Lycopodium complanatum

• Interesting Facts
• Common Names
• Description
• Biology
• Taxonomy
• Similar Species
• More Info
• Further Reading
• Notes

Interesting Facts

Common Names

Click on the language to view common names.

Description

Family Lycopodiaceae

Plants terrestrial , on rock, or epiphytic. Roots emerging near origin , or growing through cortex and emergent some distance from origin. Horizontal stems present or absent, mainly protostelic, in some species becoming actino- or plectostelic, on substrate surface or subterranean , or forming stolons. Upright shoots simple or branched, usually conspicuously leafy at least at base ; abscising gemmae formed by reduced lateral shoots. Lateral shoots present or absent, simple or branched, branching pattern dichotomous and sometimes pseudomonopodial; leaves uniform or dimorphic or trimorphic. Upright and lateral shoots round or flat in cross section ; leaves on subterranean parts flat, appressed , nonphotosynthetic, and scalelike; leaves on aerial parts appressed, ascending , or spreading , with 1 central unbranched vein , needlelike to lanceolate to ovate , remote to dense and imbricate, with or without basal and/or mucilage canals. Strobili sessile or stalked , upright, nodding , or pendent. Sporangia solitary, adaxial near leaf base or axillary ; subtending leaves (sporophylls) unmodified and photosynthetic to much modified, nonphotosynthetic, reduced, and aggregated in strobili; sporangia reniform to globose , thick-walled with hundreds of spores, outer walls variously modified. Spores all 1 kind, trilete, thick-walled, surfaces pitted to small-grooved, rugulate, or reticulate . Gametophytes subterranean and nonphotosynthetic or surficial and photosynthetic.

Genera 10--15, species 350--400 (7 genera, 27 species in the flora ) : worldwide.

The Lycopodiaceae are an extremely diverse , ancient family . The family may contain even more than the estimated 400 species because the tropical members and the very large genus Phlegmariurus are still poorly known. The relationships among genera of Lycopodiaceae are not well understood because large evolutionary gaps exist among most genera. Some of the genera, notably Diphasiastrum, Huperzia, and Lycopodiella, exhibit extensive interspecific hybridization, which has caused much taxonomic confusion in the past. Differences in expressions of many of the generic characters are subtle, and some of the characters are microscopic.[1]

Genus Lycopodium

Plants mainly trailing on ground . Roots emerging from point of origin on underside of main stems . Horizontal stems on substrate surface or subterranean , long-creeping. Upright shoots scattered along horizontal stem, 5--16 mm diam., round or flat in cross section , unbranched or with 1--4 lateral branchlets . Leaves not imbricate, linear to linear-lanceolate; leaves on horizontal stems scattered, appressed , membranous; leaves on lateral branchlets mostly 6-ranked or more, monomorphic with few exceptions, appressed, ascending to spreading , margins entire to dentate . Gemmiferous branchlets and gemmae absent. Strobili single and sessile or multiple and pedunculate , apex blunt to acute; peduncle, when present, conspicuously leafy; sporophylls extremely reduced, much shorter than peduncle or stem leaves. Sporangia reniform . Spores reticulate , sides at equator convex , angles acute. Gametophytes nonphotosynthetic, mycorrhizal, subterranean, flat and irregularly button-shaped, with ring meristem around circumference. x = 34.

Species 15--25: mainly temperate and subarctic .

In striking contrast to Diphasiastrum, Huperzia, and Lycopodiella, interspecific hybridization is practically unknown in Lycopodium. Many of the species now recognized in Lycopodium have been segregated from Lycopodium clavatum, L. annotinum, and L. jussiaei Desvaux ex Poiret. The three groups given in the key below should probably be treated as subgenera .[2]

Physical Description

Habitat

Typically found at an altitude of 0 to 4,653 meters (0 to 15,266 feet).[3]

Biology

Growth

Sunlight: Sun Exposure: Partial shade to full shade.

Taxonomy

Synonyms

Diphasiastrum complanatum (L.) Holub • Diphasium anceps< /i> (Wallr.) A. & D. Löve • Diphasium complanatum (L.) Rothm. • Diphasium complanatum montellii Kukkonen • Diphasium wallrothii H. P. Fuchs • Lepidotis Complanata • Lepidotis complanata (L.) P. Beauv. • Lycopodium anceps Wallr. • Lycopodium complanatum anceps (Wallr.) Aschers. • Lycopodium complanatum var. canadense Victorin • Stachygynandrum Complanatum

Notes

Publishing author : L. Publication : Sp. Pl. 2: 1104 1753 [1 May 1753]

Similar Species

Members of the genus Lycopodium

ZipcodeZoo has pages for 51 species, subspecies, varieties, forms, and cultivars in this genus:

