Interesting Facts
Description
Family Juglandaceae
Trees
or rarely shrubs
, deciduous, semievergreen, or evergreen
, monoecious or rarely dioecious; bark
tight (or exfoliating) . Branchlets
with solid or chambered
pith
. Terminal
buds subglobose or ovoid
to oblong
, naked or with scales
. Stipules absent. Leaves alternate (or opposite), odd- or even-pinnate, sometimes trifoliolate
, rarely simple
; leaflets
with glandular
, peltate scales, often resinous
and aromatic
, particularly conspicuous
abaxially on young leaves and twigs
, margin
serrate or rarely entire. Inflorescences pendulous or sometimes erect
, lateral
or terminal, on reduced shoots
arising on branchlets of previous year (old growth) or on current
year’s growth (new growth), of several types: androgynous panicle with male, lateral spikes and female, central spike; androgynous panicle with male, mainly lateral spikes and female, central spike male at apex; cluster
of male spikes and solitary female spike; or solitary male and female spikes. Flowers unisexual
, anemophilous
, rarely entomophilous. Male flowers subtended by an entire or 3-lobed bract; bracteoles 2 or absent; sepals 0-4, adnate
to receptacle when present; stamens 3-40(-100), inserted
on receptacle; filaments
short to nearly absent, free
or united
at base
; anthers
glabrous
or pubescent
, 2-loculed, dehiscing longitudinally. Female flowers with an entire or 3-lobed bracts; bracteoles 2 or 3 (or absent) ; sepals 0-4, adnate to ovary, free at apex; gynoecium of 2 carpels united into an inferior ovary
, 1-loculed, but at base 2-4(-8) -loculed; style 1, short or elongate
, rarely absent; stigmas 2, carinal or commissural
, sometimes 4-lobed, plumose
or fleshy
; ovule 1, orthotropous
. Fruiting spike elongate, and pendulous or short and erect, rarely conelike. Fruit a drupelike nut, 2-4(-8) -chambered at base, with a dehiscent
or indehiscent husk
, or a 2- or 3-winged or disc-winged nutlet
. Seed solitary, without endosperm. Cotyledons 4-lobed, much contorted. Germination hypogeal or epigeal. 2n = (28), 32, (64) .
Nine genera and 60 or more species: mostly in temperate
and subtropical
regions of the N hemisphere; seven genera (one endemic) and 20 species (seven endemic, one introduced
) in China.[1]
Genus Juglans
Shrubs
or trees
, 3-50 m.
Bark
light to dark gray or gray-brown, smooth
or split into ridges
or plates
. Twigs
purplish brown, terete
, stout, sparsely to densely covered with glands
and capitate-glandular hairs
, sometimes also with scales
or fasciculate hairs, early in season
with multiradiate
hairs; leaf scars
triangular or 3-lobed, large; pith
chambered
. Bud scales valvate
, densely hirsute
. Leaves usually odd-, sometimes even-pinnate; petiole
and rachis with indument
as twigs. Leaflets
5-25, sessile or subsessile
, often aromatic
, uniform
in size or median
leaflets largest, (2.5-) 4.3-15(-17.5) × 0.8-6.5 cm; surfaces usually with nonglandular hairs (simple
and/or fasciculate), glandular
hairs, sessile glands, and/or scales, sometimes glabrous
. Staminate
catkins solitary from 2d-year twigs, sessile; stamens 7-50 per flower, glabrous or pilose
. Pistillate
flowers solitary or in terminal
racemes
. Fruits nuts enclosed in husks
, not compressed
; husks thick, indehiscent; nuts tan, neither compressed nor angled
, grooved
, ridged
, rugulose
, or smooth; shells
thick. Seeds sweet. x
= 16.
Species 21: North America, Mexico, West Indies, Central America, South America, Eurasia
.
Juglans is a very important source of edible nuts, dyes, and wood
for cabinet work, furniture, and construction. Juglans regia Linnaeus, the walnut of commerce, is widely cultivated in California; it is easily distinguished from native species
by its leaves with 5-11 broad, entire leaflets and nuts with thin rugulose shells, not grooved or ridged. Because of its sensitivity to native
pathogens
, J. regia is usually grown as stem-grafts on roots
of native or hybrid walnuts (see discussion under J. hindsii). Occasional seedlings of J. regia have been reported from the vicinity of cultivated plants
, but these seldom, if ever, live to maturity.
The growth form
, bark, and fruit are important taxonomically in Juglans, but these usually are not available on herbarium
specimens. As with many woody plants
, the first one or two leaves of the season (i.e.
