Common Names
Common Names in English:
Palmleaf Rockcap Fern
Description
Family Polypodiaceae
Plants
perennial
, terrestrial
, on rock, or often epiphytic, erect
, arching
, or occasionally pendent. Stems long- to short-creeping, branched or not, bearing scales
and few to numerous
roots
, usually dictyostelic. Leaves monomorphic
to dimorphic
, circinate
in bud. Petiole
usually articulate
at base
[rarely nonarticulate
, as in Loxogramme ], lacking scales or sometimes scaly
, with usually 3 vascular bundles
. Blade simple
to often pinnatifid
, pinnatisect
, or pinnate, infrequently more divided
; rachis grooved
or not adaxially. Veins free
(and simple to several times forked
) to often anastomosing in complex
systems
, areoles with or without included
veinlets
. Indument on blade
absent, or petiole, rachis, costae, and sometimes blade tissue usually bearing hairs
(these often septate
and with reddish crosswalls) and/or scales. Sori borne abaxially on veins, round to oblong
, occasionally elongate
, rarely marginal
, rarely covering surface; paraphyses present or absent; sporangia with stalk
of 2 or 3 rows
of cells
; indusia absent. Spores usually transparent or yellowish (rarely greenish), all 1 kind, bilateral
, monolete [rarely trilete, as in some Loxogramme], surface most often smooth
, tuberculate
, verrucose
, or granulate
, occasionally spiny
, 64 per sporangium (spores globose
and 32 per sporangium in apogamous spp.
) . Gametophytes green, aboveground, cordate or elliptic
, glabrous
or sometimes glandular
; archegonia and antheridia borne on lower surface, antheridia 3-celled.
Genera ca.
40, species perhaps 500 (7 genera, 25 species in the flora
) : worldwide, especially tropics and subtropics.
Genera in this family
are variously circumscribed, and the New World species historically were placed in the single genus Polypodium . Many of the segregates
recognized here are still placed in Polypodium in recent floristic accounts. Limits
of genera in both Old World and New World are controversial and are currently under study by several workers.[1]
Genus Pecluma
Plants
terrestrial
, on rock, or epiphytic. Stems short-creeping, unbranched, 4--8 mm diam., not whitish pruinose
; scales
concolored, ovate
to linear-lanceolate, not clathrate, glabrous
or pubescent
, margins
entire. Leaves monomorphic
, crowded, narrowed toward tip
, to 90 cm. Petiole
dark brown to black, round in cross
section
except for narrow lateral
ridge
decurrent from base
of blade
. Blade narrowly oblong
to linear
, usually pectinate
, with more than (20--) 25 pairs of segments, not glaucous, pubescent, scales absent or on midrib
only; rachis scaly
and/or pubescent or glabrous abaxially, pubescent adaxially, scales basally attached and linear to cordate. Segments usually linear, 1.5--5(--8) mm wide, adnate
to costa, closely spaced, margins entire, apex acute. Veins 1--2-forked [simple
], occasionally anastomosing. Sori terminal
on veins, round; indument
of branched glandular
hairs
. Spores tuberculate
. x
= 37.
Species ca.
30: North America, Mexico, West Indies, Central America, South America.
Within Polypodiaceae, Pecluma is distinctive in its short-creeping stems, pectinate blades (usually with more than 30 pairs of segments), and ungrooved, adaxially pubescent rachises.
Many species of Pecluma are well adapted to dry seasons
. As the leaves dry out, the segments curl inwardly, presumably retarding moisture loss. After rains, the segments uncurl, apparently undamaged by the period of dryness.[2]
Physical Description
Habitat
Typically found at an altitude of 0 to 4,936 meters (0 to 16,194 feet).[3]
Biology
Reproduction
Duration: Perennial
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Moniliformopses
(
)
- Kenrick & Crane, 1997, Nom. Nud.
- Class:
Filicopsida
(
)
- Cronquist Et Al.
- Order:
Polypodiales
(
)
- Link
- Family:
Polypodiaceae
(
)
- Berchtold & J. Presl, 1820
- Polypody Family
- Genus:
Pecluma
(
)
- M. G. Price, Amer. Fern J. 73: 109. 1983.
- [Latin pectinatus, in the form of a comb, and plumula, feathery, for the leaf blades]
- Specific epithet:
ptilodon
- (Kunze) M. G. Price, Amer. Fern J. 73: 115. 1983.
- Botanical name: - Pecluma ptilodon (Kunze) M.G. Price
- Specific epithet:
ptilodon
- (Kunze) M. G. Price, Amer. Fern J. 73: 115. 1983.
