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Common Names in Arabic:
Common Names in Chinese:
Common Names in English:
Bristlecone Pine, Ancient Pine, Great Basin Bristlecone Pine, Intermountain Bristlecone Pine, Western Bristlecone Pine
Common Names in Esperanto:
Common Names in Estonian:
Common Names in French:
Common Names in German:
Grannen Kiefer, Langlebige Kiefer
Common Names in Lithuanian:
Common Names in Polish:
Common Names in Russian:
Сосна остистая межгорная
; crown usually conic when young, often rounded
or flat-topped with age. Bark
of older stems variously furrowed
and plated, plates
. Branches usually in pseudowhorls; shoots
with long shoots and short shoots; short shoots borne in close spirals
from axils of scaly bracts and bearing fascicles of leaves (needles
) . Buds ovoid
, apex pointed
), usually resinous
. Leaves dimorphic, spirally arranged
leaves (needles) (1--) 2--5(--6) per fascicle, persisting 2--12 or more years, terete
or ± 2--3-angled and rounded on abaxial
surface, sessile, sheathed at base
by 12--15 overlapping scale leaves, these (at least firmer basal ones) persisting for life of fascicle or shed after first season
; resin canals 2 or more. Pollen cones in dense, spikelike cluster
around base of current
year's growth, mostly ovoid to cylindric-conic, tan to yellow, red, blue, or lavender. Seed cones maturing in 2(--3) years, shed early or variously persistent
, pendent to ± erect
, at maturity conic or cylindric, sessile or stalked
, shedding seed soon after maturity or variously serotinous (not opening upon maturity but much later) ; scales
persistent, woody or pliable, surface of exposed apical portion of each scale (apophysis) thickened, with umbo (exposed scale surface of young cone) represented by a scar
) or extended into a hook, spur, claw
, or prickle; bracts included
. Seeds winged
or wingless; cotyledons (3--) 6--10(--18) . x
Species ca. 100 (38 in the flora with 37 native and 1 widely naturalized ) : widespread in north temperate and north tropical (mountainous) regions, North America, Mexico, West Indies, Central America, Eurasia (including 1 crossing equator in Sumatra), n Africa, Pacific Islands in Sumatra.
In many areas Pinus is a forest dominant, either early successional and thus weedy or often longer-lived and part of climax forest. Certain southern pines, especially fire successional species, have a "grass stage," i.e. , the stem of the young seedling elongates little during the first several years and bears many long, curved leaves, the plant then reminiscent of a dense clump of grass .
Nomenclature used here, and to a very large degree the taxonomy, follows Elbert L.Little Jr. (1971), former Chief Dendrologist, United States Department of Agriculture, Forest Service. Much work is being done with problematic groups, particularly complexes in Pinus contorta, the pinyons, the bristlecone pines, and P . ponderosa and related taxa. Considerable chemotaxonomic and genetic data are available on the genus, but coverage is far from comprehensive. Therefore, the conservative approach used in this treatment emphasizes external morphology.
Pine (Pinus) has been adopted by Arkansas as the state tree. Southern pine (Pinus spp.) is the state tree of Alabama.
Species Pinus longaeva
Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded , flattened (sheared), or irregular. Bark red-brown, shallowly to deeply fissured with thick, scaly , irregular, blocky ridges . Branches contorted, pendent; twigs pale red-brown, aging gray to yellow-gray, puberulent , young branches resembling long bottlebrushes because of persistent leaves. Buds ovoid-acuminate, pale red-brown, ca. 1cm, resinous . Leaves mostly 5 per fascicle, upcurved, persisting 10--30 years, 1.5--3.5cm ´ 0.8--1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate; sheath ca. 1cm, soon forming rosette, shed early. Pollen cones cylindro-ellipsoid, 7--10mm, purple-red. Seed cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading , symmetric , lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6--9.5cm, purple, aging red-brown, nearly sessile; apophyses much thickened, sharply keeled ; umbo central, raised on low buttress , truncate to umbilicate , abruptly narrowed to slender but stiff, variable prickle 1--6mm, resin exudate pale. Seeds ellipsoid-obovoid; body 5--8mm, pale brown, mottled with dark red; wing 10--12mm. [source]
Subalpine and alpine ; 1700--3400m.