L. alpinum (Alpine Club Moss) · L. annotinum (Clubmoss) · L. annotinum f. groen;andicum (Clubmoss) · L. annotinum pungens (Stiff Clubmoss) · L. apodum (Meadow Spike Moss) · L. argenteum (Northern Sennet) · L. carinatum (Club Moss) · L. cernua (Lycopodium) · L. circinale (Blackfin Barracuda) · L. clavatum (Common Club Moss) · L. clavatum contiguum (Staghorn Club-Moss) · L. clavatum f. brevipedunculatum (Staghorn Club-Moss) · L. clavatum f. incurvum (Common Club Moss) · L. clavatum f. robustinus (Staghorn Club-Moss) · L. clavatum var. aristatum (Staghorn Club-Moss) · L. clavatum var. clavatum (Running Clubmoss) · L. clavatum var. raddianum (Staghorn Club-Moss) · L. complanatum (Plants) · L. complanatum complanatum (American Clubmoss) · L. complanatum f. canadense (Christmas Green) · L. dendroideum (Tree Groundpine) · L. deuterodensum (Bushy Clubmoss) · L. digitatum (Fan Club-Moss) · L. douglasii (Douglas' Spikemoss) · L. fastigiatum (Mountain Clubmoss) · L. habereri (Haberer's Clubmoss) · L. hickeyi (Hickey's Clubmoss) · L. issleri (Issler's Clubmoss) · L. lagopus (One-Cone Clubmoss) · L. longifolium (Pacific Threadfin) · L. lycopodiella (Lycopodiella) · L. mirabile (Thread-Fishes) · L. nudum (Clubmosses) · L. obscurum (Ground Pine) · L. obscurum f. dendroideum (Flat-Branch Tree Club-Moss) · L. obscurum f. foliaceum (Flat-Branch Tree Club-Moss) · L. obscurum f. parvispicatum (Ground Pine) · L. phlegmaria (Queensland Tassel-Fern) · L. proniflorum (Mullet) · L. pungens (Stiff Clubmoss) · L. sabinaefolium sitchense (Sitka Clubmoss) · L. sabinifolium (Ground-Fir) · L. scariosum (Spreading Clubmoss) · L. sitchense (Alaskan Clubmoss) · L. tristachyum (Deep-Root Clubmoss) · L. venustulum (Hairtip Clubmoss) · L. volubile (Climbing Clubmoss) · L. x habereri (Hybrid) · L. × issleri (Issler's Clubmoss) · L. x zeilleri (Zeiller's Clubmoss) · L. zeilleri (Deeproot Clubmoss)

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Further Reading

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Notes

Contributors

• Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
• Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 11, 2012.
• Global Biodiversity Information Facility. Accessed November 27, 2007. http://www.gbif.org Mediated distribution data from 25 providers.
• Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
• The International Plant Names Index. Accessed Dec 27, 2011.

Data Sources

Accessed through GBIF Data Portal November 15, 2007:

• Australian National Herbarium
• , Australian National Herbarium
• Biologiezentrum der Oberoesterreichischen Landesmuseen, Biologiezentrum Linz
• Bundesamt für Naturschutz / Zentralstelle für Phytodiversität Deutschland, Bundesamt fuer Naturschutz / Zentralstelle fuer Phytodiversitaet Deutschland
• Canadian Museum of Nature, Canadian Museum of Nature Herbarium
• Herbario SANT, Universidade de Santiago de Compostela, SANT herbarium vascular plant collection
• Missouri Botanical Garden, Missouri Botanical Garden
• Royal Botanic Gardens, Kew, Royal Botanic Gardens, Kew
• Taiwan Biodiversity Information Facility, Magnoliophyta
• The Swedish Museum of Natural History
• , Botany
• Tiroler Landesmuseum Ferdinandeum, Tiroler Landesmuseum Ferdinandeum
• USDA PLANTS, USDA PLANTS Database
• University of Alabama Biodiversity and Systematics, Herbarium
• University of Alaska Museum of the North, University of Alaska Museum of the North Herbarium
• University of Washington Burke Museum, Vascular Plant Collection - University of Washington Herbarium
• inatura - Erlebnis Naturschau Dornbirn, inatura - Erlebnis Naturschau Dornbirn

Identifiers

• Biodiversity Heritage Library NamebankID: 2644343
• Catalogue of Life Accepted Name Code: ITS-17025
• Global Biodiversity Information Facility Taxonkey: 13728729
• Globally Unique Identifier: urn:lsid:ipni.org:names:320264-2
• International Plant Names Index (IPNI) ID: 147262-2
• Zipcode Zoo Species Identifier: 2669568

Footnotes

1. Warren H. Wagner Jr. & Joseph M. Beitel "Lycopodiaceae". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
2. "Lycopodium". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
3. Mean = 407.910 meters (1,338.287 feet), Standard Deviation = 484.310 based on 2,076 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]

©2004-2012 David Stang. All rights reserved.