Overview
Interesting Facts
Common Names
Common Names in English:
Coiled Lousewort, Curve-Beak Lousewort
Description
Family Orobanchaceae
Herbs annual
, biennial, or perennial
parasites, without chlorophyll. Stems unbranched or sometimes branched. Leaves scalelike, spirally or subimbricately arranged. Inflorescences racemose, spicate
, or subcapitate
, rarely 1-flowered; bract 1, usually similar to leaves; bractlets
2, adnate
to base
of calyx or pedicel. Flowers bisexual
, subsessile
or pedicelled. Calyx tubular, cupular, or campanulate
, (3 or) 4-6-lobed, 2-6-parted, 6-toothed, or spathelike, sometimes absent or of 3 free
sepals. Corolla bilabiate, usually curved
, sometimes tubular-campanulate or funnelform
with 5 subequal
lobes
; upper lip entire, emarginate
, or 2-lobed; lower lip 3-lobed. Stamens 4, didynamous
, inserted
at base of corolla tube
; filaments
slender; anthers
2-celled, dehiscing longitudinally, sometimes 1 cell
fertile
and another sterile
or reduced to spur. Pistil 2- or 3-carpellate; ovary superior; placentas 2-4 or 6(-10), parietal
or sometimes axile
at ovary base; ovules 2-4 or numerous
, anatropous
. Style long; stigma inflated
, discoid
, peltate, or 2-4-lobed. Capsule usually dehiscing loculicidally by 2 or 3(or 4) valves
. Seeds minute, testa pitted
or reticulate
.
About 15 genera and 150 species: North Temperate regions
, less represented in Africa, America, Asia, and Australia; nine genera and 42 species (nine endemic) in China.[1]
Genus Pedicularis
Herbs perennial
or annual
, rarely biennial, hemiparasitic. Leaves alternate, opposite, or whorled
, usually pinnatifid
to 1- or 2-pinnatisect, rarely entire or dentate
. Lower leaves usually long petiolate
; upper leaves often ± sessile. Inflorescences terminal
or flowers axillary
; bracts usually leaflike. Calyx tubular
to campanulate
, often ± bilabiate, usually deeply cleft
anteriorly, (2--) 5-lobed. Corolla purple, red, yellow, or white, strongly bilabiate; upper lip (galea) hooded
, enclosing anthers
, laterally compressed
, rounded
or truncate
, or terminating in teeth or in a beak
; lower lip 3-lobed, usually spreading
, external to upper lip in bud. Stamens 4, didynamous
; filaments
glabrous
or pubescent
; anthers mucronate
or not. Stigma capitate. Capsule moderately compressed
or not, loculicidal. Seeds numerous
, reticulate
or costate
.
About 600 species: best represented in the mountains of SW China and widespread in the frigid and alpine
zones of the northern hemisphere; 352 species (271 endemic) occur in China.
From the limited available data, the species of Pedicularis in China are pollinated by bumblebees gathering nectar and/or pollen on nectariferous
non-beaked species, and pollen only, by scraping or vibrating,on beaked
species including those with long corolla tubes
(Macior, L. W. 1988. Plant Species Biol. 3: 61--66.; Macior, L. W. & S. K
. Sood. 1991. Plant Species Biol. 6: 75--81.; Macior, L. W., pers. comm.
).
The rank of series has been traditionally used in Pedicularis since the time of Prain's work (Ann. Bot. Gard. (Calcutta) 3: 1--196. 1890). Sections
(and subgenera
) have also been used (e.g.
, by Bunge in Walpers, Repert. Bot. Syst. 3: 409--433. 1844; Li, H. L. Proc. Acad. Nat. Sci. Philadelphia 101: 1--214. 1949; Yamazaki, T. 1988. A revision
of the genus Pedicularis in Nepal. in: Ohba, H. & Malla, S.B. (eds.), The Himalayan Plants 1: 91--161, and others). [2]
Physical Description
Habit: Forb/herb
Biology
Reproduction
Duration: Perennial
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- (Auct.) Cavalier-Smith, 1998
- Infraphylum:
Angiospermae
(
)
- Auct.
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Lamianae
(
)
- Takhtajan, 1967
- Order:
Lamiales
(
)
- Bromhead, 1838
- Family:
Orobanchaceae
(
)
- Ventenat, 1799
- broomrape
- Tribe:
Rhinantheae
(
)
- Genus:
Pedicularis
(
)
- C. Linnaeus, 1753
- Lousewort
- Specific epithet:
contorta
- Benth.
- Botanical name: - Pedicularis contorta Benth.
- Specific epithet:
contorta
- Benth.
