Common Names
Click on the language to view common names.
Common Names in Chinese:
Gao Shan Mei Ji, Gao Shan Zao Shu He
Common Names in Dutch:
Alpenbeemdgras
Common Names in English:
Alpine Bluegrass, Alpine Meadow Grass, Bluegrass
Common Names in Estonian:
Alpi Nurmikas
Common Names in French:
Pâturin Des Alpes
Common Names in German:
Alpen-Rispengras, Romeie
Common Names in Italian:
Fienarola Delle Alpi
Common Names in Polish:
Wiechlina Alpejska, Wyklina Alpejska
Common Names in Russian:
мятлик аль пийский, Mjatlik Al' Pijskij
Description
Family Poaceae
Annual
or perennial
herbs, or tall woody bamboos
. Flowering stems (culms
) jointed
, internodes hollow or solid; branches arising singly from nodes and subtended by a leaf sheath
and 2-keeled prophyll, often fascicled in bamboos. Leaves arranged alternately in 2 ranks
, differentiated into sheath, blade
, and an adaxial
erect
appendage
at sheath/blade junction (ligule) ; leaf sheath surrounding and supporting culm-internode, split to base
or infrequently tubular
with partially or completely fused margins
, modified with reduced blade in bamboos (culm sheaths) ; leaf blades divergent, usually long, narrow and flat, but varying from inrolled
and filiform
to ovate
, veins parallel, sometimes with cross-connecting veinlets
(especially in bamboos) ; ligule membranous or a line
of hairs
. Inflorescence terminal
or axillary
, an open, contracted
, or spikelike panicle, or composed of lax
to spikelike racemes
arranged along an elongate
central axis, or digitate, paired
, or occasionally solitary; axillary inflorescences often many, subtended by spatheoles
(specialized bladeless leaf sheaths) and gathered into a leafy compound
panicle; spikelets
often aggregated into complex
clusters
in bamboos. Spikelets composed of distichous bracts arranged along a slender axis (rachilla) ; typically 2 lowest bracts (glumes
) empty, subtending
1 to many florets
; glumes often poorly differentiated from accompanying bracts in bamboos. Florets composed of 2 opposing bracts enclosing a single small flower, outer bract (lemma) clasping
the more delicate, usually 2-keeled inner bract (palea) ; base of floret often with thickened prolongation articulated with rachilla (callus) ; lemma often with apical or dorsal bristle
(awn
), glumes also sometimes awned
. Flowers bisexual
or unisexual
; lodicules (small scales
representing perianth) 2, rarely 3 or absent, 3 to many in bamboos, hyaline
or fleshy
; stamens 3 rarely 1, 2, 6, or more in some bamboos, hypogynous, filaments
capillary
, anthers
versatile; ovary 1-celled, styles (1 or) 2(rarely 3), free
or united
at base, topped by feathery stigmas, exserted from sides or apex of floret. Fruit normally a dry indehiscent caryopsis with thin pericarp firmly adherent
to seed, pericarp rarely free, fleshy in some bamboos; embryo small or large; hilum
punctate
to linear
.
About 700 genera and 11,000 species: widely distributed in all regions of the world.[1]
Genus Poa
Annuals
or perennials
. Culm
bases
infrequently swollen, or with bulbous sheath
bases; new shoots
intravaginal or extravaginal
, rarely (in China) pseudointravaginal, intravaginal but with reduced or rudimentary
lower leaf blades
and weakly differentiated prophyl. Uppermost culm leaf sheath closed from 1/20th to entire length
; ligule hyaline
, membranous or infrequently papery
; blade flat, folded, or involute
, abaxially keeled
, adaxially with 1 groove
on either side of the midvein
, apex prow-tipped. Inflorescence a terminal
panicle; branches 1-9 per node; flowers all bisexual
, or mixed bisexual and female (rarely male), with distal female flowers within spikelets
, or with partially to wholly female spikelets or inflorescences. Spikelets laterally compressed
, florets
(1-) 2-8(-10), rachilla disarticulating
above glumes
and between florets, uppermost floret vestigial; vivipary sometimes present; glumes mostly strongly keeled, unequal, or subequal
, lower glume 1- or 3-veined, upper glume 3(or 5) -veined; lemmas laterally compressed, usually distinctly keeled, 5(-7) -veined, distal margins and apex membranous, apex awnless, rarely minutely mucronate
; floret callus short, truncate
, blunt
, glabrous
or webbed
(with a dorsal tuft of woolly
hairs
), rarely with a line
of hairs around base of lemma; palea subequal or infrequently to 2/3 as long as lemma, not gaping
, keels green, distinctly separated, usually scabrid
, smooth
in Poa sect. Micrantherae, sometimes pilulose to villous
, margins
usually smooth, glabrous. Lodicules 2. Stamens 3, anthers
sometimes vestigial. Ovary glabrous. Caryopsis oblong
to fusiform
, triangular to oval
in cross
section
, sometimes grooved
, free
or adhering to the palea. 2n = 14-266. x = 7.