Great Basin BristleconePpine occurs in montane , subalpine , and timberline communities. It occurs in pure stands, but is also frequently codominant with Limber Pine (Pinus flexilis). Other associated species, depending on geographic location and site characteristics, include Single Leaf Pinyon (Pinus monophylla) at lower elevations , Quaking Aspen (Populus tremuloides) on mesic sites, and Engelmann Spruce (Picea engelmannii), Subalpine Fir (Abies lasiocarpa), Rocky Mountain White Fir (Abies concolor var. concolor), and Rocky Mountain Douglas-fir (Pseudotsuga menziesii var. glauca) in eastern Nevada and Utah.
The understorey in Great Basin bristlecone pine communities is typically sparse. Shrub associates include Big Sagebrush (Artemisia tridentata), Low Sagebrush (Artemisia arbuscula), Wax Currant (Ribes cereum), Curl-leaf Mountain Mahogany (Cercocarpus ledifolius) and others. Common associated herbaceous species include Prairie Junegrass (Koeleria macrantha), Bottlebrush Squirreltail (Elymus elymoides), King’s Sandwort (Arenaria kingii), and Granite Prickly Phlox (Leptodactylon pungens). In general, stands become increasingly diverse eastward through the range of the species, with a corresponding decrease in altitudinal range. Overall, plant diversity in these Bristlecone Pine communities is greater on limestone-derived soils than on quartzite-derived soils.
Great Basin Bristlecone Pine has the longest life span of any nonclonal species in the world. It is believed that the longevity of Bristlecone Pines is directly related to site adversity, with a high proportion of dead: live wood reducing respiration and water loss, thereby extending the life span of the tree . A relationship between tree age and proportion of dead stemwood suggests that the great ages of some individuals are related to their capacity to survive partial die-back while maintaining a constant ratio of photosynthesizing and non-photosynthesizing live tissue . In addition, high-elevation, arid environments are poor habitats for insects and root-decaying fungi that might otherwise reduce the life span of these ancient trees.
Great Basin Bristlecone Pine communities are highly drought-tolerant, generally found on very dry, mid- to high-elevation exposed slopes and ridges , with no evidence of Pleistocene glaciation. Slopes are typically steep, ranging from 10% to 50%. Stands are typically very open at high elevations, with a sparse understory . At lower elevations, Great Basin Bristlecone Pine is generally found in denser, mixed forests . Great Basin Bristlecone Pine is shade intolerant and cannot establish in very dense forest environments. Canopy cover may range from approximately 15-50%, with more open stands on harsher higher elevation sites containing massive multi-trunked trees, and tall upright trees with more tapered single trunks characterizing lower elevation sites with higher canopy density.
Soils are shallow lithosols, usually derived from limestone or dolomite , though occasionally sandstone or quartzite soils support Great Basin Bristlecone Pine. Great Basin Bristlecone Pine is found in arid climates with cold winters and droughty summers. Annual precipitation ranges from 300-600 mm, with temperatures as low as -18°C in January to 34°C in July.
List of Habitats:
- 1 Forest
- 1.4 Forest - Temperate [more info]
Culture: Space 20-30' apart.
Sunlight: Sun Exposure: Full Sun .
Temperature: Cold Hardiness: 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b. (map)
- Chatton, 1925
- Haeckel, 1866
- Cavalier-Smith, 1981
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Kenrick & Crane, 1997, nom. inval.
- Kenrick & Crane, 1997
- Novák ex Takht. (1967)
- Subclass: Pinidae () - Cronquist et al., 1966
- Class: Spermatopsida () - Novák ex Takht. (1967)
- Infraphylum: Radiatopses () - Kenrick & Crane, 1997
- Subphylum: Euphyllophytina () - Kenrick & Crane, 1997, nom. inval.
