Overview
Ornamental shrub from tropical Africa with leaves 3-4 compound , often coppery or purplish; flowers purplish to red; and fruit dark red, about half an inch in diameter.
Interesting Facts
Common Names
Common Names in Portuguese (Brazil):
Léia, Léia-Alaranjada
Description
Family Vitaceae
Woody climbers
, sometimes vines
, rarely small succulent trees
, hermaphroditic
or polygamo-monoecious
to polygamo-dioecious
. Stems unarmed
, sometimes with conspicuous
lenticels
, or bark
sometimes shredding
(in most species of Vitis) ; branches often swollen at 3-7-lacunar nodes; pith
continuous or interrupted
by diaphragms
at nodes; tendrils
simple
, bifurcate
to trifurcate
, or 4-12-branched (in Parthenocissus), usually leaf-opposed, rarely tendrils absent. Raphide
sacs
present in parenchymatous tissues
. Leaves simple, lobed
or unlobed, or digitately or pedately compound
to 1-3-pinnately compound, alternate, distichous, variously toothed
, commonly with multicellular
, stalked
, caducous
spherical
structures known as "pearl" glands
; stipules 2 or rarely absent, often caducous. Flowers small, with prophylls, in panicles, corymbs, or rarely spikes, often leaf-opposite, pseudo-terminal
, or axillary
(in Cayratia and Tetrastigma), actinomorphic
, hypogynous, 4- or 5(-7 as in Rhoicissus) -merous. Calyx with 4 or 5(-7) small teeth or lobes
or a continuous ring. Petals valvate
, 4 or 5(-7), free
or basally connate
, or distally connate forming a calyptra (e.g.
, in Vitis) . Stamens 4 or 5(-7), antepetalous
; anthers
introrse
, dehiscing longitudinally, tetrasporangiate
or rarely bisporangiate
. Floral
disk intrastaminal
, ring-shaped, cupular, or gland-shaped. Ovary superior, 2-loculed; ovules 2 per locule; placentation axile
, appearing nearly basal, apotropous
or anatropous
, bitegmic, crassinucellar; style simple, connate; stigma discoid
or capitate, rarely 4-lobed (Tetrastigma), not papillate
. Fruit a berry, 1-4-seeded. Seeds endotestal, with an abaxial
chalazal
knot
and an adaxial
raphe with 2 furrows
, one on each side; embryo straight, small; endosperm oily, proteinaceous, copious
, ruminate
.
About 14 genera and ca.
900 species: worldwide, but mostly in tropical
and subtropical
regions; eight genera and 146 species (87 endemic, two introduced
) in China.
The family
is important for grapes, wine, and raisins (especially Vitis vinifera, and several other species and hybrids of Vitis) . There are a few ornamental
climbers in the genera Ampelopsis, Cissus, Parthenocissus, and Tetrastigma, of which Parthenocissus tricuspidata of China and Japan and P. quinquefolia of eastern North America are well-known examples. Some plants
(e.g., Ampelopsis japonica, Cissus quadrangularis, and Tetrastigma hemsleyanum) are highly regarded medicines.[1]
Genus Leea
Erect
shrubs
or small trees
. Leaves alternate, 1-4-pinnate, rarely simple
or 3-foliolate. Inflorescences compound
dichasial or umbelliform. Flowers 4- or 5-merous, bisexual
. Lower part of petals adnate
to staminodial tube
on disk. Apex of staminodial tube 5-lobed, connate
to each other by thinner tissue
to form sinuses; apex of lobes
retuse
or 2-lobed, retusely apiculate
to bifid. Filaments
flattened, light brown, alternating with staminodial lobes, curved
inward; anthers
inverted
and packed within staminodial tube in bud; filaments straight and anthers long-exserted in open flower. Disk deeply cupulate
. Ovary discoid
, 4-6(-10) -locular; ovule 1 per locule; style short; stigma slightly thickened. Berry depressed-subglobose, 4-6(-10) -seeded. Endosperm ruminate
.
Geographic distribution is the same as that of the family
.[2]
Physical Description
Species Leea guineensis
Shrubs or small trees . Branchlets terete , almost glabrous . Leaves 2- or 3-pinnate; petiole 6-13 cm, central petiolules 1.5-4 cm, lateral petiolules 0.5-1.5 cm, glabrous; leaflets oval elliptic to long and roundly lanceolate, 5-15 × 2.5-8 cm, base broadly cuneate, or rarely suborbicular , margin with acute teeth, apex acuminate, glabrous; lateral veins 6-11 pairs, abaxial veinlets conspicuous but not protruding. Inflorescences a corymboselike compound dichasium, ca. 50 cm in diam. Pedicel very short or nearly none, sparsely with papillary hairs ; buds ca. 3 mm. Calyx tube cupulate ; sepal triangular, with an acute apex, glabrous. Petals 5, elliptic, red. Stamens 5; filaments 1.2-1.6 mm; anthers yellow. Ovary ovate ; stigma expanded slightly. Berry subglobose, ca. 0.8 cm in diam. [source]
Habit: Evergreen .
