Overview
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Critically Endangered |
|
Interesting Facts
Common Names
Click on the language to view common names.
Common Names in English:
Northern California Black Walnut, Hinds´ Black Walnut, Hinds´ Walnut, Hinds's Black Walnut, Northern California walnut, Northern California Walnut, Walnut
Common Names in German:
Kalifornische Walnuß
Common Names in unspecified:
Northern California Walnut
Description
Family Juglandaceae
Trees
or rarely shrubs
, deciduous, semievergreen, or evergreen
, monoecious or rarely dioecious; bark
tight (or exfoliating) . Branchlets
with solid or chambered
pith
. Terminal
buds subglobose or ovoid
to oblong
, naked or with scales
. Stipules absent. Leaves alternate (or opposite), odd- or even-pinnate, sometimes trifoliolate
, rarely simple
; leaflets
with glandular
, peltate scales, often resinous
and aromatic
, particularly conspicuous
abaxially on young leaves and twigs
, margin
serrate or rarely entire. Inflorescences pendulous or sometimes erect
, lateral
or terminal, on reduced shoots
arising on branchlets of previous year (old growth) or on current
year’s growth (new growth), of several types: androgynous panicle with male, lateral spikes and female, central spike; androgynous panicle with male, mainly lateral spikes and female, central spike male at apex; cluster
of male spikes and solitary female spike; or solitary male and female spikes. Flowers unisexual
, anemophilous
, rarely entomophilous. Male flowers subtended by an entire or 3-lobed bract; bracteoles 2 or absent; sepals 0-4, adnate
to receptacle when present; stamens 3-40(-100), inserted
on receptacle; filaments
short to nearly absent, free
or united
at base
; anthers
glabrous
or pubescent
, 2-loculed, dehiscing longitudinally. Female flowers with an entire or 3-lobed bracts; bracteoles 2 or 3 (or absent) ; sepals 0-4, adnate to ovary, free at apex; gynoecium of 2 carpels united into an inferior ovary
, 1-loculed, but at base 2-4(-8) -loculed; style 1, short or elongate
, rarely absent; stigmas 2, carinal or commissural
, sometimes 4-lobed, plumose
or fleshy
; ovule 1, orthotropous
. Fruiting spike elongate, and pendulous or short and erect, rarely conelike. Fruit a drupelike nut, 2-4(-8) -chambered at base, with a dehiscent
or indehiscent husk
, or a 2- or 3-winged or disc-winged nutlet
. Seed solitary, without endosperm. Cotyledons 4-lobed, much contorted. Germination hypogeal or epigeal. 2n = (28), 32, (64) .
Nine genera and 60 or more species: mostly in temperate
and subtropical
regions of the N hemisphere; seven genera (one endemic) and 20 species (seven endemic, one introduced
) in China.[1]
Genus Juglans
Shrubs
or trees
, 3-50 m.
Bark
light to dark gray or gray-brown, smooth
or split into ridges
or plates
. Twigs
purplish brown, terete
, stout, sparsely to densely covered with glands
and capitate-glandular hairs
, sometimes also with scales
or fasciculate hairs, early in season
with multiradiate
hairs; leaf scars
triangular or 3-lobed, large; pith
chambered
. Bud scales valvate
, densely hirsute
. Leaves usually odd-, sometimes even-pinnate; petiole
and rachis with indument
as twigs. Leaflets
5-25, sessile or subsessile
, often aromatic
, uniform
in size or median
leaflets largest, (2.5-) 4.3-15(-17.5) × 0.8-6.5 cm; surfaces usually with nonglandular hairs (simple
and/or fasciculate), glandular
hairs, sessile glands, and/or scales, sometimes glabrous
. Staminate
catkins solitary from 2d-year twigs, sessile; stamens 7-50 per flower, glabrous or pilose
. Pistillate
flowers solitary or in terminal
racemes
. Fruits nuts enclosed in husks
, not compressed
; husks thick, indehiscent; nuts tan, neither compressed nor angled
, grooved
, ridged
, rugulose
, or smooth; shells
thick. Seeds sweet. x
= 16.
Species 21: North America, Mexico, West Indies, Central America, South America, Eurasia
.
Juglans is a very important source of edible nuts, dyes, and wood
for cabinet work, furniture, and construction. Juglans regia Linnaeus, the walnut of commerce, is widely cultivated in California; it is easily distinguished from native species
by its leaves with 5-11 broad, entire leaflets and nuts with thin rugulose shells, not grooved or ridged. Because of its sensitivity to native
pathogens
, J. regia is usually grown as stem-grafts on roots
of native or hybrid walnuts (see discussion under J. hindsii). Occasional seedlings of J. regia have been reported from the vicinity of cultivated plants
, but these seldom, if ever, live to maturity.
The growth form
, bark, and fruit are important taxonomically in Juglans, but these usually are not available on herbarium
specimens. As with many woody plants
, the first one or two leaves of the season (i.e.