- Phylum: Tracheophyta () - Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae () - Cavalier-Smith, 1981
- Kingdom: Plantae () - Haeckel, 1866 - Plants
Pinus aristata Engelmann var. longaeva (D. K. Bailey) Little • Pinus aristata longaeva (D. K. Bailey) E. murray • Pinus aristata var. longaeva (D. K. Bailey) Little • Pinus balfouriana longaeva (D. K. Bailey) E. Murray
Status: Accepted Name
Comment: Conservation status: VU (B1+2e), Habit: Tree
Last scrutiny: 5-Jun-2008
In some references it has been treated as a variety of Pinus aristata (Pinus aristata Engelm. var. longaeva (D.K. Bailey) Little)..
Members of the genus Pinus
ZipcodeZoo has pages for 356 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
P. albicaulis (Whitebark Pine) · P. albicaulis 'Nana' (Whitebark Pine) · P. amamiana (Yakushima White Pine) · P. aristata (Bristlecone Pine) · P. aristata 'Sherwood Compact' (Sherwood Compact Pine) · P. arizonica (Arizona Pine) · P. arizonica Engelm. var. arizonica (Arizona Pine) · P. arizonica Engelm. var. stormiae Martinez (Arizona Pine) · P. arizonica var. stormiae (Arizona Pine) · P. armandii (Davids Pine) · P. attenuata (Knob-Cone Pine) · P. attenuatis (Knobcone Pine) · P. attenuradiata (Pine) · P. ayacahuite (Mexican White PinePinus Ayacahuite) · P. balfouriana (Foxtail Pine) · P. balfouriana austrina (Foxtail Pine) · P. balfouriana balfouriana (Foxtail Pine) · P. banksiana (Black Pine) · P. banksiana 'Chippewa' (Jack Pine) · P. banksiana 'Girards Weeper' (Jack Pine) · P. banksiana 'Manomet' (Jack Pine) · P. banksiana 'Potters' (Jack Pine) · P. bhutanica (Bhutan White Pine) · P. brutia (Afghan Pine) · P. brutia var. eldarica (Afganistan Pine) · P. brutia var. pityusa (Yel' Vostochnaya) · P. bungeana (Lace-Bark Pine) · P. bungeana 'Rowe Arboretum' (Chinese Lacebark Pine) · P. canariensis (Canary Island Pine) · P. canariensis f. brevifolia (Canary Island Pine) · P. caribaea (Caribbean Pine) · P. caribaea bahamensis (Caribbean Pine) · P. caribaea var. bahamensis (Caribbean Pine) · P. caribaea var. caribaea (Caribbean Pine) · P. caribaea var. hondurensis (Caribbean Pine) · P. cembra (Arolla Pine) · P. cembra L. 'Pygmaea' (Swiss Stone Pine) · P. cembra witches' broom (Swiss Stone Pine) · P. cembra 'Blue Mound' (Swiss Stone Pine) · P. cembra 'Chalet' (Arolla Pine) · P. cembra 'Compacta' (Swiss Stone Pine) · P. cembra 'Compacta Glauca' (Swiss Pine Or Arolla Pine) · P. cembra 'Glauca Nana' (Swiss Pine Or Arolla Pine) · P. cembra 'Klein' (Silver Whispers" Swiss Stone Pine) · P. cembra 'Nana' (Dwarf Swiss Stone Pine) · P. cembra 'Silver Sheen' (Swiss Pine Or Arolla Pine) · P. cembroides (Stoneseed Pinyon) · P. cembroides cembroides (Border Pinyon) · P. chiapensis (Chiapas White Pine) · P. clausa (Sand Pine) · P. contorta (Bolander Beach Pine) · P. contorta bolanderi (Bolander Beach Pine) · P. contorta Douglas ex Loudon var. contorta (Lodgepole Pine) · P. contorta murrayana (Sierra Lodgepole Pine) · P. contorta var. bolanderi (Bolander Beach Pine) · P. contorta var. contorta (Shore Pine) · P. contorta var. latifolia (Tall Lodgepole Pine) · P. contorta var. murrayana (Murray Lodgepole Pine) · P. contorta 'Spaan's Dwarf' (Coast Pine) · P. coulteri (Big Cone Pine) · P. cubensis (Oriente Pine) · P. culminicola (Potosi Pinyon Pine) · P. dalatensis (Vietnamese White PinePinus Dalatensis) · P. densata (Chinese Mountain Pine) · P. densiflora (Japanese Red Pine) · P. densiflora Siebold & Zucc. var. funebris (Kom.) T.N.Liou & Q.L.Wang ex Silba (Japanese Red Pine) · P. densiflora Siebold & Zucc. 