Description
Family Ericaceae
Plants
usually woody, or herbs, sometimes lacking chlorophyll. Leaves spiral
, sometimes decussate, margin
often toothed
. Inflorescence racemose; bracteoles paired
, basal. Flowers (4 or) 5-merous. Calyx imbricate. Corolla connate
, imbricate. Stamens 10, sometimes with spurs or awns
, dehiscing by pores
, pollen in tetrads
, rarely single. Ovary superior or inferior, placentation axile
, rarely parietal
, often many ovules per locule. Style
± as long as corolla, slender. Fruit a capsule or berry, rarely a drupe; calyx persistent
.
About 125 genera and 4000 species: widely distributed in temperate
and subarctic
regions, also at high elevations
in tropical regions
; 22 genera and 826 species (524 endemic) in China.
The Monotropoideae are here included
in the Ericaceae; previously in FRPS (56: 157 216. 1990), they were treated as the Pyrolaceae. Chiogenes, recorded from China in FRPS (57(3) : 69 71. 1991), is here included in Gaultheria. Over the last half century, the Empetraceae have usually been separated from, but closely associated with, the Ericaceae. In their ecology, leaf morphology and insertion
, rusts, embryology, stamen anatomy, etc.
, they largely agree with that family
. Molecular data place Empetrum and its relatives firmly within the Ericaceae, and in particular within the subfamily
Ericoideae, in agreement with phytochemical and palynological data, and there they are best recognized as a separate tribe
. There are distinctive features of the Empetreae that were responsible for their past familial status, e.g.
, reduced perianth with separate members
, low ovule number, enlarged stigmas, etc. However, these are likely to be derived features associated with wind pollination. See Kron et al.
(Bot. Rev. 68: 335 423. 2002) and the recent treatment of the Ericaceae by Stevens et al. (in Kubitzki, Fam. Gen. Vasc. Pl. 6: 145 194. 2004) .
Several genera and many species are ornamentals
. Some fruits of Vaccinium in N China are sweet and edible, but of no particular value to humans. Some species of Chamaedaphne, Craibiodendron, Leucothoë, Lyonia, Pieris, and Rhododendron contain more or less toxic
diterpenes, which are harmful to humans or domestic animals.Mingyuan Fang, Ruizheng Fang, Mingyou He, Linzheng Hu, Hanbi Yang, Haining Qin, Tianlu Min, David F. Chamberlain, Peter Stevens, Gary D. Wallace & Arne Anderberg "Ericaceae". in Flora of China Vol. 14 Page 242. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
Genus Pieris
Shrubs
or trees
, evergreen
[or woody vines
in E North America]. Leaves spirally arranged
or pseudoverticillate
[or in whorls of 3 in NE Asia], petiolate
; leaf blade
leathery, margin
entire
to strongly toothed
. Inflorescences terminal
or axillary
, paniculate
or racemose. Pedicels ± pubescent
and with glandular
hairs
. Flowers 5-merous. Calyx valvate
, abaxially glandular, adaxially pubescent. Corolla white, urceolate
to tubular-urceolate, shortly 5-lobed. Stamens included
; filaments
swollen at base
, straight or geniculate
; anthers
on back with a pair of spurs at anther-filament junction, dehiscing by introrse-terminal and elliptic
pores
. Ovary superior, with many ovules per locule. Stigma truncate
. Capsule loculicidal, with 5 unthickened sutures
. Seeds small.
Seven species: E Asia, Caribbean Region, E North America; three species (one endemic) in China.Fang Rhui-cheng, Peter F. Stevens "Pieris". in Flora of China Vol. 14 Page 460. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Magnoliophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Flowering Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Dilleniidae
(
)
- Takhtajan, 1967
- Superorder:
Ericanae
(
)
- Takhtajan, 1967
- Order:
Ericales
(
)
- Dumortier, 1829
- Family:
Ericaceae
(
)
- Durande, 1782, Nom. Cons.
- Heath Family
- Subfamily:
Vaccinioideae
(
)
- Tribe:
Andromedeae
(
)
- Genus:
Pieris
(
)
- D. Don
- Fetterbush
- Cultivar:
Mouwsvila
- Botanical name: - Pieris 'Mouwsvila'
- Cultivar:
Mouwsvila
- Genus:
Pieris
(
- Tribe:
Andromedeae
(
- Subfamily:
Vaccinioideae
(
- Family:
Ericaceae
(
- Order:
Ericales
(
- Superorder:
Ericanae
(
- Subclass:
Dilleniidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Magnoliophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Parentage: sport of 'Forest Flame'
An accepted name
in the RHS Horticultural Database.
