Overview
Clusters of female flowers develop into the familiar dark purplish to black mulberries that were a fairly dependable source of sustenance for pioneers in western North America.
Interesting Facts
- Mulberry leaves provide the natural food for silkworms. Commercially cultivated mulberries are varieties of Morus alba; they are prized as shade trees with edible fruits. [source]
- Native Americans used infusions made from the bark of Morus alba medicinally in various ways: as a laxative , as a treatment for dysentary, and as a purgative (D. E. Moerman 1986. [source]
Common Names
Click on the language to view common names.
Common Names in Afrikaans:
Gewone Moerbei, Witmoerbei
Common Names in Arabic:
El Ttuut, Tuth
Common Names in Bengali:
Tut
Common Names in Bulgarian:
Chernitsia Biala
Common Names in Burmese:
Posa
Common Names in Chinese:
Bai Sang, Sang, Sang Shu
Common Names in Danish:
Morbær (Fruit), Morbær (Plant)
Common Names in Dutch:
Moerbei (Fruit), Moerbezie (Plant), Witte Moerbeiboom
Common Names in English:
Common Mulberry, Black-Fruited Mulberry, Chinese White Mulberry, Fruitless White Mulberry, Mulberry, Mulberry Bush, Mulberry Tree, Russian Mulberry, Silkworm Mulberry, Silkworm Tree, White Mulberry, White Mulberry Tree, White-Fruited Mulberry
Common Names in French:
Mûre De Murier (Fruit), Mûrier (Plant), Mûrier Blanc, Moral Blanco
Common Names in German:
Maulbeerbaum (Plant), Maulbeere (Fruit), Weiße Maulbeere, Weißer Maulbeerbaum
Common Names in Greek:
Aspri Moria, Aspromuria
Common Names in Gujarati:
Shetun
Common Names in Hindi:
Shahtut, Tut, टूट
Common Names in Italian:
Gelso (Plant), Gelso Bianco, Gelso Comune, Mora Di Gelso (Fruit), Moral Blanco, Morera Blanco, Moro (Fruit), Moro Bianco, Moro Da Carta, Morus
Common Names in Japanese:
Guwa, Kara Guwa, Kara Yama Guwa, Kuwa, Ma Guwa
Common Names in Korean:
Ppong, Ppong Na Mu
Common Names in Malay:
Bebesaran (Indonesia), Bebesaran Lampung (Indonesia, Java), Murbei (Indonesia)
Common Names in Nepalese:
Kimbu
Common Names in Polish:
Morwa Biala
Common Names in Portuguese:
Amora Da Amoreira (Fruit), Amoreira (Plant), Amoreira Branca, Amoreira-Branca
Common Names in Romanian:
Dud Alb
Common Names in Russian:
šelkovica Belaja, Shelkovitsa Belaia, Tut Belyi, шелковица белая
Common Names in Sanskrit:
Tula
Common Names in Spanish:
Mora, Mora (Fruit), Mora Blanca, Mora De árbol, Moral, Moral Blanco, Morera (Plant), Morera Blanca
Common Names in Swahili:
Mforsadi, Mfurusadi
Common Names in Swedish:
Vitt Mullbär
Common Names in Tagalog:
Amingit, Amoras
Common Names in Tamil:
Kambli Chedi, Mussuketi, Musukette
Common Names in Thai:
Mon
Common Names in unspecified:
Mulberry, White Mulberry
Common Names in Vietnamese:
Dâu-Tàm, D[aa]u, D[awf]m Tang, T[awf]m Tang
Description
Family Moraceae
Trees
, shrubs
, vines
, or rarely herbs, frequently with milky
or watery latex, sometimes spiny
. Stipules present, frequently caducous
. Leaves alternate, rarely opposite; petiole
often present and well-defined; leaf blade
simple
, sometimes with cystoliths
, margin
entire or palmately lobed
, venation
pinnate or palmate. Inflorescences axillary
, frequently paired
, racemose, spicate
, capitate, or rarely cymose
, sometimes a fig or syconium with flowers completely enclosed within a hollow receptacle. Flowers unisexual
(plants
monoecious or dioecious), small to very small. Calyx lobes
(1 or) 2-4(-8), free
or connate
, imbricate or valvate
. Corolla absent. Male flowers: stamens as many as and opposite to calyx lobes (except in Artocarpus), straight or inflexed
in bud; anthers
1- or 2-loculed, crescent-shaped to top-shaped; pistillode
(rudimentary
sterile
pistil) often present. Female flowers: calyx lobes usually 4; ovary superior, semi-inferior, or inferior, 1(or 2) -loculed; ovules 1 per locule, anatropous
or campylotropous; style branches 1 or 2; stigmas usually filiform
. Fruit usually a drupe, rarely an achene, enveloped by an enlarged calyx and/or immersed
in a fleshy
receptacle, often joined into a syncarp. Seed solitary; endosperm present or absent.