More than 500 species: throughout Arctic
and N and S temperate regions
and extending to most subtropical
and tropical
mountains, in habitats
such as temperate forests, mountain slopes
, grasslands, wetlands, steppes
, alpine
areas and tundra
, deserts, and around human habitation, on acidic to sub-basic or subsaline, dry to wet soils, from sea level to the upper limits
of vegetation; 81 species (14 endemic, at least one introduced
) in China.
Poa includes many species useful and important for forage
, soil stabilization, and lawns, and several widespread weeds
. Five of six recognized subgenera
are present in China. (1) Poa subg. Arctopoa: stout plants
with thick rhizomes, scabrid to ciliate
lemma margins, and glabrous calluses, found in subsaline to subalkaline wetlands. (2) Poa subg. Ochlopoa: plants with bulbous sheathed culm bases (spikelets then often viviparous), or if not bulbous then commonly quite smooth throughout, with shortly villous palea keels and no callus hairs, sometimes annuals. (3) Poa subg. Pseudopoa: slender annuals with scabrid-angled panicle branches, shortish glumes, uppermost culm sheaths closed
for 1/15-1/10 their length, glabrous calluses, and scabrid rachillas. (4) Poa subg. Poa: the largest and most diverse
subgenus
, including annuals and perennials, with or without rhizomes, but generally with the uppermost culm sheaths closed for over 1/4 their length. (5) Poa subg. Stenopoa: commonly tufted
perennials generally with the uppermost culm sheaths closed for only 1/15-1/5(-1/4) their length, with mainly extravaginal shoots, mostly without rhizomes, mostly with panicle branches that are scabrid angled from the base, and with 3-veined first glumes.
Some species have races with florets that develop into bulbils that can readily send down
roots
as soon as they drop from the inflorescence (i.e.
, they are viviparous). Viviparous spikelets often have fairly normal-looking proximal
florets. Pubescence
on the lemmas and calluses of such florets is often poorly developed relative to that in normal spikelets, or absent. Identification is easiest with plants having normal spikelets.
Hybridization and facultative
apomixis are common in some subgenera, especially Poa subg. Poa and P. subg. Stenopoa, and the vast majority of species studied are polyploid.[2]
Physical Description
Species Poa alpina
Perennials
, densely tufted
; shoots
intravaginal. Culms
erect
or obliquely
ascending
, (5-) 10-30(-45) cm tall, usually several per tuft, smooth
,
nodes often 2, 1 exserted. Leaf sheath
smooth, glabrous
, 2 or more
× as long as blade
, basal ones persistent
, investing
culm bases,
uppermost closed
for 1/4 length
; ligule white, 2-4(-5) mm, abaxially
smooth, of tillers 1-2 cm long; blade grayish green, flat or folded,
thickly papery
, withering, 3-10(-16) cm, 2-6 mm wide, surfaces glabrous,
margins
smooth or sparsely scabrid
, apex prow-tipped. Panicle loosely
contracted
to open, ovoid
to oblong
(pyramidal
at anthesis
), 2-7
× 2-3 cm, purple tinged; branches ascending to spreading
, 2
per node, rounded
, smooth or distally sparsely scabrid, longest 2(-3)
cm, divaricately rebranched with moderately crowded spikelets
in
distal 1/2. Spikelets broadly ovate
, 4-8 mm, florets
3-5(-7) ; vivipary
absent in China; rachilla internodes ca.
0.5 mm, smooth, glabrous
(rarely slightly pilulose) ; glumes
broadly ovate, membranous-papery,
subequal
, faintly 3-veined, keel arched, scabrid, surfaces smooth,
margins membranous, smooth, apex acute, lower glume 2.5-3(-4) mm,
upper glume 3.4-4.5 mm; lemmas broadly ovate, membranous-papery,
apex and margins broadly membranous, keel arched, keel villous for
2/3 of length, marginal
veins for 1/2 length, intermediate veins
indistinct, area between veins pilulose to short villous
; callus
glabrous; palea glabrous or proximally infrequently pilulose between
keels, keels scabrid, often medially pilulose to shortly villous.
Anthers
1.2-2 mm.
Fl.
and fr. Jul-Sep. 2n = 22, 28, 32, 33,
34, 35, 42, 44, 58. [source]
This species has spikelets broadly ovate, lower glumes 3-veined,
lemma proximally pubescent
between veins, callus glabrous, old sheaths
persistent and closely overlapping, anthers more than 1.2 mm, and
palea keels shortly villous, together making it quite distinct
from
other species. Gatherings from arctic
regions and European mountains
are often viviparous, but such plants
have not been recorded from
China. [source]
Habit: Graminoid
Habitat
Low arctic
to subalpine
meadows, sporadic
in taiga, slopes
, crevices
along ditch
banks, sandy places; 2400-3800 m
[3].
