Interesting Facts
Common Names
Common Names in English:
Hybrid, Haberer's Clubmoss
Description
Family Lycopodiaceae
Plants
terrestrial
, on rock, or epiphytic. Roots
emerging near origin
, or growing through cortex and emergent some distance
from origin. Horizontal stems present or absent, mainly protostelic, in some species becoming actino- or plectostelic, on substrate surface or subterranean
, or forming stolons. Upright shoots
simple
or branched, usually conspicuously leafy at least at base
; abscising gemmae formed by reduced lateral
shoots. Lateral shoots present or absent, simple or branched, branching pattern
dichotomous and sometimes pseudomonopodial; leaves uniform
or dimorphic
or trimorphic. Upright and lateral shoots round
or flat in cross
section
; leaves on subterranean parts flat, appressed
, nonphotosynthetic, and scalelike; leaves on aerial
parts appressed, ascending
, or spreading
, with 1 central unbranched vein
, needlelike to lanceolate to ovate
, remote
to dense and imbricate, with or without basal and/or mucilage canals. Strobili sessile or stalked
, upright, nodding
, or pendent. Sporangia solitary, adaxial
near leaf base or axillary
; subtending
leaves (sporophylls) unmodified and photosynthetic to much modified, nonphotosynthetic, reduced, and aggregated in strobili; sporangia reniform to globose
, thick-walled with hundreds
of spores, outer walls variously modified. Spores all 1 kind, trilete, thick-walled, surfaces pitted
to small-grooved, rugulate, or reticulate
. Gametophytes subterranean and nonphotosynthetic or surficial
and photosynthetic.
Genera 10--15, species 350--400 (7 genera, 27 species in the flora
) : worldwide.
The Lycopodiaceae are an extremely diverse
, ancient family
. The family may contain even more than the estimated 400 species because the tropical
members
and the very large genus Phlegmariurus are still poorly known. The relationships
among genera of Lycopodiaceae are not well understood because large evolutionary gaps
exist among most genera. Some of the genera, notably Diphasiastrum, Huperzia, and Lycopodiella, exhibit
extensive interspecific
hybridization, which has caused much taxonomic
confusion in the past. Differences in expressions of many of the generic
characters are subtle, and some of the characters are microscopic.[1]
Genus Lycopodium
Plants
mainly trailing
on ground
. Roots
emerging from point
of origin
on underside of main stems
. Horizontal stems on substrate surface or subterranean
, long-creeping. Upright shoots
scattered
along horizontal stem, 5--16 mm diam., round
or flat in cross
section
, unbranched or with 1--4 lateral
branchlets
. Leaves not imbricate, linear
to linear-lanceolate; leaves on horizontal stems scattered, appressed
, membranous; leaves on lateral branchlets mostly 6-ranked or more, monomorphic
with few exceptions, appressed, ascending
to spreading
, margins
entire to dentate
. Gemmiferous
branchlets and gemmae absent. Strobili single and sessile or multiple
and pedunculate
, apex blunt
to acute; peduncle, when present, conspicuously leafy; sporophylls extremely reduced, much shorter than peduncle or stem leaves. Sporangia reniform
. Spores reticulate
, sides at equator convex
, angles
acute. Gametophytes nonphotosynthetic, mycorrhizal, subterranean, flat and irregularly button-shaped, with ring
meristem around circumference. x
= 34.
Species 15--25: mainly temperate
and subarctic
.
In striking contrast to Diphasiastrum, Huperzia, and Lycopodiella, interspecific
hybridization is practically unknown in Lycopodium. Many of the species now recognized in Lycopodium have been segregated from Lycopodium clavatum, L. annotinum, and L. jussiaei Desvaux ex
Poiret. The three groups given in the key
below should probably be treated as subgenera
.[2]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Lycophytina
(
)
- Infraphylum:
Lycophytae
(
)
- (Auct.) Cavalier-Smith, 1998
- Class:
Lycopodiopsida
(
)
- Bartl.