, the lowermost leaves on the twig
) are sometimes atypical
in structure, having broader, blunter leaflets. The fasciculate hairs on the veins normally have more rays than those on the blade
. In addition to the types of hairs described below, small multiradiate hairs are found on the immature
twigs, petioles, rachises, and midribs
. They are usually gone when the leaf is fully expanded, but they may persist for a short time afterwards.[2]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Juglandanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Fagales
(
)
- Engler, 1892
- Family:
Juglandaceae
(
)
- DC. ex Perleb, 1818, nom. cons.
- walnuts
- Subfamily:
Juglandoideae
(
)
- Tribe:
Juglandeae
(
)
- Genus:
Juglans
(
)
- C. Linnaeus, 1753
- Walnut
- Specific epithet:
compressa
- Muhl.
- Botanical name: - Juglans compressa Muhl.
- Specific epithet:
compressa
- Muhl.
- Genus:
Juglans
(
- Tribe:
Juglandeae
(
- Subfamily:
Juglandoideae
(
- Family:
Juglandaceae
(
- Order:
Fagales
(
- Superorder:
Juglandanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author : Muhl. Publication : in Ges. Naturf. Fr. Berl. Neue Schr. iii. (1801) 390; Willd. Sp. Pl. iv . 458
Similar Species
Members of the genus Juglans
ZipcodeZoo has pages for 82 species, subspecies, varieties, forms, and cultivars in this genus:
J. ailanthifolia (Japanese Walnut) · J. ailanthifolia var. cordiformis (Heartnut) · J. ailantifolia (Japanese Walnut) · J. australis (Nogal Criollo) · J. bixbyi (Bixby Walnut) · J. boliviana (Bolivian Walnut) · J. californica (Northern California Black Walnut) · J. californica S.Watson var. californica S.Watson (Southern California Black Walnut) · J. californica var. californica (California Walnut) · J. cinerea (Butternut) · J. cinerea 'Bountiful' (Butternut) · J. cinerea 'Loumis' (Butternut) · J. cordiformis 'Campbell Cw1' (Heartnut) · J. cordiformis 'Campbell Cw3' (Heartnut) · J. cordiformis 'Campbell Cww' (Heartnut) · J. cordiformis 'Frank' (Heartnut) · J. cordiformis 'Imshu' (Heartnut) · J. cordiformis 'Locket' (Heartnut) · J. cordiformis 'Marvel' (Heartnut) · J. cordiformis 'Mitchell Hybrid' (Heartnut) · J. cordiformis 'Rhodes' (Heartnut) · J. cordiformis 'Simcoe' (Heartnut) · J. cordiformis 'Stealth' (Heartnut) · J. cordiformis 'Wright' (Heartnut) · J. hindsii (Northern California Black Walnut) · J. illinoinensis (Pecan) · J. intermedia (Intermediate Walnut) · J. jamaicensis (Walnut) · J. major (Arizona Black Walnut) · J. mandshurica (Manchurian Walnut) · J. microcarpa (Little Walnut) · J. microcarpa var. microcarpa (Little Walnut) · J. microcarpa var. stewartii (Stewarts Little Walnut) · J. microcarpa var. stewartii (I.M.Johnst.) W.Manning (Stewart's Little Walnut) · J. microcarpa subsp. major (Little Walnut) · J. neotropica (Andean Walnut) · J. nigra (Black Walnut) · J. nigra 'Beineke 11' (Black Walnut 'beineke 11') · J. nigra 'Black Gem' (Black Walnut) · J. nigra 'Daniels' (Black Walnut) · J. nigra 'Emma Kay' (Black Walnut) · J. nigra 'Football 2' (Black Walnut) · J. nigra 'Krause' (Black Walnut) · J. nigra 'Kwik Krop' (Black Walnut) · J. nigra 'Laciniata' (Black Walnut) · J. nigra 'Lamb's Curly' (Black Walnut) · J. nigra 'Ridgeway' (Black Walnut) · J. nigra 'Rowher' (Black Walnut) · J. nigra 'Sauber 1' (Black Walnut) · J. nigra 'Schrieber' (Black Walnut) · J. nigra 'Sparrow' (Black Walnut) · J. nigra 'Surprise' (Black Walnut) · J. nigra 'Thomas Black' (Black Walnut) · J. nigra 'Thomas Myers' (Black Walnut) · J. regia (Carpathian Walnut) · J. regia var. Carpathian (Carpathian Walnut) · J. regia 'Allegheny' (Carpathian Walnut) · J. regia 'Bedco 1' (Carpathian Walnut) · J. regia 'Broadview' (Dwarf English Walnut) · J. regia 'Cascade' (Carpathian Walnut) · J. regia 'Champion' (Carpathian Walnut) · J. regia 'China-B' (Carpathian Walnut) · J. regia 'Coble #2' (Carpathian Walnut) · J. regia 'Colby' (Carpathian Walnut) · J. regia 'Greenhaven' (Carpathian Walnut) · J. regia 'Hansen' (Carpathian Walnut) · J. regia 'Kaiser' (Carpathian Walnut) · J. regia 'Ky Giant' (Carpathian Walnut) · J. regia 'Lake' (Carpathian Walnut) · J. regia 'Looking Glass' (Carpathian Walnut) · J. regia 'Mckinster' (Carpathian Walnut) · J. regia 'Northern Prize' (Carpathian Walnut) · J. regia 'Perry' (Carpathian Walnut) · J. regia 'Purpurea' (Carpathian Walnut) · J. regia 'Reda' (Carpathian Walnut) · J. regia 'S-1' (Carpathian Walnut) · J. regia 'Shiawassee' (Carpathian Walnut) · J. regia 'Somers' (Carpathian Walnut) · J. regia 'Utah Giant' (Carpathian Walnut) · J. × bixbyi (Bixby Walnut) · J. x intermedia (Intermediate Walnut) · J. 'Royal' (Walnut)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Agricultural botany an enumeration and description of useful plants and weeds, which merit the notice, or require the attention, of American agriculturists / New York: J.W. Moore; 1847. url p. 156.
