenia dulcis is also grown for its edible, fleshy inflorescence stalks
. Species of Hovenia, Paliurus, and Rhamnus are cultivated as ornamentals
.[1]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Rhamnanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Rosales
(
)
- Perleb, 1826
- Family:
Rhamnaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- buckthorn, nerpruns
- Tribe:
Colletieae
(
)
- Genus:
Retanilla
(
)
- (A.P. de Candolle) A.T. Brongniart, 1827
- Specific epithet:
glauca
- Phil.
- Botanical name: - Retanilla glauca Phil.
- Specific epithet:
glauca
- Phil.
- Genus:
Retanilla
(
- Tribe:
Colletieae
(
- Family:
Rhamnaceae
(
- Order:
Rosales
(
- Superorder:
Rhamnanae
(
- Subclass:
Rosidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Trevoa glauca Reiche
Notes
Publishing author : Phil.
Similar Species
Members of the genus Retanilla
ZipcodeZoo has pages for 0 species, subspecies, varieties, forms, and cultivars in this genus:
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Further Reading
- Chen Yi-ling and Chou Pan-kai. 1982. Rhamnaceae. In: Chen Yi-ling, ed., Fl. Reipubl. Popularis Sin. 48(1): i-vi, 1-169.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed February 5, 2012.
Identifiers
- Biodiversity Heritage Library NamebankID: 8679549
- Global Biodiversity Information Facility Taxonkey: 15817073
- Globally Unique Identifier: urn:lsid:ipni.org:names:718079-1
- International Plant Names Index (IPNI) ID: 218768-2
- Zipcode Zoo Species Identifier: 3673767
Footnotes
- Yilin Chen & Carsten Schirarend "Rhamnaceae". in Flora of China Vol. 12 Page 115,355. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
