Lycopodium volubile

• Interesting Facts
• Common Names
• Description
• Taxonomy
• Similar Species
• More Info
• Further Reading
• Notes

Interesting Facts

Common Names

Description

Family Lycopodiaceae

Plants terrestrial , on rock, or epiphytic. Roots emerging near origin , or growing through cortex and emergent some distance from origin. Horizontal stems present or absent, mainly protostelic, in some species becoming actino- or plectostelic, on substrate surface or subterranean , or forming stolons. Upright shoots simple or branched, usually conspicuously leafy at least at base ; abscising gemmae formed by reduced lateral shoots. Lateral shoots present or absent, simple or branched, branching pattern dichotomous and sometimes pseudomonopodial; leaves uniform or dimorphic or trimorphic. Upright and lateral shoots round or flat in cross section ; leaves on subterranean parts flat, appressed , nonphotosynthetic, and scalelike; leaves on aerial parts appressed, ascending , or spreading , with 1 central unbranched vein , needlelike to lanceolate to ovate , remote to dense and imbricate, with or without basal and/or mucilage canals. Strobili sessile or stalked , upright, nodding , or pendent. Sporangia solitary, adaxial near leaf base or axillary ; subtending leaves (sporophylls) unmodified and photosynthetic to much modified, nonphotosynthetic, reduced, and aggregated in strobili; sporangia reniform to globose , thick-walled with hundreds of spores, outer walls variously modified. Spores all 1 kind, trilete, thick-walled, surfaces pitted to small-grooved, rugulate, or reticulate . Gametophytes subterranean and nonphotosynthetic or surficial and photosynthetic.

Genera 10--15, species 350--400 (7 genera, 27 species in the flora ) : worldwide.

The Lycopodiaceae are an extremely diverse , ancient family . The family may contain even more than the estimated 400 species because the tropical members and the very large genus Phlegmariurus are still poorly known. The relationships among genera of Lycopodiaceae are not well understood because large evolutionary gaps exist among most genera. Some of the genera, notably Diphasiastrum, Huperzia, and Lycopodiella, exhibit extensive interspecific hybridization, which has caused much taxonomic confusion in the past. Differences in expressions of many of the generic characters are subtle, and some of the characters are microscopic.[1]

Genus Lycopodium

Plants mainly trailing on ground . Roots emerging from point of origin on underside of main stems . Horizontal stems on substrate surface or subterranean , long-creeping. Upright shoots scattered along horizontal stem, 5--16 mm diam., round or flat in cross section , unbranched or with 1--4 lateral branchlets . Leaves not imbricate, linear to linear-lanceolate; leaves on horizontal stems scattered, appressed , membranous; leaves on lateral branchlets mostly 6-ranked or more, monomorphic with few exceptions, appressed, ascending to spreading , margins entire to dentate . Gemmiferous branchlets and gemmae absent. Strobili single and sessile or multiple and pedunculate , apex blunt to acute; peduncle, when present, conspicuously leafy; sporophylls extremely reduced, much shorter than peduncle or stem leaves. Sporangia reniform . Spores reticulate , sides at equator convex , angles acute. Gametophytes nonphotosynthetic, mycorrhizal, subterranean, flat and irregularly button-shaped, with ring meristem around circumference. x = 34.

Species 15--25: mainly temperate and subarctic .

In striking contrast to Diphasiastrum, Huperzia, and Lycopodiella, interspecific hybridization is practically unknown in Lycopodium. Many of the species now recognized in Lycopodium have been segregated from Lycopodium clavatum, L. annotinum, and L. jussiaei Desvaux ex Poiret. The three groups given in the key below should probably be treated as subgenera .[2]

Physical Description

Taxonomy

Synonyms

Pseudodiphasium Volubile

Notes

Publishing author : G.Forst. Publication : Florulae Insularum Australium Prodromus 1786 (Oct. - Nov. 1786)

Name Status: Accepted Name .