Between 37 and 43 genera and 1100â1400 species: widespread in tropical
and subtropical
areas, less common in temperate
areas; nine genera and 144 species (26 endemic, five introduced
) in China.
Economically, the most important species are those of Morus and Maclura associated with the production
of silk
. Some species in Broussonetia, Maclura, and Morus are important for paper making; some species in Artocarpus, Ficus, and Morus have edible fruit; and some species of Artocarpus and Broussonetia are used for furniture or timber.[1]
Genus Morus
Trees
or shrubs
, deciduous, with latex; monoecious or dioecious. Winter buds
with 3-6 bud scales; scales
imbricate. Stipules free
, sublateral, caducous
. Leaves alternate; leaf blade
simple
to deeply palmately lobed
, margin
toothed
; primary
veins 3-5 from base
, secondary veins pinnate. Male inflorescences axillary
, spicate
, many-flowered, shortly pedunculate
. Female inflorescences shortly spicate to capitate. Male flowers: calyx lobes
4, imbricate; stamens inflexed
in bud; pistillode
top-shaped. Female flowers: sessile; calyx lobes 4, imbricate, fleshy
in fruit; ovary 1-loculed; style present or not; stigma 2-branched, abaxially pubescent
or papillose
. Fruit with enlarged, succulent calyx usually aggregated into juicy syncarp. Syncarp with achenes enclosed by enlarged and succulent calyx; endocarp shell-like; exocarp
fleshy. Seed globose
; endosperm fleshy; embryo incurved
; cotyledon elliptic
.
About 16 species: widespread in all temperate
areas, also in the mountains of tropical
Africa, Indonesia, and South America; 11 species (five endemic, one introduced
) in China.
Morus species are cultivated in many temperate and tropical countries. The leaves are used as food for silkworms and the fruit for human consumption
. Male material
is often very difficult to name
.[2]
Physical Description
Species Morus alba
Shrubs
or trees
, to 15 m.
Bark
brown tinged with red or yellow, thin,
shallowly furrowed
, with long, narrow ridges
. Branchlets
orange-brown
or dark green with reddish cast, pubescent
or occasionally glabrous
;
lenticels
reddish brown, elliptic
, prominent
. Buds ovoid
, 4-6 mm,
apex acute to rounded
; outer scales
yellow-brown with dark margins
,
glabrous or with a few marginal
trichomes
; leaf scars
half round
,
bundle scars
numerous
, in circle. Leaves: stipules ovate
to lanceolate,
5-9 mm, pubescent; petiole
2.5-5 cm, short-pubescent. Leaf blade
ovate, often deeply and irregularly lobed
, (6-) 8-10 × 3-6
cm, base
cuneate, truncate
, or cordate, margins coarsely serrate
to crenate
, apex acute to short-acuminate; surfaces abaxially glabrous
or sparingly pubescent along major veins or in tufts in axils of
principal lateral
veins and midribs
, adaxially glabrous to sparsely
pubescent. Catkins: peduncle and axis pubescent; staminate
catkins
2.5-4 cm; pistillate
catkins 5-8 mm.
Flowers: staminate and pistillate
on same or different plants
. Staminate flowers
: sepals distinct
,
green with red tip
, ca.
1.5 mm, pubescent; filaments
ca. 2.7 mm.
Pistillate flowers: ovary green, ovoid, slightly compressed
, ca.
2 mm, glabrous; style branches divergent, red-brown, 0.5-1 mm; stigma
papillose
. Syncarps red when immature
, becoming black, purple, or
nearly white, cylindric
, 1.5-2.5 × 1 cm; achenes light brown,
ovoid, 2-3 mm. [source]
Morus alba and M. rubra are both highly variable and are often confused.
Both species have deeply lobed to entire leaves and are variable
in pubescence
. Some individuals are intermediate in leaf pubescence,
suggesting the possibility of hybridization. [source]
ID Features: Alternate leaf arrangement. Variable leaf shape with serrate leaf margins. Fleshy drupe fruit. Imbricate buds with fine hairs or scale tips. Terminal bud absent. Rough leaves. Bark sweet when chewed.
Habit: A dense, rounded deciduous, medium to large tree .
