Common Names
Common Names in English:
Benguet Pine, Khasia Pine
Description
Family Pinaceae
Trees
or rarely shrubs
, evergreen
or deciduous, monoecious. Branchlets
often dimorphic
: long branchlets with clearly spirally arranged
, sometimes scalelike leaves; short branchlets often reduced to slow growing lateral
spurs bearing dense clusters
of leaves at apex. Leaves solitary or in bundles of (1 or) 2-5(-8) when basally subtended by a leaf sheath
; leaf blade
linear
or needlelike, not decurrent. Cones unisexual
. Pollen cones solitary or clustered, with numerous
spirally arranged microsporophylls
; microsporophyll with 2 microsporangia; pollen usually 2-saccate (nonsaccate in Cedrus, Larix, Pseudotsuga, and most species of Tsuga) . Seed cones erect
or pendulous, maturing in 1st, 2nd, or occasionally 3rd year, dehiscent
or occasionally indehiscent, with many spirally arranged ovulate
scales
and bracts; ovulate scales usually smaller than bracts at pollination, with 2 upright ovules adaxially, free
or only basally adnate
with bracts, maturing into seed scales. Seed scales appressed, woody or leathery, variable in shape
and size, with 2 seeds adaxially, persistent
or deciduous after cone maturity. Bracts free or adnate basally with seed scales, well developed or rudimentary
, exserted or included
. Seeds terminally winged
(except in some species of Pinus) . Cotyledons 2-18. Germination hypogeal or epigeal. 2n = 24* (almost always) .
Ten or eleven genera and ca.
235 species: N hemisphere; ten genera (two endemic) and 108 species (43 endemic, 24 introduced
) in China.
Species of the Pinaceae are among the most valuable and commercially important plants
in the world. Most species are trees, and are often excellent sources of lumber, wood
products, and resins; many are cultivated for afforestation
and as ornamentals
.[1]
Genus Pinus
Trees
or shrubs
aromatic
, evergreen
; crown usually conic when young, often rounded
or flat-topped with age. Bark
of older stems variously furrowed
and plated, plates
and/or ridges
layered
or scaly
. Branches usually in pseudowhorls; shoots
dimorphic
with long shoots and short shoots; short shoots borne in close spirals
from axils of scaly bracts and bearing fascicles of leaves (needles
) . Buds ovoid
to cylindric
, apex pointed
(blunt
), usually resinous
. Leaves dimorphic, spirally arranged
; foliage
leaves (needles) (1--) 2--5(--6) per fascicle, persisting 2--12 or more years, terete
or ± 2--3-angled and rounded on abaxial
surface, sessile, sheathed at base
by 12--15 overlapping scale leaves, these (at least firmer basal ones) persisting for life of fascicle or shed after first season
; resin canals 2 or more. Pollen cones in dense, spikelike cluster
around base of current
year's growth, mostly ovoid to cylindric-conic, tan to yellow, red, blue, or lavender. Seed cones maturing in 2(--3) years, shed early or variously persistent
, pendent to ± erect
, at maturity conic or cylindric, sessile or stalked, shedding seed soon after maturity or variously serotinous (not opening upon maturity but much later) ; scales
persistent, woody or pliable, surface of exposed apical portion of each scale (apophysis) thickened, with umbo (exposed scale surface of young cone) represented by a scar
(sometimes apiculate
) or extended into a hook, spur, claw
, or prickle; bracts included
. Seeds winged
or wingless; cotyledons (3--) 6--10(--18) . x
=12.
Species ca.
100 (38 in the flora
with 37 native
and 1 widely naturalized
) : widespread in north temperate
and north tropical
(mountainous) regions, North America, Mexico, West Indies, Central America, Eurasia
(including 1 crossing equator in Sumatra), n Africa, Pacific Islands in Sumatra.
In many areas Pinus is a forest
dominant, either early successional and thus weedy or often longer-lived and part of climax forest. Certain southern pines, especially fire successional species, have a "grass stage," i.e.
, the stem of the young seedling elongates
little during the first several years and bears many long, curved
leaves, the plant then reminiscent of a dense clump
of grass
.