- Genus:
Pedicularis
(
- Tribe:
Rhinantheae
(
- Family:
Orobanchaceae
(
- Order:
Lamiales
(
- Superorder:
Lamianae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Infraphylum:
Angiospermae
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name
Status: Accepted Name
.
Last scrutiny: 15-Mar-2000
Similar Species
Members of the genus Pedicularis
ZipcodeZoo has pages for 80 species, subspecies, varieties, forms, and cultivars in this genus:
P. alopecuroides (Canadian Lousewort) · P. angustifolia (Mogollon Mountain Lousewort) · P. attollens (Attol Lousewort) · P. attollens subsp. protogyna (Little Elephantshead) · P. aurantiaca (Indian Warrior) · P. bracteosa (Bracted Lousewort) · P. bracteosa paysoniana (Payson's Lousewort) · P. bracteosa var. atrosanguinea (Bracted Lousewort) · P. bracteosa var. bracteosa (Bracted Lousewort) · P. bracteosa var. canbyi (Canby Lousewort) · P. bracteosa var. flavida (Towering Lousewort) · P. bracteosa var. latifolia (Widepetal Lousewort) · P. bracteosa var. pachyrhiza (Towering Lousewort) · P. bracteosa var. paysoniana (Payson Lousewort) · P. canadensis (Canadian Lousewort) · P. canadensis canadensis (Smoothflower Lousewort) · P. canadensis fluviatilis (Canadian Lousewort) · P. canadensis subsp. fluviatilis (Canadian Lousewort) · P. capitata (Capitate Lousewort) · P. centranthera (Dwarf Lousewort) · P. chamissonis (Chamisso Lousewort) · P. contorta (Coiled Lousewort) · P. contorta var. contorta (Coiled Lousewort) · P. contorta var. ctenophora (Coil-Beaked Lousewort) · P. contorta var. rubicunda (Coil-Beaked Lousewort) · P. corydaloides (Canadian Lousewort) · P. crenulata (Meadow Lousewort) · P. cyathophylloides (Canadian Lousewort) · P. cystopteridifolia (Fern-Leaved Lousewort) · P. densiflora (Indian Warrior) · P. densiflora aurantiaca (Indian Warrior) · P. densiflora subsp. aurantiaca (Indian Warrior) · P. dudleyi (Dudley's Lousewort) · P. flammea (Red-Tip Lousewort) · P. flammea (Scarletfruit Passionflower) · P. furbishiae (Furbish Lousewort) · P. groenlandica (Bull Elephant's-Head) · P. hirsuta (Hairy Lousewort) · P. howellii (Howell Lousewort) · P. kanei (Woolly Lousewort) · P. labradorica (Labrador Lousewort) · P. labradorica var. labradorica (Labrador Lousewort) · P. labradorica var. sulphurea (Labrador Lousewort) · P. lanceolata (Lousewort) · P. langsdorfii (Langsdorf's Lousewort) · P. langsdorfii arctica (Langsdorf's Lousewort) · P. lapponica (Lapland Lousewort) · P. macrodonta (Muskeg Lousewort) · P. oederi (Oeder's Lousewort) · P. ornithorhyncha (Bird's-Beak Lousewort) · P. palustris (European Purple Lousewort) · P. palustris palustris (European Purple Lousewort) · P. parryi (Parry Cinchweed) · P. parryi subsp. mogollonica (Mogollon Lousewort) · P. parryi subsp. purpurea (Rocky Mountain Lousewort) · P. parviflora (Small-Flower Lousewort) · P. parviflora pennellii (Smallflower Lousewort) · P. parviflora subsp. pennellii (Pennell's Lousewort) · P. pennellii pennellii (Pennell's Lousewort) · P. procera (Giant Lousewort) · P. pulchella (Mountain Lousewort) · P. racemosa alba (Leafy Lousewort) · P. racemosa alba var. alba (Sickletop Betony) · P. racemosa subsp. alba (Sickletop Lousewort) · P. rainierensis (Mount Rainier Lousewort) · P. semibarbata (Bearded Lousewort) · P. semibarbata var. charlestonensis (Charleston Lousewort) · P. semibarbata var. semibarbata (Pinewoods Lousewort) · P. sudetica (Sudetan Lousewort) · P. sudetica albolabiata (Sudetic Lousewort) · P. sudetica interioides (Sudetic Lousewort) · P. sudetica interior (Sudetic Lousewort) · P. sudetica scopulorum (Sudetic Lousewort) · P. sudetica subsp. albolabiata (Sudetic Lousewort) · P. sudetica subsp. interior (Sudetic Lousewort) · P. sudetica subsp. pacifica (Pacific Lousewort) · P. sudetica subsp. scopulorum (Sudetic Lousewort) · P. surrecta (Greenland Lousewort) · P. sylvatica (Small Lousewort) · P. verticillata (Whorled Lousewort)
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Further Reading
- ... Flora of Glacier National Park, Montana, by Paul C. Standley. Washington, Govt. Print. Off., 1921. url p. 248.