'Octo-partitu' (Japanese Red Pine) · P. densiflora Siebold & Zucc. 'Tanyosho' (Japanese Red Pine) · P. densiflora 'Aurea' (Golden Japanese Red Pine) · P. densiflora 'Glitzer's Weeping' (Japanese Red Pine) · P. densiflora 'Haybud' (Japanese Red Pine Haybud') · P. densiflora 'Oculis Draconis' (Dragon's Eye Pine) · P. densiflora 'Oculus-Draconis' (Dragon Eye Pine) · P. densiflora 'Pendula' (Weeping Japanese Red Pine) · P. densiflora 'Pygmaea' (Japanese Red Pine) · P. densiflora 'Umbraculifera' (Tanyosho Pine) · P. densiflora 'Umbraculifera Compacta' (Compact Tanyosho Pine) · P. densiflora 'Vibrant' (Japanese Red Pine) · P. devoniana (Pino Escobetón) · P. discolor (Border Pinyon) · P. douglasiana (Douglas Pine) · P. durangensis (Durango Pine) · P. echinata (Arkansas Pine) · P. edulis (Colorado Pinyon) · P. edulis var. edulis (Twoneedle Pinyon) · P. elliottii (Honduras Pine) · P. elliottii Engelm. var. densa Little & K.W.Dorman (South Florida Slash Pine) · P. elliottii Engelm. var. elliottii (Honduras Pine) · P. elliottii var. densa (South Florida Slash Pine) · P. elliottii var. elliottii (Slash Pine) · P. engelmannia 'Dwarf Form' (Apache Pine) · P. engelmannii (Apache Pine) · P. faeda 'CF L3791' (Loblolly Pine) · P. fenzeliana (Hainan White PinePinus Fenzeliana) · P. flexilis (Limber Pine) · P. flexilis var. reflexa (Limber Pine) · P. flexilis 'Cesarini Blue' (Limber Pine 'cesarini Blue') · P. flexilis 'Glauca' (Limber Pine) · P. flexilis 'Glauca Pendula' (Limber Pine) · P. flexilis 'Piute' (Limber Pine 'piute')
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- 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 25.
- Annual report - Western Society of Malacologists. Pomona, Calif., Western Society of Malacologists. url p. 21.
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- California women in botany: interviews conducted by Carol Holleuffer; with introductions by Lincoln Constance, Betty Gilchrist, Peter Rowlands, and John Hunter; The Bancroft Library, Regional Oral History Office, Berkeley, Calif.: University of California, c1987. url p. 15.
- Great Basin naturalist memoirs. 1978 [Provo, Utah]Brigham Young University, 1976-1992. url p. 109, p. 148, p. 32, p. 644.
- Our living resources: a report to the nation on the distribution, abundance, and health of U.S. plants, animals, and ecosystems / [edited by], Edward T. LaRoe. .. [et al.]. Washington, DC: U.S. Dept. of the Interior, National Biological Service, 1995. url p. 515.
- Phytologia. Bronx Park, New York, H.A. Gleason and H.N. Moldenke, url p. 221, p. 222.
- The Great Basin naturalist. 38 1978 Provo, Utah: M.L. Bean Life Science Museum, Brigham Young University, 1939-1999. url , , p. 113, p. 126, p. 144, p. 178, ranne, baker, andrews & ryan, page 21, reid, page 235, p. 24, p. 261, medin, page 272, p. 35, p. 359, p. 393, hiebert & hamrick, page 487, p. 489, kay & oviatt, fig. 1, page 49, fig. 3, page 492, fig. 4, page 493, p. 494, p. 50, p. 55, p. 620, p. 626, welsh & neese, page 701, p. 702, p. 706, p. 707, p. 709.