Typically found at an altitude of 0 to 3,037 meters (0 to 9,964 feet).[4]
Biology
Reproduction
Duration: Perennial
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Magnoliophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Flowering Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Liliopsida
(
)
- Scopoli, 1760
- Subclass:
Commelinidae
(
)
- Takhtajan, 1967
- Superorder:
Poanae
(
)
- (Small, 1903) Takhtajan, 1997 Ex Reveal & Doweld, 1999
- Superorder:
Poanae
(
- Subclass:
Commelinidae
(
- Class:
Liliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Magnoliophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Poa alpina var. vivipara L.
- Poa alpina vivipara (L.) Arcang.
- Poa vivipara (L.) Willd.
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: 15-Mar-2000
Place of publication
: Sp.
pl. 1:67. 1753
Name verified on 13-May-1992 by ARS Systematic Botanists. Last updated: 19-Apr-2000
Similar Species
Members of the genus Poa
ZipcodeZoo has pages for 1626 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
P. abbreviata (Northern Bluegrass) · P. abietina · P. abortiva · P. abyssinica · P. achrochaeta · P. acicularifolia · P. acicularis · P. acinaciphylla · P. acmocalyx · P. acrochaeta · P. acroleuca · P. acroleuca var. ryukyuensis · P. acuminata · P. acuticaulis · P. acutiflora · P. acutifolia · P. acutiglumis · P. acutissima · P. adjarica · P. adspersa · P. adusta · P. aegyptiaca · P. aemensis · P. aequatoriensis · P. aequigluma · P. aestivalis · P. affinis · P. afghanica · P. agassizensis · P. agrostidea · P. airoides · P. aitchisonii · P. akmanii · P. aksuensis · P. alascana · P. alata · P. alba · P. alberti · P. albescens · P. albida · P. alcea · P. alexeenkoi · P. alexeji · P. algida · P. almasovii · P. alopecurioides 'Woodside' · P. alopecurus · P. alopecurus alopecurus · P. alopecurus subsp · P. alpicola · P. alpigena · P. alpina (Alpine Meadow Grass) · P. alpina f. vivipara (Alpine Bluegrass) · P. alpina minor · P. alpina nodosa · P. alpina alpina · P. alpina var. bivonae · P. alpina var. brevifolia · P. alpina var. minor · P. alpina var. purpurascens · P. alpina vivipara · P. alsodes (Grove Bluegrass) · P. alta · P. altaica · P. alternans · P. altissima · P. altoperuana · P. amabilis · P. ambigua · P. amboinensis · P. amboinicia · P. ammophila · P. amoena · P. ampla · P. amplexicaulis · P. anadyrica · P. anae · P. anceps · P. anceps condensata · P. andicola · P. andina · P. andina spicata · P. andina var. major · P. androgyna · P. anfamensis · P. angusta · P. angustata · P. angustifolia · P. angustifolia var. angustiglumis · P. angustiglumis · P. ankaratrensis · P. annua (Annual Meadow-Grass) · P. annua f. purpurea · P. annua f. reptans (Creeping Bluegrass) · P. annua pilantha · P. annua reptans · P. annua rigidiuscula · P. annua var. nepalensis · P. annua var. rigidiuscula · P. annua var. stricta
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Further Reading
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Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Global Biodiversity Information Facility. Accessed November 16, 2007. http://www.gbif.org Mediated distribution data from 29 providers.
- "Poa alpina". in Flora of China Vol. 22 Page 261. Published by Missouri Botanical Garden Press. Online at EFloras.org.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 30, 2008)
Data Sources
Accessed through GBIF Data Portal November 16, 2007:
- European Environment Agency, EUNIS
- GBIF-Spain, Real Jardin Botanico
- , Vascular Plant Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2661203
- Catalogue of Life Accepted Name Code: ITS-41076
- Global Biodiversity Information Facility Taxonkey: 14376784
- Globally Unique Identifier: urn:lsid:ipni.org:names:86836-3
- GRIN Nomen Number: 28896
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 41076
- U.S.D.A. Plant Symbol: POVI16
- Zipcode Zoo Species Identifier: 57051
Footnotes
- Shou-liang Chen, De-Zhu Li, Guanghua Zhu, Zhenlan Wu, Sheng-lian Lu, Liang Liu, Zheng-ping Wang, Bi-xing Sun, Zheng-de Zhu, Nianhe Xia, Liang-zhi Jia, Zhenhua Guo, Wenli Chen, Xiang Chen, Yang Guangyao, Sylvia M. Phillips, Chris Stapleton, Robert J. Soreng, Susan G. Aiken, Nikolai N. Tzvelev, Paul M. Peterson, Stephen A. Renvoize, Marina V. Olonova & Klaus Ammann "Poaceae". in Flora of China Vol. 22. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Guanghua Zhu, Liang Liu, Robert J. Soreng & Marina V. Olonova "Poa". in Flora of China Vol. 22 Page 1, 225, 257, 312, 315. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Poa alpina". in Flora of China Vol. 22 Page 261. Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = 535.600 meters (1,757.218 feet), Standard Deviation = 384.870 based on 11,116 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