- 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 560.
- A flora of northwest America: containing brief descriptions of all the known indigenous and naturalized plants growing without cultivation north of California, west of Utah, and south of British Columbia / by Thomas Howell. Vol. 1, Phanerogamae. Portland, Or.: [s.n.], 1903. url p. 538.
- A manual of the flora of northern Idaho /Carl Epling and Joe Ewan. 1 1941 1941. url p. 27.
- An illustrated flora of the Pacific States: Washington, Oregon, and California. Stanford University, Stanford University Press, 1923-[60] url p. 804, p. 807, p. 808.
- Appendix C., vegetation/agricultural resources of the Beal Project area / prepared for Beal Mountain Mining, Inc.; prepared by Ken L. Scow, L. Dean Culwell. Helena, Mont.: Western Technology and Engineering, [1988] url .
- Botanical survey of the Goat Flat proposed research natural area, Deerlodge National Forest/ by Peter L. Achuff and Lisa S. Roe. Helena, Mont.: Montana Natural Heritage Program[1992] url p. 20.
- Budd's flora of the Canadian Prairie Provinces / [Ottawa]: Research Branch, Agriculture Canada, 1987. url p. 648, p. 649.
- Bulletin of the Torrey Botanical Club. 46 1919 New York: Torrey Botanical Club, 1870-1996 url p. 316, p. 437, p. 449.
- Conserving globally rare plants on lands administered by the Dillon Office of the Bureau of Land Management / Helena, MT: Natural Heritage Program, c2003. url .
- Contributions from the Botanical Laboratory. Philadelphia: University of Pennsylvania Press, 1892- url .
- Contributions from the Department of Botany of Columbia University. [New York, N.Y.: s.n.], 1896- url p. 293.
- Contributions from the New York Botanical Garden. New York: The Garden, 1899- url p. 27, p. 353.
- Contributions from the United States National Herbarium 3 1892-1896 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 245, p. 248, p. 409, p. 410, p. 507, p. 508, p. 511, p. 512, p. 572, p. 70.
- Contributions from the herbarium of the Geological Survey of Canada / by James M. Macoun. [Ottawa?: s.n.], 1894-1906. url p. 473.
- Contributions to Canadian botany / by Jas. M. Macoun. Ottawa: [s. n.], 1894-1906. url p. 473.
- Elementary flora of the Northwest, Cincinnati [etc.]American Book Company[c1914] url .
- Flora of southeastern Washington and adjacent Idaho / by Charles V. Piper and R. Kent Beattie. Lancaster, Pa.: New Era, 1914. url p. 229.
- Flora of southern British Columbia and Vancouver Island: with many references to Alaska and northern species / by Joseph Kaye Henry. Toronto: W.J. Gage, c1915. url p. 273.
- Flora of the Rocky Mountains and adjacent plains, Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and neighboring parts of Nebraska, South Dakota, North Dakota, and British Columbia, by P.A. Rydberg. .. New York, The author, 1922. url p. 798.
- Flora of the northwest coast, including the area west of the summit of the Cascade Mountains, from the forty-ninth parallel south to the Calapooia Mountains on the south border of Lane County, Oregon. Lancaster, Pa., Press of the New Era Printing Company, 1915. url .
- Flora of the northwest coast: including the area west of the summit of the Cascade Mountains, from the forty-ninth parallel south to the Calapooia Mountains on the south border of Lane County, Oregon / by Charles V. Piper and R. Kent Beattie. Lancaster, Pa.: Press of the New era printing company, 1915. url p. 326.
- Flora of the southeastern Washington and adjacent Idaho, Lancaster, Pa., Press of the New Era Printing Company, 1914. url .
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- Flora of the state of Washington. By Charles V. Piper. Washington, Govt. Print. Off., 1906. url p. 512.
- Great Basin naturalist memoirs. 1987 [Provo, Utah]Brigham Young University, 1976-1992. url p. 578.
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- Mountain wild flowers of America: a simple and popular guide to the names and descriptions of the flowers that bloom above the clouds / by Julia W. Henshaw. Boston: Ginn, 1906. url p. 379, p. 96, p. 97, p. xv.
- Mountain wild flowers of America; a simple and popular guide to the names and descriptions of the flowers that bloom above the clouds, by Julia W. Henshaw. Toronto, Briggs, 1906. url , p. 379, p. 379.