- Genus:
Pecluma
(
- Family:
Polypodiaceae
(
- Order:
Polypodiales
(
- Class:
Filicopsida
(
- Infraphylum:
Moniliformopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Polypodium ptilodon Kunze, Linnaea 9: 42. 1834
Notes
Publishing author : Lellinger Publication : Amer. Fern J. 77: 101 1987 Basionym author: (Jenman) Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000
Similar Species
Members of the genus Pecluma
ZipcodeZoo has pages for 65 species, subspecies, varieties, forms, and cultivars in this genus:
P. absidata · P. alfredii · P. atra · P. bermudiana · P. boliviana · P. bolivianum · P. camptophyllaria (Colombian Rockcap Fern) · P. camptophyllaria abbreviata (Colombian Rockcap Fern) · P. camptophyllaria camptophyllaria (Colombian Rockcap Fern) · P. camptophyllaria subsp. abbreviata (Colombian Rockcap Fern) · P. camptophyllaria var. abbreviata · P. camptophyllaria var. lachnifera · P. camptophyllaria var. macedoi · P. chiapensis · P. choquetanensis · P. choquetangense · P. choquetangensis · P. consimile · P. consimilis (Montane Rockcap Fern) · P. cupreolepis · P. curvans · P. dispersa (Widespread Rockcap Fern) · P. divaricata · P. eurybasis · P. eurybasis var. glabrescens · P. eurybasis var. villosa · P. eurybasis var. villosum · P. ferruginea · P. filicula · P. funicula · P. hoehnei · P. hoehnii · P. hygometrica · P. hygrometrica · P. hygrometricum · P. imbeana · P. insularis · P. macedoi · P. oranense · P. oranensis · P. paradieae · P. paradiseae · P. pastazensis · P. pectinata (Combleaf Rockcap Fern) · P. pectinatiforme · P. pectinatiformis · P. perpinnata · P. pilosa · P. plumula (Plumed Rockcap Fern) · P. ptilodon (Palmleaf Rockcap Fern) · P. ptilodon caespitosum (Palmleaf Rockcap Fern) · P. ptilodon ptilodon (Palmleaf Rockcap Fern) · P. ptilodon var. caespitosa · P. ptilodon var. pilosa · P. ptilodon var. robusta · P. ptilodon var. robustum · P. recurvata · P. reducida · P. robusta · P. schkuhrii · P. sicca · P. singeri · P. sursumcurrens · P. truncorum · P. venturii
More Info
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Further Reading
- Evans, A. M. 1969. Interspecific relationships in the Polypodium pectinatum-plumula complex. Ann. Missouri Bot. Gard. 55: 193--293.
- Price, M. G. 1983. Pecluma, a new tropical American fern genus. Amer. Fern J. 73: 109--116.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-2005. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. [http://sn2000.taxonomy.nl/Taxonomicon/]. Access date: Nov 23, 2005
- Global Biodiversity Information Facility. Accessed February 28, 2008. http://www.gbif.org Mediated distribution data from 6 providers.
- "Pecluma ptilodon". in Flora of North America Vol. 2. Published by Oxford University Press. Online at EFloras.org.
- The International Plant Names Index. Accessed Jan 19, 2007.
- USDA, NRCS. 2005. The PLANTS Database, Version 3.5 (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA.
Data Sources
Accessed through GBIF Data Portal February 28, 2008:
- Australian National Herbarium (CANB)
- GBIF-Spain: CUBA:Herbario del JardĂn Botánico Nacional, La Habana, Cuba: HAJB-Pteridophyta
- Herbarium of the University of Aarhus: The AAU Herbarium Database
- Instituto Nacional de Biodiversidad (Costa Rica): Biodiversidad de Costa Rica
- Missouri Botanical Garden
- USDA PLANTS: USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2668447
- Catalogue of Life Accepted Name Code: ITS-504155
- Globally Unique Identifier: urn:lsid:ipni.org:names:17514900-1
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 504155
- International Plant Names Index (IPNI) ID: 186188-2
- Natural Heritage Network Species Identifier: PPPOL06050
- U.S.D.A. Plant Symbol: PEPT2
- Zipcode Zoo Species Identifier: 49813
Footnotes
- Alan R. Smith "Polypodiaceae". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
- A. Murray Evans "Pecluma". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
- Mean = 538.790 meters (1,767.684 feet), Standard Deviation = 973.440 based on 78 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