- Agricultural botany: Philadelphia, J. W. Moore;1847. url p. 156.
- An encyclopaedia of trees and shrubs: being the Arboretum et fruticetum britannicum abridged: containing the hardy trees and shrubs of Britain, native and foreign, scientifically and popularly described; with their propagation, culture, by J. C. Loudon. London: F. Warne; 1869. url p. 739.
- Annotated list of the ferns and flowering plants of New York state, by Homer D. House. Albany, The University of the state of New York, 1924. url p. 256.
- Bartonia;proceedings of the Philadelphia botanical club. .. 1946 Philadelphia, Philadelphia Botanical Club, Academy of Natural Sciences. url p. 54, p. 89.
- Botany of the United States north of Virginia; comprising descriptions of the flowering and fern-like plants hitherto found in those states, arranged according to the natural system. With a synopsis of the genera according to the Linnean system, a New York, Harper, 1868. url p. 337.
- Botany of the northern and middle states, or, A description of the plants found in the United States, north of Virginia arranged according to the natural system, with a synopsis of the genera according to the Linnaean system--a sketch of the rudiments of botany, and a glossary of terms / by Lewis C. Beck. Albany: Webster and Skinners, 1833. url p. 336.
- Compendium floræ philadelphicæ: containing a description of the indigenous and naturalized plants found within a circuit of ten miles around Philadelphia / by William P.C. Barton. Philadelphia: published by M. Carey, 1818. url p. 178, p. 179.
- Florula Bostoniensis. A collection of plants of Boston and its vicinity, with their generic and specific characters, principal synonyms, descriptions, places of growth, and time of flowering, and occasional remarks. Boston, C. C. Little and J. Brown, 1840. url p. 380.
- Florula bostoniensis. A collection of plants of Boston and its environs, with their generic and specific characters, synonyms, descriptions, places of growth, and time of flowering, and occasional remarks. Boston, Cummings and Hilliard, 1814. url p. 229.
- Hortus lignosus londinensis: or, A catalogue of all the ligneous plants, indigenous and foreign, hardy and half-hardy, cultivated in the gardens and grounds in the neighbourhood of London: with all their synonymes, scientific and By J. C. Loudon. London, The author, 1838. url p. 95.
- Loudon, J. C. Loudon's Hortus britannicus.A catalogue of all the plants indigenous, cultivated in, or introduced to Britain. Part I. The Linnaean arrangement... Part II. The Jussieuean arrangement... /Edited by J. C. Loudon. 1890 London: Printed for Longman, Rees, Orme, Brown, and Green, 1830. url p. 384.
- Natural history of New York. Albany: D. Appleton: 1842-1894. url p. 181, p. 181.
- Pursh, F. T. Flora Americae Septentrionalis: or, A systematic arrangement and description of the plants of North america. Containing, besides what have been described by preceding authors, many new and rare species, collected during twelve years travels and residence in that country /by Frederick Pursh. .. 2 1814 London: Printed for White, Cochrance, and co., 1814. url p. 637.
- Report on the forests of North America (exclusive of Mexico) / by Charles S. Sargent. Washington [D.C.]: G.P.O., 1884. url p. 132.
- Reports of the Survey. Botanical series. Minneapolis [etc.]1892- url p. 793.
- Sweet's Hortus britannicus: or a catalogue of plants cultivated in the gardens of Great Britain, arranged in natural orders / by Robert Sweet. London: J. Ridgway, 1826. url p. 97.
- The American farmer's encyclopedia and dictionary of rural affairs: embracing all the most recent discoveries in agricultural chemistry / adapted to the comprehension of unscientific readers by Gouverneur Emerson. Philadelphia: Carey and Hart, 1844. url p. 621.