Nomenclature
used here, and to a very large degree
the taxonomy, follows Elbert L.Little Jr. (1971), former Chief Dendrologist, United
States Department of Agriculture, Forest Service. Much work is being done with problematic groups, particularly complexes in Pinus contorta, the pinyons, the bristlecone pines, and P . ponderosa and related taxa. Considerable chemotaxonomic and genetic data are available on the genus, but coverage is far from comprehensive. Therefore, the conservative approach used in this treatment emphasizes external morphology.
Pine (Pinus) has been adopted by Arkansas as the state tree. Southern pine (Pinus spp.) is the state tree of Alabama.[2]
Physical Description
Habit: Evergreen .
Flowers: Flower Color: green
Size/Age/Growth
Size: over 40' tall.
Biology
Growth
Culture: Space 15-20' apart.
Sunlight: Sun Exposure: Full Sun .
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name Status: Accepted Name . Latest taxonomic scrutiny: Farjon A., 11-Jan-2006
Similar Species
Members of the genus Pinus
ZipcodeZoo has pages for 1428 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
P. abchasica · P. abies · P. abies 'Pumila Glauca' · P. acapulcensis · P. adunca · P. africana · P. ajanensis · P. albicaulis (Alpine Whitebark Pine) · P. albicaulis 'Flinck' · P. albicaulis 'Nana' (Whitebark Pine) · P. albicaulis 'No. 3' · P. albicaulis 'Noble's Dwarf' · P. alopecuroides · P. altamirani · P. altissima · P. alvordensis · P. amabilis · P. amamiana · P. amiltroni · P. andraei · P. apacheca · P. aracanensis · P. arctica · P. aristata (Rocky Mountain Bristlecone Pine) · P. aristata 'Cecilia' · P. aristata 'Grumpy' · P. aristata 'Kohout's Mini' · P. aristata 'Sherwood Compact' (Sherwood Compact Pine) · P. aristata 'So Tight' · P. arizonica (Arizona Pine) · P. arizonica Engelm. var. arizonica (Arizona Pine) · P. arizonica Engelm. var. stormiae Martinez (Arizona Pine) · P. arizonica var. arizonica · P. arizonica var. cooperi · P. arizonica var. stormiae (Arizona Pine) · P. armandi · P. armandii (Davids Pine) · P. armandii Franch. var. armandii · P. armandii 'Gold Tip' · P. armandii var. armandii · P. armandii var. dabeshanensis · P. armandii var. mastersiana · P. armata · P. astecaensis · P. atrovirens · P. attenuata (Knob-Cone Pine) · P. attenuatis (Knobcone Pine) · P. attenuradiata (Pine) · P. aucklandii · P. australis · P. austriaca · P. ayacahuite (Ayacahuite Pine) · P. ayacahuite brachyptera · P. ayacahuite Ehrenb. ex Schltdl. var. brachyptera Shaw · P. ayacahuite var. ayacahuite · P. ayacahuite var. veitchii · P. babarensis · P. baciano · P. backhousiana · P. baileyii · P. balfouriana (Foxtail Pine) · P. balfouriana austrina (Foxtailed Pine) · P. balfouriana balfouriana (Foxtailed Pine) · P. balfouriana 'Dwarf Form' · P. balfouriana subsp. austrina · P. balfouroides · P. balsamea · P. banatica · P. banksiana (Northern Scrub Pine) · P. banksiana 'Arctis' · P. banksiana 'Chippewa' (Jack Pine) · P. banksiana 'Compacta' · P. banksiana 'Girards Weeper' (Jack Pine) · P. banksiana 'H.J. Welch' · P. banksiana 'Manomet' (Jack Pine) · P. banksiana 'Neponset' · P. banksiana 'Potters' (Jack Pine) · P. banksiana 'Schneverdingen' · P. banksiana 'Schoodic' · P. banksiana 'Uncle Fogy' · P. banksiana 'Wisconsin' · P. beardsleyi · P. benthamiana · P. bessereriana · P. bhutanica · P. binato-folio · P. bothiana · P. brachyptera · P. brachyptera var. ponderosa · P. brachypthera · P. bracteata · P. brunoniana · P. brutia (Calabrian Cluster Pine) · P. brutia Ten. var. eldarica (Medw.) Nahal · P. brutia Ten. var. pityusa (Steven) Silba · P. brutia var. brutia · P. brutia var. eldarica (Afganistan Pine) · P. brutia var. pendulifolia · P. brutia var. pityusa · P. bruttia
More Info
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Further Reading
- Farjon, A. (1998a): World Checklist and Bibliography of Conifers, Royal Botanical Garden, Kew
- Farjon, A. 2001. World Checklist and Bibliograpy of Conifers. 2nd edition. The Royal Botanic Gardens, Kew.