Flowers: Fleshy white to red fruit. Drupe. Up to 1" long. Ripens in June to July. Edible. • Bloom Period: February, March, April, May, June, July. • Flower Color: Green • Flower Conspicuous: Tiny cup-shaped flowers in clusters
Seeds: Fruit: Similar in shape and size to a long blackberry. A multiple fruit composed of numerous small, fleshy drupes. Typically a mix of white, pink, red and purple. Edible. Relished by birds. Fruits and bird droppings will stain sidewalks badly.
Foliage: Toothed , deeply lobed , glossy • Foliage Shape: Ovate • Normal foliage color: Green • Underside foliage: Green • Juvenile foliage: Green • Mature foliage: Green • New foliage: Green • Spring foliage: Green • Summer foliage: Alternate leaf arrangement . Leaves have many shapes , can be lobed or not. Serrate leaf margins. Rounded cordate leaf base. Pubescent leaf give a rough texture . Dark green leaf color. • Fall foliage: Dull yellow fall color. Color develops late. Color quality varies.
Size/Age/Growth
Growth Rate: Fast. • Size: to 30 to 35 ft . tall, 35 to 40 ft. wide.
Landscaping
Landscape Uses: For extreme tolerance. Difficult sites. Raise silkworms. Fruit attracts birds. Fruitless and weeping forms more useful than straight species. • Liabilities: Considered "trash tree ". Suckers . Invasive. Bacterial blight , leaf spot, canker , powdery mildewy, scale, witches'. broom . Tree looks unkept. Fruit can be extremely messy, don't plant near walkways or parking lots . • Care: Follow a regular watering schedule during the first growing season to establish a deep, extensive root system . Watering can be reduced after establishment. Feed with a general purpose fertilizer before new growth begins in spring .
Habitat
Disturbed
areas, woodland margins
, fencerows, dry to moist thickets;
0-1500 m
[3].
Hardy
to zone 5, and warmer parts of zone 4.
Typically found at an altitude of 0 to 2,998 meters (0 to 9,836 feet).[4]
Biology
Reproduction
Duration: Perennial
Growth
Culture: Transplants readily. Tolerant of drought , urban and seaside conditions. Full sun . Prune in winter. Prefers moist, well-drained fertile soil. PH adaptable.
Soil: Minimum pH: 6.1 • Maximum pH: 6.5
Sunlight: Sun Exposure: Full sun
Moisture: Water Requirements: Once established needs only occasional water.
Temperature: Heat Zones: High: 8 (>90 to 120 days) Low:3 (>7 to 14 days) (map) • Cold Hardiness: High:8 (10 to 20 F) Low:4 (-30 to -20 F) (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Morus alba var. tatarica (Linnaeus) Seringe • Morus tatarica L.
Notes
Name
Status: Accepted Name
.
Last scrutiny: 15-Mar-2000
Similar Species
Members of the genus Morus
ZipcodeZoo has pages for 43 species, subspecies, varieties, forms, and cultivars in this genus:
M. alba (Common Mulberry) · M. alba 'Beautiful Day' (White Mulberry Tree) · M. alba 'Bellaire' (White Mulberry Tree) · M. alba 'Chaparral' (Fruitless Weeping Mulberry) · M. alba 'Contorted' (Mulberry) · M. alba 'Nuclear Blast' (White Mulberry Tree) · M. alba 'Oscar's' (Mulberry) · M. alba 'Pakistan' (Mulberry) · M. alba 'Pendula' (Weeping White Mulberry) · M. alba 'Shangri La' (Mulberry) · M. alba 'Sweet Lavender' (Mulberry) · M. alba 'White Shahtoot' (Mulberry) · M. australis (Southern Mulberry Tree) · M. bombycis (Wild Korean Mulberry) · M. cathayana (Mulberry) · M. excelsa (African Teak) · M. latifolia (Large-Leaved Mulberry) · M. macroura (Long-Fruited Mulberry) · M. microphylla (Little-Leaf Mulberry) · M. mongolica (Mongolian Mulberry) · M. multicaulis (Multi-Trunked Mulberry) · M. nigra (Black Mulberry) · M. nigra 'Bachuus Noir' (Mulberry) · M. nigra 'Black Beauty' (Black Mulberry) · M. nigra 'Black Tartarian' (Mulberry) · M. nigra 'Buzza Black' (Mulberry) · M. nigra 'Chelsea' (Black Mulberry) · M. nigra 'Shah' (Mulberry) · M. nigra 'Silk Hope' (Mulberry) · M. nigra 'Superberry' (Mulberry) · M. nigra 'Wellington' (Black Mulberry) · M. rubra (Red Mulberry) · M. rubra var. tomentosa (Red Mulberry) · M. rubra 'Gelato' (Mulberry) · M. serrata (Himalayan Mulberry) · M. tiliaefolia (Wild Korean Mulberry) · M. tinctoria (Indian Mulberry) · M. 'Collier' (Mulberry) · M. 'Geraldi Dwarf' (Dwarf Mulberry) · M. 'Illinois Everbearing' (Mulberry Tree) · M. 'Paradise' (Mulberry) · M. 'Red Shahtoot' (Mulberry) · M. 'Unryu' (Contorted Mulberry Tree)
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Further Reading
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- A gardener's year, by H. Rider Haggard. With plan and twenty-five illustrations. London, Longmans, Green, 1905. url .