Size/Age/Growth
Size: 6-8' tall.
Habitat
Forests , shrublands.[3].
Biology
Growth
Sunlight: Sun Exposure: Light Shade.
Temperature: Cold Hardiness: 9a, 9b, 10a, 10b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Leea manillensis Walpers.
Notes
Publishing author
: G.Don Publication
: Gen. Hist. 1: 712 1831 [early
Aug 1831]
An accepted name
in the RHS
Horticultural Database.
Place of publication: Gen. hist. 1:712. 1831
Name
verified on 22-Aug-2006 by ARS Systematic Botanists. Last
updated: 22-Aug-2006
Similar Species
Members of the genus Leea
ZipcodeZoo has pages for 3 species, subspecies, varieties, forms, and cultivars in this genus:
L. coccinea (Hawaiian Holly) · L. coccinea 'Rubra' (Hawaiian Holly) · L. rubra (Hawaiian Holly)
More Info
- Search for Pictures: images.google.com
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Further Reading
- Bulletin of the British Museum (Natural History). London: The Museum, 1951-1992. url p. 365.
- Catalogue of the African plants London, Printed by order of the Trustees, 1896-1901. url , , .
- Catalogue of the African plants collected by Dr. Friedrich Welwitsch in 1853-61. .. London, Printed by order of the Trustees, 1896-1901. url p. 153, p. 153, p. 164, p. 164.
- Flora Malesiana. general editor, C.G.G.J. van Steenis. Djakarta: Noordhoff-Kolff, 1950- url p. 756, p. 760, p. 777, p. 778, p. 781, p. 782.
- Liberia / by Sir Harry Johnson; with an appendix on the flora of Liberia by Dr. Otto Stapf. London: Hutchinson, 1906. url p. 590, p. 590.
- Liberia, by Sir Harry Johnston. With an appendix on the flora of Liberia, by Dr. Otto Stapf; 28 coloured illustrations by Sir Harry Johnston, 24 botanical drawings by Miss Matilda Smith, 402 black and white il London, Hutchinson, 1906. url p. 590.
- Memoirs of the California Academy of Sciences. San Francisco: The Academy, 1868- url p. 61.
- Oliver, D. Flora of tropical Africa /by Daniel Oliver. .. assisted by other botanists. 1 1868 London: L. Reeve and co., 1868-1999. url p. 472.
- Sketch of the forestry of West Africa with particular reference to its present principal commercial products. London, S. Low, Marston, Searle, & Rivington, 1887. url .
- Wissenschaftliche ergebnisse der Deutschen Zentral-Africa-Expedition, 1907-1908: unter Führung Adolf Friedrichs, herzogs zu Mecklenburg. Leipzig: Klinkhardt & Biermann, 1910- url p. 494, p. 706.
- Li Chaoluan. 1998. Vitaceae (excluding Leea). In: Li Chaoluan, ed., Fl. Reipubl. Popularis Sin. 48(2): ii-vii, 1-3, 12-208.
- Li Chaoluan. 1998. Leea. In: Li Chaoluan, ed., Fl. Reipubl. Popularis Sin. 48(2): 3-12.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 15, 2012.
Data Sources
Accessed through GBIF Data Portal December 01, 2007:
- Herbarium of the University of Aarhus, The AAU Herbarium Database
- Missouri Botanical Garden, Missouri Botanical Garden
- Taiwan Biodiversity Information Facility, Magnoliophyta
Identifiers
- Biodiversity Heritage Library NamebankID: 3477736
- Global Biodiversity Information Facility Taxonkey: 14368475
- GRIN Nomen Number: 315504
- International Plant Names Index (IPNI) ID: 68307-1
- Zipcode Zoo Species Identifier: 707253
Footnotes
- Prof. Zhiduan Chen, Hui Ren & Jun Wen "Vitaceae". in Flora of China Vol. 12 Page 33, 115, 173. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Leea". in Flora of China Vol. 12 Page 169, 173. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Leea guineensis". in Flora of China Vol. 12 Page 169, 170. Missouri Botanical Garden Press. Online at EFloras.org. [back]