- Cheng Wan-chün, Fu Li-kuo, Law Yu-wu, Fu Shu-hsia, Wang Wen-tsai, Chu Cheng-de, Chao Chi-son & Chen Chia-jui. 1978. Pinaceae. In: Cheng Wan-chün & Fu Li-kuo, eds., Fl. Reipubl. Popularis Sin. 7: 32-281.
- Bailey, D.K. 1970. Phytogeography and taxonomy of Pinus subsection Balfourianae. Ann. Missouri Bot. Gard. 57: 210--249.
- Bailey, D.K. 1987. A study of Pinus subsection Cembroides I: The single-needle pinyons of the Californias and the Great Basin. Notes Roy. Bot. Gard. Edinburgh 44: 275--310.
- Bailey, D.K. and F.G. Hawksworth. 1979. Pinyons of the Chihuahuan Desert region. Phytologia 44: 129--133.
- Critchfield, W.B. and E.L. Little Jr. 1966. Geographic Distribution of the Pines of the World. Washington. [U.S.D.A., Misc. Publ. 991.]
- Duffield, J.W. 1952. Relationships and species hybridization in the genus Pinus. Silvae Genet. 1: 93--97.
- Fowells, H.A. 1965. Silvics of Forest Trees of the United States. Washington. [Agric. Handb. 271.]
- Kurz,H. and R.K. Godfrey. 1962. Trees of Northern Florida. Gainesville.
- Little, E.L. Jr. and W.B. Critchfield. 1969. Subdivisions of the genus Pinus (pines). Washington. [U.S.D.A., Misc. Publ. 1144.]
- Mirov, N.T. 1967. The Genus Pinus. New York.
- Peattie, D.C. 1953. A Natural History of Western Trees. Boston.
- Perry, J.P.Jr. 1991. The Pines of Mexico and Central America. Portland.
- Preston, R.J. 1976. North American Trees (Exclusive of Mexico and Tropical United States), ed. 3. Ames.
- Price, R.A. 1989. The genera of Pinaceae in the southeastern United States. J. Arnold Arbor. 70: 247--305.
- Sargent, C.S. 1922. Manual of the Trees of North America (Exclusive of Mexico), ed. 2. Boston and New York. [Facsimile edition in 2 vols. 1961, reprinted 1965, New York.]
- Shaw, G.R. 1914. The Genus Pinus. Cambridge, Mass. [Publ. Arnold Arbor. 5.]
- Sudworth, G.B. 1908. Forest Trees of the Pacific Slope. Washington.
- Sudworth, G.B. 1917. The Pine Trees of the Rocky Mountain Region. Washington. [U.S.D.A. Bull. 460.]
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-2006. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed April 19, 2007.
- Conifer Database 2006.
- Conifer Specialist Group 1998. In IUCN 2008. 2008 IUCN Red List of Threatened Species. IUCNRedList.org. Downloaded July 19, 2008.
- Global Biodiversity Information Facility. Accessed May 25, 2007. http://www.gbif.org Mediated distribution data from provider.
Data Sources
Accessed through GBIF Data Portal November 27, 2007:
- Missouri Botanical Garden, Missouri Botanical Garden
- National Herbarium of New South Wales, NSW herbarium collection
- National Herbarium of New South Wales, Plants of Papua New Guinea
Identifiers
- Biodiversity Heritage Library NamebankID: 3497330
- Global Biodiversity Information Facility Taxonkey: 14706264
- Globally Unique Identifier: urn:lsid:ipni.org:names:677057-1
- International Plant Names Index (IPNI) ID: 677057-1
- IUCN ID: 42372
- Zipcode Zoo Species Identifier: 882001
Footnotes
- Liguo Fu, Nan Li, Thomas S. Elias & Robert R. Mill "Pinaceae". in Flora of China Vol. 4 Page 11. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Robert Kral "Pinus". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