Last scrutiny: 12-Mar-2003

Similar Species

Members of the genus Lycopodium

ZipcodeZoo has pages for 51 species, subspecies, varieties, forms, and cultivars in this genus:

L. alpinum (Alpine Club Moss) · L. annotinum (Clubmoss) · L. annotinum f. groen;andicum (Clubmoss) · L. annotinum pungens (Stiff Clubmoss) · L. apodum (Meadow Spike Moss) · L. argenteum (Northern Sennet) · L. carinatum (Club Moss) · L. cernua (Lycopodium) · L. circinale (Blackfin Barracuda) · L. clavatum (Common Club Moss) · L. clavatum contiguum (Staghorn Club-Moss) · L. clavatum f. brevipedunculatum (Staghorn Club-Moss) · L. clavatum f. incurvum (Common Club Moss) · L. clavatum f. robustinus (Staghorn Club-Moss) · L. clavatum var. aristatum (Staghorn Club-Moss) · L. clavatum var. clavatum (Running Clubmoss) · L. clavatum var. raddianum (Staghorn Club-Moss) · L. complanatum (Plants) · L. complanatum complanatum (American Clubmoss) · L. complanatum f. canadense (Christmas Green) · L. dendroideum (Tree Groundpine) · L. deuterodensum (Bushy Clubmoss) · L. digitatum (Fan Club-Moss) · L. douglasii (Douglas' Spikemoss) · L. fastigiatum (Mountain Clubmoss) · L. habereri (Haberer's Clubmoss) · L. hickeyi (Hickey's Clubmoss) · L. issleri (Issler's Clubmoss) · L. lagopus (One-Cone Clubmoss) · L. longifolium (Pacific Threadfin) · L. lycopodiella (Lycopodiella) · L. mirabile (Thread-Fishes) · L. nudum (Clubmosses) · L. obscurum (Ground Pine) · L. obscurum f. dendroideum (Flat-Branch Tree Club-Moss) · L. obscurum f. foliaceum (Flat-Branch Tree Club-Moss) · L. obscurum f. parvispicatum (Ground Pine) · L. phlegmaria (Queensland Tassel-Fern) · L. proniflorum (Mullet) · L. pungens (Stiff Clubmoss) · L. sabinaefolium sitchense (Sitka Clubmoss) · L. sabinifolium (Ground-Fir) · L. scariosum (Spreading Clubmoss) · L. sitchense (Alaskan Clubmoss) · L. tristachyum (Deep-Root Clubmoss) · L. venustulum (Hairtip Clubmoss) · L. volubile (Climbing Clubmoss) · L. x habereri (Hybrid) · L. × issleri (Issler's Clubmoss) · L. x zeilleri (Zeiller's Clubmoss) · L. zeilleri (Deeproot Clubmoss)

