Interesting Facts
Common Names
Common Names in English:
Gromwell
Description
Family Boraginaceae
Herbs perennial
, biennial, or annual
, less often lianas, shrubs
, or trees
, usually bristly
or scabrous-pubescent. Leaves simple
, exstipulate
, alternate, rarely opposite, entire or serrate at margin
. Inflorescences often double
scorpioid cymes, rarely solitary; bracts present or absent. Flowers bisexual
, actinomorphic
, rarely zygomorphic. Calyx usually 5-parted or lobed
, mostly persistent
. Corolla tubular
, campanulate
, rotate, funnelform
, or salverform
; tube
appendages
5, rarely more, mostly trapeziform, rarely absent, sometimes a ring
of hairs
present; limb usually 5-parted; lobes
overlapping, rarely twisted in bud. Stamens 5, inserted
on corolla tube or rarely at throat
, included
or rarely exserted; anthers
introrse
, 2-loculed, usually dorsifixed
at base
, less often medifixed
, dehiscence longitudinal
. Nectaries at base of corolla tube or on disc below ovary. Ovary superior, 2-carpellate; locules 2 and each with 2 ovules, or 4 and each with 1 ovule; ovules nearly atropous
, semianatropous, or anatropous
. Style terminal
or gynobasic
, branched or not. Gynobase
flat, fastigiate
, or subulate
. Fruit 1-4-seeded drupes or nutlets
(mericarps) ; nutlets mostly dry, often ornamented with wings
, prickles and/or glochids (stiff bristles
with barbed
or anchorlike tips
) . Seeds vertical
or oblique
, coat
membranous; embryo straight, less often curved
; cotyledons flat, fleshy
.
About 156 genera and 2500 species: temperate
and tropical regions
, centered in the Mediterranean region; 47 genera and 294 species in China, of which four genera and 156 species are endemic.[1]
Genus Lithospermum
Herbs annual
or perennial
, short strigose
. Leaves alternate. Cymes terminal
or flowers solitary, bracteate
. Calyx 5-parted nearly to base
, slightly enlarged in fruit. Corolla white, yellow, or violet, actinomorphic
, funnelform
or salverform
; throat
with appendages
or bands
of hairs
, or longitudinally crispate
; limb usually campanulate
, 5-parted; lobes
spreading
. Stamens included
; filament
very short; anthers
oblong-linear, apex obtuse
, mucronulate
. Style filiform
, not exserted; stigma entire or indistinctly 2-cleft, capitate. Gynobase
flat. Nutlets
white or gray, ovoid
, smooth
, shiny or tuberculate
; attachment scar
at base adaxially.
About 50 species: Africa, Asia, Europe, North America, South America; five species in China.[2]
Physical Description
Habit: Evergreen .
Flowers: Bloom Period: March, April, May. • Flower Color: medium blue
Size/Age/Growth
Size: under 6" tall.
Biology
Growth
Soil: Minimum pH: 5.6 • Maximum pH: 7.5
Sunlight: Sun Exposure: Sun to Partial Shade.
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Solananae
(
)
- R. Dahlgren Ex Reveal, 1992
- Family:
Boraginaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- borage, bourraches
- Subfamily:
Boraginoideae
(
)
- Tribe:
Lithospermeae
(
)
- Genus:
Lithospermum
(
)
- C. Linnaeus, 1753
- Stoneseed
- Specific epithet:
purpureocoeruleum
- Botanical name: - Lithospermum purpureocoeruleum
- Specific epithet:
purpureocoeruleum
- Genus:
Lithospermum
(
- Tribe:
Lithospermeae
(
- Subfamily:
Boraginoideae
(
- Family:
Boraginaceae
(
- Superorder:
Solananae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Similar Species
Members of the genus Lithospermum
ZipcodeZoo has pages for 27 species, subspecies, varieties, forms, and cultivars in this genus:
L. arvense arvense (Field Gromwell) · L. californicum (California Gromwell) · L. calycosum (Chinati Gromwell) · L. canescens (Gromwell) · L. caroliniense (Carolina Gromwell) · L. caroliniense var. caroliniense (Carolina Puccoon) · L. caroliniense var. croceum (Carolina Puccoon) · L. cobrense (Mountain Gromwell) · L. confine (Arizona Gromwell) · L. diffusum (Scrambling Gromwell) · L. erythrorhizon (Lithospermum) · L. incisum (Fringed Gromwell) · L. latifolium (American Gromwell) · L. matamorense (Rough Gromwell) · L. mirabile (San Antonio Gromwell) · L. multiflorum (Many-Flowered Gromwell) · L. obtusifolium (Rounded-Leaf Gromwell) · L. officinale (European Gromwell) · L. parksii (Parks Stoneseed) · L. parksii I.M.Johnst. var. parksii I.M.Johnst. (Parks' Gromwell) · L. parksii I.M.Johnst. var. rugulosum I.M.Johnst. (Parks' Gromwell) · L. parksii var. parksii (Parks' Stoneseed) · L. parksii var. rugulosum (Parks Stoneseed) · L. purpureocoeruleum (Gromwell) · L. ruderale (Columbia Puccoon) · L. tuberosum (Tuberous Gromwell) · L. viride (American Bugleweed)
More Info
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Further Reading
- Wang Wen-tsai, Liu Yu-lan, Zhu Ge-ling, Lian Yong-shan, Wang Jing-quan & Wang Qing-rui in Kung Xian-wu & Wang Wen-tsai, eds. 1989. Boraginaceae. Fl. Reipubl. Popularis Sin. 64(2): 1-253.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Identifiers
- Zipcode Zoo Species Identifier: 965689
Footnotes
- Gelin Zhu, Harald Riedl & Rudolf V. Kamelin "Boraginaceae". in Flora of China Vol. 16 Page 329. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Lithospermum". in Flora of China Vol. 16 Page 342. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