- Order:
Lycopodiales
(
)
- Dumortier, 1829
- Family:
Lycopodiaceae
(
)
- Palisot de Beauvois ex Mirbel, in Lamarck & Mirbel, 1802
- club mosses
- Genus:
Lycopodium
(
)
- C. Linnaeus, 1753
- Club-moss [Greek lykos, wolf, and pous, podes, foot; in reference to the resemblance of the branch tips to a wolf's paw]
- Subspecies:
habereri
- Botanical name: - Lycopodium x habereri
- Subspecies:
habereri
- Genus:
Lycopodium
(
- Family:
Lycopodiaceae
(
- Order:
Lycopodiales
(
- Class:
Lycopodiopsida
(
- Infraphylum:
Lycophytae
(
- Subphylum:
Lycophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Lycopodium habereri House
Similar Species
Members of the genus Lycopodium
ZipcodeZoo has pages for 51 species, subspecies, varieties, forms, and cultivars in this genus:
L. alpinum (Alpine Club Moss) · L. annotinum (Clubmoss) · L. annotinum f. groen;andicum (Clubmoss) · L. annotinum pungens (Stiff Clubmoss) · L. apodum (Meadow Spike Moss) · L. argenteum (Northern Sennet) · L. carinatum (Club Moss) · L. cernua (Lycopodium) · L. circinale (Blackfin Barracuda) · L. clavatum (Common Club Moss) · L. clavatum contiguum (Staghorn Club-Moss) · L. clavatum f. brevipedunculatum (Staghorn Club-Moss) · L. clavatum f. incurvum (Common Club Moss) · L. clavatum f. robustinus (Staghorn Club-Moss) · L. clavatum var. aristatum (Staghorn Club-Moss) · L. clavatum var. clavatum (Running Clubmoss) · L. clavatum var. raddianum (Staghorn Club-Moss) · L. complanatum (Plants) · L. complanatum complanatum (American Clubmoss) · L. complanatum f. canadense (Christmas Green) · L. dendroideum (Tree Groundpine) · L. deuterodensum (Bushy Clubmoss) · L. digitatum (Fan Club-Moss) · L. douglasii (Douglas' Spikemoss) · L. fastigiatum (Mountain Clubmoss) · L. habereri (Haberer's Clubmoss) · L. hickeyi (Hickey's Clubmoss) · L. issleri (Issler's Clubmoss) · L. lagopus (One-Cone Clubmoss) · L. longifolium (Pacific Threadfin) · L. lycopodiella (Lycopodiella) · L. mirabile (Thread-Fishes) · L. nudum (Clubmosses) · L. obscurum (Ground Pine) · L. obscurum f. dendroideum (Flat-Branch Tree Club-Moss) · L. obscurum f. foliaceum (Flat-Branch Tree Club-Moss) · L. obscurum f. parvispicatum (Ground Pine) · L. phlegmaria (Queensland Tassel-Fern) · L. proniflorum (Mullet) · L. pungens (Stiff Clubmoss) · L. sabinaefolium sitchense (Sitka Clubmoss) · L. sabinifolium (Ground-Fir) · L. scariosum (Spreading Clubmoss) · L. sitchense (Alaskan Clubmoss) · L. tristachyum (Deep-Root Clubmoss) · L. venustulum (Hairtip Clubmoss) · L. volubile (Climbing Clubmoss) · L. x habereri (Hybrid) · L. × issleri (Issler's Clubmoss) · L. x zeilleri (Zeiller's Clubmoss) · L. zeilleri (Deeproot Clubmoss)
More Info
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Further Reading
- Øllgaard, B. 1990. Lycopodiaceae. In: K. Kubitzki et al., eds. 1990+. The Families and Genera of Vascular Plants. 1+ vol. Berlin etc. Vol. 1, pp. 31--39.
- Øllgaard, B. 1987. A revised classification of the Lycopodiaceae s. lat. Opera Bot. 92: 153--178.
- Øllgaard, B. 1989. Index of the Lycopodiaceae. Biol. Skr. 34: 1--135.
- Wagner, W. H. Jr. and J. M. Beitel. 1992. Generic classification of modern North American Lycopodiaceae. Ann. Missouri Bot. Gard. 79: 676--686.
- Wagner, F. S. 1992. Cytological problems in Lycopodium sens. lat. Ann. Missouri Bot. Gard. 79: 718--729.
- Wagner, W. H. Jr. 1993. A new combination for a North American lycopod. Novon 3: 305.
- Hickey, R. J. 1977. The Lycopodium obscurum complex in North America. Amer. Fern J. 67: 45--49.
- Wagner, W. H. Jr., J. M. Beitel, and R. C. Moran. 1989. Lycopodium hickeyi: A new species of North American clubmoss. Amer. Fern J. 79: 119--121.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 11, 2012.
Identifiers
- Biodiversity Heritage Library NamebankID: 446476
- Natural Heritage Network Species Identifier: PPLYC011D0
- Zipcode Zoo Species Identifier: 167533
Footnotes
- Warren H. Wagner Jr. & Joseph M. Beitel "Lycopodiaceae". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
- "Lycopodium". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