- The World List of Threatened Trees WCMC, IUCN url p. 425.
- Bailey, D.K. 1970. Phytogeography and taxonomy of Pinus subsection Balfourianae. Ann. Missouri Bot. Gard. 57: 210--249.
- Bailey, D.K. 1987. A study of Pinus subsection Cembroides I: The single-needle pinyons of the Californias and the Great Basin. Notes Roy. Bot. Gard. Edinburgh 44: 275--310.
- Bailey, D.K. and F.G. Hawksworth. 1979. Pinyons of the Chihuahuan Desert region. Phytologia 44: 129--133.
- Critchfield, W.B. and E.L. Little Jr. 1966. Geographic Distribution of the Pines of the World. Washington. [U.S.D.A., Misc. Publ. 991.]
- Duffield, J.W. 1952. Relationships and species hybridization in the genus Pinus. Silvae Genet. 1: 93--97.
- Fowells, H.A. 1965. Silvics of Forest Trees of the United States. Washington. [Agric. Handb. 271.]
- Kurz,H. and R.K. Godfrey. 1962. Trees of Northern Florida. Gainesville.
- Little, E.L. Jr. and W.B. Critchfield. 1969. Subdivisions of the genus Pinus (pines). Washington. [U.S.D.A., Misc. Publ. 1144.]
- Mirov, N.T. 1967. The Genus Pinus. New York.
- Peattie, D.C. 1953. A Natural History of Western Trees. Boston.
- Perry, J.P.Jr. 1991. The Pines of Mexico and Central America. Portland.
- Preston, R.J. 1976. North American Trees (Exclusive of Mexico and Tropical United States), ed. 3. Ames.
- Price, R.A. 1989. The genera of Pinaceae in the southeastern United States. J. Arnold Arbor. 70: 247--305.
- Sargent, C.S. 1922. Manual of the Trees of North America (Exclusive of Mexico), ed. 2. Boston and New York. [Facsimile edition in 2 vols. 1961, reprinted 1965, New York.]
- Shaw, G.R. 1914. The Genus Pinus. Cambridge, Mass. [Publ. Arnold Arbor. 5.]
- Sudworth, G.B. 1908. Forest Trees of the Pacific Slope. Washington.
- Sudworth, G.B. 1917. The Pine Trees of the Rocky Mountain Region. Washington. [U.S.D.A. Bull. 460.]
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed March 27, 2012.
- IUCN 2012. IUCN Red List of Threatened Species. Version 2011.2. . Downloaded on January 28, 2012.
- Stritch, L., Mahalovich, M. & Nelson, K.G. 2011. Pinus longaeva. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloadedon 04February2012.
Accessed through GBIF Data Portal November 21, 2007:
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- Missouri Botanical Garden, Missouri Botanical Garden
- USDA PLANTS, USDA PLANTS Database
- Utah State University, USU-UTC Specimen Database
- Utah Valley State College
- , Utah Valley State College Herbarium
- Biodiversity Heritage Library NamebankID: 2663754
- Catalogue of Life Accepted Name Code: Con-3229
- Global Biodiversity Information Facility Taxonkey: 2481187
- Globally Unique Identifier: urn:lsid:ipni.org:names:677064-1
- GRIN Nomen Number: 406450
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 183352
- International Plant Names Index (IPNI) ID: 197012-2
- IUCN ID: 232969
- Natural Heritage Network Species Identifier: PGPIN04180
- U.S.D.A. Plant Symbol: PIARL
- Zipcode Zoo Species Identifier: 56333
- Robert Kral "Pinus". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
- "Pinus longaeva". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
- Stritch, L., Mahalovich, M. & Nelson, K.G. 2011. Pinus longaeva. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloaded on 04 February 2012. [back]