More Info

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Further Reading

• A contribution to the flora of Australia / by William Woolls. Sydney: F. White, 1867. url p. 88.
• A contribution to the phytogeography and flora of the Arfak mountains /by L.S. Gibbs. 1917 London: Taylor and Francis, 1917. url p. 32.
• A general system of botany, descriptive and analytical in two parts; tr. from the original by Mrs. Hooker. .. with additions, an appendix on the natural method, and a synopsis of the orders by J.D. Hooker. London, Longmans, 1876. url p. 911.
• An outline of plant geography, by Douglas Houghton Campbell. New York, The Macmillan company, 1926. url p. 232, p. 382.
• Bulletin of the Torrey Botanical Club. 1914 41 New York: Torrey Botanical Club, 1870-1996 url p. 55.
• Contributions from the Hull Botanical Laboratory. [Chicago: University of Chicago Press, 1895- url p. 57, p. 59, p. 63, p. 65.
• Contributions from the University of Michigan Herbarium. 22 1999 Ann Arbor: University Herbarium, University of Michigan, 1939- url p. 144.
• Dutch N. W. New Guinea. A contribution to the phytogeography and flora of the Arfak mountains, & c. By L. S. Gibbs. London: Taylor and Francis, 1917. url p. 32, p. 78.
• Embryogenesis in plants. London, Methuen;[1955] url p. 101, p. 105.
• Flora of the Hawaiian islands: a description of their phanerogams and vascular cryptogams /by William Hillebrand; annotated and published after the author's death by W.F. Hillebrand. 1888 London: New York: Williams & Norgate; B. Westermann, 1888. url p. 642.
• Fossil plants: for students of botany and geology / Cambridge: University Press, 1898-1919. url .
• Hooker, J. D. The botany of the Antarctic voyage of H.M. discovery ships Erebus and Terror in the Years 1839-1843: under the command of Captain Sir James Clark Ross /by Joseph Dalton Hooker. London: Reeve Brothers, 1844-60. url p. 112, p. 55.
• Journal of botany, British and foreign. 4 1866 London: Robert Hardwicke, 1863-1942. url p. 231.
• Journal of the Royal Microscopical Society. Oxford [etc.]Royal Microscopical Society. url p. 210, p. 314, p. 611, p. 62.
• Malayan fern allies. Handbook to the determination of the fern allies of the Malayan islands (incl. those of the Malay peninsula, the Philippines and New Guinea). By Capt. C.R.W.K. van Alderwerelt van Rosenburgh. Pub. by the Dept. of agriculture, industry and commerce, Netherlands India. Batavia: Landsdrukkerij, 1915. url p. 50, p. 52.
• Malayan fern allies: handbook to the determination of the fern allies of the Malayan Islands (incl. those of the Malay Peninsula, the Philippines and New Guinea) / by C.R.W.K. van Alderwerelt van Rosenburgh. Batavia [Djakarta]: Landsdrukkerij, 1915. url p. 50.
• Manual of the New Zealand flora. By T. F. Cheeseman. .. Wellington, N. Z., J. Mackay, Govt. Printer, 1906. url p. 1040, p. 1110.
• Memoirs of the Torrey Botanical Club. 17 1918 Durham, N.C.: Published for the Club by the Seeman Printery, 1889- url p. 96.
• My New Zealand garden / by a Suffolk lady. London: Elliot Stock, 1905. url p. 54.
• Organography of plants, especially of the archegoniatae and spermaphyta, ed. by Isaac Bayley Balfour. Oxford, Clar. Press, 1900-5. url , p. 346, p. 419.
• Phytologia. Bronx Park, New York, H.A. Gleason and H.N. Moldenke, url p. 203.
• Proceedings of the American Academy of Arts and Sciences. Boston: Metcalf and Co., 1846-1958 url p. 222.
• Science progress. London: John Murray, 1916- url p. 42.
• Seemann, B. Flora vitiensis: a description of the plants of the Viti or Fiji islands, with an account of their history, uses, and properties /By Berthold Seemann;. .. plates by Walter Fitch. 1865 - 18 London: L. Reeve, 1865-73. url p. vi.
• The Gardeners' chronicle and agricultural gazette. London: published for the proprietors, 1844-1873. url p. 365.
• The Gardeners' chronicle: a weekly illustrated journal of horticulture and allied subjects. London: [Gardeners Chronicle], 1874-1955. url p. 503.
• The Journal of the Linnean Society. Botany. 42 1914 London: the Society: Longman, Green, Longman, Roberts & Green: ||Williams and Norgate, 1865-1968. url p. 20, p. 207, p. 247.
• The London journal of botany: containing figures and descriptions of such plants as recommend themselves by their novelty, rarity, history, or uses: together with botanical notices and information and and occasional portraits and memoirs of eminent botanists. 3 1844 London: H. Baillie?re, 1842-1848. url p. 412.