- Mountain wild flowers of Canada; a simple and popular guide to the names and descriptions of the flowers that bloom above the clouds, Julia W. Henshaw. Toronto, Briggs, 1906. url p. 379.
- New manual of botany of the central Rocky Mountains, vascular plants / revised by Aven Nelson. New York: American Book Co., [c1909] url p. 455.
- North American fauna. Washington: Fish and Wildlife Service; for sale by the Supt. of Docs., U. S. Govt. Print. Off. url p. 53.
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- Rare plant inventory of the outstanding natural areas of the Rocky Mountain Front / prepared for United States Department of Interior, Bureau of Land Management; prepared by Lisa Ann Schassberger. Helena, Mont.: Montana Natural Heritage Program, [1988] url p. 31.
- Report on the conservation status of Cirsium longistylum, a candidate threatened species / Lisa A. Schassberger. Helena, Mont.: Montana Natural Heritage Program, [1991]. url p. 11.
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- Sensitive plant survey in the Horse Prairie Creek drainage, Beaverhead County, Montana, Butte District, Bureau of Land Management / prepared by Jim Vanderhorst; prepared for United States Department of Interior, Bureau of Land management, Dillon Resource Area. Helena, Mont.: Montana Natural Heritage Program, c1995. url p. 42.
- Sensitive plant survey in the Tendoy Mountains, Beaverhead County, Montana / prepared by Jim Vanderhorst and Peter Lesica; prepared for Bureau of Land Management, Butte District. Helena, Mt.: Montana Natural Heritage Program, [1994]. url .
- Sensitive plant surveys in the Big Belt and Elkhorn Mountains, U.S.D.A. Forest Service, Region 1, Helena National Forest, Montana / prepared by Jackie M. Poole and Bonnie L. Heidel; prepared for Helena National Forest. Helena, Mont.: Montana Natural Heritage Program, [1993] url .
- Sensitive, threatened, and endangered vascular plants of Montana / Peter Lesica, J. Stephen Shelly. Helena, Mont.: Montana Natural Heritage Program, Montana State Library, 1991. url p. 72, p. 8, p. 87.
- Status review and taxonomic studies of Astragalus molybdenus, U.S.D.A. Forest Service, Region 1, Lewis and Clark National Forest / prepared by Lisa Ann Schassberger and J. Stephen Shelly. Helena, Mont.: Montana Natural Heritage Program, [1990] url p. 32.
- Status review of Allotropa virgata, U.S.D.A. Forest Service, Region 1, Bitterroot and Deerlodge National Forests, Montana / prepared by Lisa Schassberger Roe. Helena, Mont.: Montana Natural Heritage Program, [1992] url p. 19, p. 26, p. 27, p. 28, p. 29, p. 31, p. 37, p. 38, p. 6.
- Status review of Cirsium longistylum, U.S.D.A. Forest Service, Region 1 Lewis and Clark National Forest, Montana / Lisa A. Schassberger and Peter L. Achuff. Helena, Mont.: Montana Natural Heritage Program, [1991]. url p. 7.
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- Vascular plant and sensitive plant species inventory for the highland mountains, Deerlodge National Forest; prepared for Deerlodge National Forest / prepared by Peter Lesica. Helena, Mont.: Montana Natural Heritage Program, [1992] url p. 25, p. 47.
- Vegetation and flora of the Line Creek Plateau area, Carbon County, Montana / prepared by Peter Lesica and the Montana Natural Heritage Program; prepared for USDA Forest Service, Intermountain Research Station. Helena, MT: Montana Natural Heritage Program, [1993] url p. 28.
- Wild flowers of the North American mountains. New York, McBride, 1917. url p. 189, p. 190, p. 37, p. 373.
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Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 11, 2012.
Data Sources
Accessed through GBIF Data Portal February 02, 2008:
- Oregon State University, Vascular Plant Collection
- The New York Botanical Garden, Vascular Plant Type Specimens
- USDA PLANTS, USDA PLANTS Database
- University of Washington Burke Museum, Vascular Plant Collection - University of Washington Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2655782
- Catalogue of Life Accepted Name Code: ITS-33369
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 33369
- Natural Heritage Network Species Identifier: PDSCR1K090
- U.S.D.A. Plant Symbol: PECOR
- Zipcode Zoo Species Identifier: 53978
Footnotes
- Zhi-Yun Zhang & Nikolai N. Tzvelev "Orobanchaceae". in Flora of China Vol. 18 Page 229. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Yang Han-bi, Noel H. Holmgren, Robert R. Mill "Pedicularis". in Flora of China Vol. 18 Page 97. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