- A guide to the trees / by Alice Lounsberry...with sixty-four coloured and one hundred and sixty-four black-and-white plates and fifty diagrams, by Mrs. Ellis Rowan...with and introduction by Dr. N. L. Britton. New York: Frederick A. Stokes Company Publishers, [c1900] url , p. 309.
- A guide to the trees by Alice Lounsberry; with sixty-four coloured and one hundred and sixty-four black-and-white plates and fifty-five diagrams by Ellis Rowan; with an introduction by N.L. Britton. Toronto: W. Briggs, c1900. url , .
- A guide to the trees; with sxity-four coloured and one hundred and sixty-four black-and-white plates and fifty-five diagrams, New YorkF.A. Stokes[c1900] url p. 309, p. xi, p. xv.
- A handbook of systematic botany; with a revision of the fungi, by E. Knoblauch; tr. and ed. by M.C. Potter. London, Swan[1904] url p. 354.
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- A manual of Indian timbers; an account of the growth, distribution, and uses of the trees and shrubs of India and Ceylon with descriptions of their wood-structure, by J. S. Gamble... London: S. Low, Marston & co. ltd., 1902. url p. 634.
- A manual of poisonous plants, chiefly of eastern North America, with brief notes on economic and medicinal plants, and numerous illustrations, by L.H. Pammel Cedar Rapids, Ia., The Torch press, 1911. url p. 629, p. 954.
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- A naturalist in western China, with vasculum, camera, and gun; being some account of eleven years' travel, exploration, and observation in the more remote parts of the Flowery kingdom, by Ernest Henry Wilson...with an introduction by Charles Sprague Sargent...With one hundred and one full-page illustrations and a map. London, Methuen & co., ltd.[1913] url p. 220, p. 23, p. 77.
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Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed March 27, 2012.
- Global Biodiversity Information Facility. Accessed November 16, 2007. http://www.gbif.org Mediated distribution data from 32 providers.
- "Morus alba". in Flora of North America Vol. 3. Published by Oxford University Press. Online at EFloras.org.
- Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 29, 2008)
Data Sources
Accessed through GBIF Data Portal November 16, 2007:
- Conservatoire botanique national du Bassin parisien, Conservatoire botanique national du Bassin parisien
- Forest Research Institute, Department of Natural Forests, Herbarium
- International Plant Genetic Resources Institute(IPGRI), EURISCO
- Missouri Botanical Garden, Missouri Botanical Garden
- Oregon State University, Vascular Plant Collection
- School of Life Sciences, Arizona State University, Arizona State University Vascular Plant Herbarium
- The New York Botanical Garden, Bronx River Bioblitz
- The Swedish Museum of Natural History
- , Herbarium of Oskarshamn
- The Swedish Museum of Natural History
- , Lund Botanical Museum
- Tiroler Landesmuseum Ferdinandeum, Tiroler Landesmuseum Ferdinandeum
- US National Plant Germplasm System, United States National Plant Germplasm System Collection
- USDA PLANTS, USDA PLANTS Database
- University of Alabama Biodiversity and Systematics, Herbarium
- University of Washington Burke Museum, Vascular Plant Collection - University of Washington Herbarium
- Utah Valley State College
- , Utah Valley State College Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 1
- Catalogue of Life Accepted Name Code: ITS-19066
- Global Biodiversity Information Facility Taxonkey: 4490920
- Globally Unique Identifier: urn:lsid:ipni.org:names:291504-2
- GRIN Nomen Number: 24607
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19066
- International Plant Names Index (IPNI) ID: 854616-1
- Natural Heritage Network Species Identifier: PDMOR0D010
- U.S.D.A. Plant Symbol: MOTA
- Zipcode Zoo Species Identifier: 50628
Footnotes
- Zhengyi Wu, Zhe-Kun Zhou & Michael G. Gilbert "Moraceae". in Flora of China Vol. 5 Page 21. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Zhengyi Wu, Zhe-Kun Zhou & Michael G. Gilbert "Morus". in Flora of China Vol. 5 Page 22. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Morus alba". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- Mean = 324.880 meters (1,065.879 feet), Standard Deviation = 363.760 based on 1,160 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