Similar Species
Members of the genus Pieris
There are approximately 238 species in this genus. Here are just 100 of them:
P. 'Balls of Fire' · P. 'Bert Chandler' · P. 'Chaccone' · P. 'Firecrest' · P. 'Flaming Silver' · P. 'Forest Flame' · P. 'Mount Congreve Coral' · P. 'Mount Congreve Form' · P. 'Mount Congreve Pink' · P. 'Mouwsvila' · P. 'Spring Candy' · P. 'Tilford' · P. angelika (Arctic White) · P. annamensis · P. balcana · P. beckerii · P. bodinieri · P. bracteata · P. brassicae brassicae · P. brassicae cheiranthi · P. brassicae wollastoni · P. buxifolia · P. callidice · P. cavaleriei · P. compta · P. coreana · P. cubensis · P. daplidice · P. divaricata · P. doyonensis · P. elliptica · P. esquirolii · P. fauriei · P. floribunda (Fetterbush) · P. floribunda 'Elongata' · P. floribunda 'Forest Flame' · P. floribunda 'Millstream' · P. floribunda x japonica · P. formosa · P. formosa 'Charles Williams' · P. formosa 'Gill' · P. formosa 'Werrington' · P. formosa (Wall.) D.Don. var. forrestii (R.L.Harrow) Airy Shaw · P. formosa var. forrestii · P. formosa var. forrestii 'Ball of Fire' · P. formosa var. forrestii 'Brown Buds' · P. formosa var. forrestii 'Charles Michael' · P. formosa var. forrestii 'Fota Pink' · P. formosa var. forrestii 'Henry Price' · P. formosa var. forrestii 'Jenkinsii' · P. formosa var. forrestii 'Jermyns' · P. formosa var. forrestii 'Rowallane' · P. formosa var. forrestii 'Wakehurst' · P. formosa x japonica · P. formosana · P. forrestii · P. fortunati · P. gagnepainiana · P. Havila = 'Mouwsvila' · P. henryi · P. hieracioides · P. hybrid · P. jakusimanum · P. japonica · P. japonica 'Amamiana' (Andromeda) · P. japonica 'Astrid' · P. japonica 'Aurea' · P. japonica 'Bisbee Dwarf' (Andromeda) · P. japonica 'Blush' · P. japonica 'Boltman's Pink' · P. japonica 'Bonfire' · P. japonica 'Bonsai' · P. japonica 'Brookside Miniature' · P. japonica 'Buchanan's Dwarf' · P. japonica 'Carnaval' · P. japonica 'Christmas Cheer' (Andromeda) · P. japonica 'Coleman' · P. japonica 'Compact Crimson' · P. japonica 'Compacta' (Compact Andromeda) · P. japonica 'Crispa' · P. japonica 'Crystal' · P. japonica 'Cupido' · P. japonica 'Daisen' · P. japonica 'Daisy Hill' · P. japonica 'Debutante' · P. japonica 'Doctor Ticknor' · P. japonica 'Don' · P. japonica 'Dorothy Wyckoff' (Andromeda) · P. japonica 'Elaine' · P. japonica 'Firecrest' · P. japonica 'Flaming Silver' (Flaming Silver Andromeda) · P. japonica 'Flaming Star' · P. japonica 'Flamingo' (Andromeda) · P. japonica 'Forest Flame' (Forest Flame Japanese Andromeda) · P. japonica 'Geisha' · P. japonica 'Glenroy Pink Plenty' · P. japonica 'Grayswood' · P. japonica 'Green Pillar' · P. japonica 'Hino Crimson' · P. japonica 'Iseli'
Bibliography
- Chou Yiliang & Zhou Ruichang. 1990. Pyrolaceae. In: Fang Wenpei & Hu Wenkuang, eds., Fl. Reipubl. Popularis Sin. 56: 157216
- Fang Rhuicheng, Yang Hanbi & Chin Tsenli. 1999. Ericaceae (1). In: Fang Rhuicheng, ed., Fl. Reipubl. Popularis Sin. 57(1): 1213
- Hsu Tingzhi, Gao Baochun, Fang Rhuicheng & Huang Shuhua. 1991. Ericaceae (3). In: Fang Rhuicheng, ed., Fl. Reipubl. Popularis Sin. 57(3): 1207
- Hu Wenkuang, Hu Lincheng, Fang Mingyuan & He Mingyou. 1994. Ericaceae (2). In: Hu Lincheng, Fang Mingyuan, ed., Fl. Reipubl. Popularis Sin. 57(2): 1438
- Ming Tien lu. 1980. Empetraceae. In: Cheng Mien & Ming Tien lu, eds., Fl. Reipubl. Popularis Sin. 45(1): 6062.
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Notes
Identifiers
- Zipcode Zoo Species Identifier: 1849343