• The Philippine journal of science. 5 1910 Manila. url p. 324, p. 70, p. 74, p. 75.
• The Plant world. Baltimore [etc.]Plant World Association [etc.] url p. 226.
• The derivation of the flora of Hawaii. California, Stanford University, 1919. url p. 27.
• The ferns and fern allies of New Zealand, with instructions for their collection and hints on their cultivation. Melbourne, G. Robertson, 1882. url .
• The gardens of the sun: or A naturalist's journal on the mountains and in the forests and swamps of Borneo and the Sulu archipelago. By F. W. Burbidge. .. London, J. Murray, 1880. url p. 349, p. 349.
• The morphology of pteridophytes; the structure of ferns and allied plants. London, Hutchinson University Library[1962] url p. 55, p. 56, p. 67.
• The origin of a land flora, a theory based upon the facts of alternation, London, Macmillan and Co., Ltd., 1908. url .
• The origin of a land flora: a theory based upon the facts of alternation / by F. O. Bower. With numerous illustrations. London: Macmillan, 1908. url p. 329.
• The structure and development of mosses and ferns (Archegoniatae) / by Douglas Houghton Campbell. New York: Macmillan, 1928 [c1905] url p. 493, p. 496.
• The vegetation of the Chatham Islands, Melbourne, J. Ferres, 1864. url .
• Transactions and proceedings of the New Zealand Institute. Wellington: New Zealand Institute. url , p. 100, p. 109, p. 110, p. 146, p. 150, p. 161, p. 162, p. 195, p. 21, p. 211, p. 233, p. 236, p. 237, p. 238, p. 238, p. 261, p. 263, p. 266, p. 267, p. 268, p. 271, p. 282, p. 284, p. 287, p. 295, p. 296, p. 347, p. 351, p. 351, p. 356, p. 361, p. 363, p. 365, p. 367, p. 367, p. 369, p. 378, p. 383, p. 394, p. 4, p. 40, p. 78, p. 81, p. 82, p. 93, p. 94, p. 95, p. 97, p. 99.
• Transactions of the Linnean Society of London, 2nd series: Botany 4 1894-1896 London. url p. 109, p. 255.
• Transactions of the Royal Society of Victoria. Melbourne: The Society, 1888- url p. 39.
• Øllgaard, B. 1990. Lycopodiaceae. In: K. Kubitzki et al., eds. 1990+. The Families and Genera of Vascular Plants. 1+ vol. Berlin etc. Vol. 1, pp. 31--39.
• Øllgaard, B. 1987. A revised classification of the Lycopodiaceae s. lat. Opera Bot. 92: 153--178.
• Øllgaard, B. 1989. Index of the Lycopodiaceae. Biol. Skr. 34: 1--135.
• Wagner, W. H. Jr. and J. M. Beitel. 1992. Generic classification of modern North American Lycopodiaceae. Ann. Missouri Bot. Gard. 79: 676--686.
• Wagner, F. S. 1992. Cytological problems in Lycopodium sens. lat. Ann. Missouri Bot. Gard. 79: 718--729.
• Wagner, W. H. Jr. 1993. A new combination for a North American lycopod. Novon 3: 305.
• Hickey, R. J. 1977. The Lycopodium obscurum complex in North America. Amer. Fern J. 67: 45--49.
• Wagner, W. H. Jr., J. M. Beitel, and R. C. Moran. 1989. Lycopodium hickeyi: A new species of North American clubmoss. Amer. Fern J. 79: 119--121.

Notes

Contributors

• Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
• Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 11, 2012.
• Global Biodiversity Information Facility. Accessed November 26, 2007. http://www.gbif.org Mediated distribution data from 7 providers.
• Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
• The International Plant Names Index. Accessed Dec 27, 2011.
• USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (May 05, 2008)

Data Sources

Accessed through GBIF Data Portal November 26, 2007:

• Australian National Herbarium
• , Australian National Herbarium
• Missouri Botanical Garden, Missouri Botanical Garden
• National Herbarium of New South Wales, Plants of Papua New Guinea

Identifiers

• Biodiversity Heritage Library NamebankID: 2644352
• Catalogue of Life Accepted Name Code: ITS-17058
• Global Biodiversity Information Facility Taxonkey: 13728284
• Globally Unique Identifier: urn:lsid:ipni.org:names:132940-3
• GRIN Nomen Number: 412255
• International Plant Names Index (IPNI) ID: 132940-3
• Zipcode Zoo Species Identifier: 406156

Footnotes

1. Warren H. Wagner Jr. & Joseph M. Beitel "Lycopodiaceae". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
2. "Lycopodium". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]

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