- The American farmer's encyclopedia: embracing all the recent discoveries in agricultural chemistry, and the use of mineral, vegetable and animal manures, with descriptions and figures of American insects, injurious to vegetation. Being by Gouverneur Emerson, of Pennsylvania, upon the basis of Johnson's Farmer's encyclopedia. New York: A.O. Moore, Agricultural Book Publisher, 1858, [c1857] url p. 621.
- The American farmer's encyclopedia; embracing all the recent discoveries in agricultural chemistry, New York, A. O. Moore, 1858. url p. 621.
- The Metaspermae of the Minnesota Valley: a list of the higher seed-producing plants indigenous to the drainage-basin of the Minnesota River / by Conway MacMillan. Minneapolis: Geological and Natural History Survey of Minnesota, 1892. url p. 793.
- The Metaspermae of the Minnesota valley; a list of the higher seed-producing plants indigenous to the drainage-basin of the Minnesota river. Minneapolis[Harrison & Smith, State Printers]1892 url p. 793.
- The North American sylva, or, A description of the forest trees of the United States, Canada, and Nova Scotia, not described in the work of F. Andrew Michaux and containing all the forest trees discovered in the Rocky Mountains, the territory of Oregon, down to the shores of the Pacific and into the confines of California, as well as in various parts of th Philadelphia: J. Dobson, 1842. url .
- The North American sylva; or, A description of the forest trees of the United States, Canada and Nova Scotia. Considered particularly with respect to their use in the arts and their introduction into commerce. To which is ad Tr. from the French of F. Andrew Michaux. .. with notes by J. Jay Smith. Philadelphia, Rice, Rutter & co., 1865. url p. 55.
- The Universal herbal; or, Botanical, medical, and agricultural dictionary; containing an account of all the known plants in the world, arranged according to the Linnean system. Specifying the uses to which they are or may be applied, whether as food, as medicine, or in the a London, Caxton Press[1824] url p. 856, p. 859.
- The farmer's and planter's encyclopaedia of rural affairs; embracing all the most recent discoveries in agricultural chemistry, suited to the comprehension of unscientific readers. By Cuthbert W. Johnson. .. With extensive additions adapting the work to the United States. .. By Gouverneur Emerson. .. Philadelphia, J. B. Lippincott & Co., 1868. url p. 621.
- The farmer's encyclopædia, and dictionary of rural affairs; embracing all the most recent discoveries in agricultural chemistry. Adapted to the comprehension of unscientific readers. .. By Cuthbert W. Johnson. .. Adapted to the United States by Gouverneur Emerson. Philadelphia, Carey and Hart, 1844. url p. 621.
- The gardener's magazine and register of rural & domestic improvement. London: Longman, Rees, Orme, Brown and Green, 1826-1844. url p. 483.
- The nut culturist: a treatise on the propagation, planting and cultivation of nut-bearing trees and shrubs, adapted to the climate of the United States. .. / New York: Orange Judd, 1896. url , p. 157.
- The silva of North America: a description of the trees which grow naturally in North America exclusive of Mexico /by Charles Sprague Sargent. .. illustrated with figures and analyses drawn from nature by Charles Edward Faxon. .. 14 1902 Boston, New York: Houghton, Mifflin and company, 1891-1902. url p. 128, p. 153, p. 170.
- The trees of Great Britain & Ireland / by Henry John Elwes and Augustine Henry. Edinburgh: Priv. print., 1906-13. url p. 601.
- Trees and shrubs; an abridgment of the Arboretum et fruticetum britannicum: containing the hardy trees and shrubs of Britain, native and foreign, scientifically and popularly described; with their propagation, culture LondonF. Warne1875 url p. 739.
- Kuang Ko-zen & Lu An-ming. 1979. Juglandaceae. In: Kuang Ko-zen & Li Pei-chun, eds., Fl. Reipubl. Popularis Sin. 21: 6-44.
- McGranahan, G. H. and P. B. Catlin. 1987. Juglans rootstocks. In: R. C. Rom and R. F. Carlson, eds. 1987. Rootstocks for Fruit Crops. New York. Pp. 411-450.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Identifiers
- Biodiversity Heritage Library NamebankID: 10683559
- Global Biodiversity Information Facility Taxonkey: 15529382
- Globally Unique Identifier: urn:lsid:ipni.org:names:442339-1
- International Plant Names Index (IPNI) ID: 442339-1
- Zipcode Zoo Species Identifier: 3466534
Footnotes
- Anmin Lu, Donald E. Stone & L. J. Grauke "Juglandaceae". in Flora of China Vol. 4 Page 277. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Alan T. Whittemore & Donald E. Stone "Juglans". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