, the lowermost leaves on the twig
) are sometimes atypical
in structure, having broader, blunter leaflets. The fasciculate hairs on the veins normally have more rays than those on the blade
. In addition to the types of hairs described below, small multiradiate hairs are found on the immature
twigs, petioles, rachises, and midribs
. They are usually gone when the leaf is fully expanded, but they may persist for a short time afterwards.[2]
Physical Description
Species Juglans hindsii
Trees
, 6-23 m.
Bark
light or medium gray, split into smooth
or ±
scaly
plates
. Twigs
with distal edge
of leaf scar
shallowly to deeply
notched
, not bordered
by well-defined band
of pubescence
; pith
light
brown. Terminal
buds ellipsoid
to oblong
, flattened, 6-8 mm.
Leaves
22-45 cm; petiole
3-8 cm. Leaflets
13-21, narrowly triangular to
lanceolate, symmetric
or weakly falcate
, (5.6-) 7.3-13 × (1-)
1.9-2.8 cm, margins
serrate, apex acuminate; surfaces abaxially glabrous
or with sparse glands
, sparse glands and few capitate-glandular hairs
scattered
along major veins, fasciculate hairs conspicuously tufted
in axils of proximal
veins, sometimes also on adjacent
blade
and
edges of midrib
, adaxially glabrous or with scattered scales, major
veins glabrous or with sparse scattering of glands and few capitate-glandular
hairs, without nonglandular hairs; terminal leaflet well developed.
Staminate
catkins 6-15 cm; stamens 20-40 per flower; pollen sacs
1-1.4 mm. Fruits 1-2, globose
, 3.5-5 cm; nuts ovoid
to ovoid-globose,
2.4-3.2 cm, smooth or nearly so or shallowly and indistinctly ridged
or grooved
. [source]
Before 1850, Juglans hindsii was restricted
to a few locations
(J. R. Griffin and W. B
. Critchfield 1972). It has been widely used
as a rootstock
for grafting J. regia and has been planted extensively
in many parts of California for this purpose. It is now naturalized
in many areas where it apparently did not occur before the introduction
of commercial
walnut growing. Possibly some of these naturalized
populations are introgressed with J. nigra, since spontaneous hybridization
between J. hindsii and J. nigra has been reported in areas where
both species have been planted. These hybrids are difficult to distinguish
from J. hindsii unless fruit are present. Currently most commercial
walnut orchards use hybrid rootstocks, usually J. hindsii ×
J. regia (G. H. McGranahan and P. B. Catlin 1987). [source]
Flowers: Bloom Period: February, March, April, May.
Habitat
Along streams , sometimes on disturbed slopes ; of conservation concern; 0-300 m [3].
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Juglandanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Fagales
(
)
- Engler, 1892
- Family:
Juglandaceae
(
)
- DC. ex Perleb, 1818, nom. cons.
- walnuts
- Subfamily:
Juglandoideae
(
)
- Tribe:
Juglandeae
(
)
- Genus:
Juglans
(
)
- C. Linnaeus, 1753
- Walnut
- Specific epithet:
hindsii
- (Jeps.) Jeps. ex R.E.Sm.
- Botanical name: - Juglans hindsii (Jeps.) Jeps. ex R.E.Sm.
- Specific epithet:
hindsii
- (Jeps.) Jeps. ex R.E.Sm.
- Genus:
Juglans
(
- Tribe:
Juglandeae
(
- Subfamily:
Juglandoideae
(
- Family:
Juglandaceae
(
- Order:
Fagales
(
- Superorder:
Juglandanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Juglans californica hindsii Jepson • Juglans californica S. Watson Var. hindsii Jepson • Juglans californica var. hindsii Jepson
Notes
Basionym
: Juglandaceae Juglans californica S.Watson var. hindsii
Jeps.
Basionym author: (Jeps.)
Basionym: Juglandaceae Juglans californica S.Watson var. hindsii
Jeps.
Basionym author: (Jeps.)
Name
Status: Accepted Name
.
Last scrutiny: 15-Mar-2000
Similar Species
Members of the genus Juglans
ZipcodeZoo has pages for 82 species, subspecies, varieties, forms, and cultivars in this genus:
J. ailanthifolia (Japanese Walnut) · J. ailanthifolia var. cordiformis (Heartnut) · J. ailantifolia (Japanese Walnut) · J. australis (Nogal Criollo) · J. bixbyi (Bixby Walnut) · J. boliviana (Bolivian Walnut) · J. californica (Northern California Black Walnut) · J. californica S.Watson var. californica S.Watson (Southern California Black Walnut) · J. californica var. californica (California Walnut) · J. cinerea (Butternut) · J. cinerea 'Bountiful' (Butternut) · J. cinerea 'Loumis' (Butternut) · J. cordiformis 'Campbell Cw1' (Heartnut) · J. cordiformis 'Campbell Cw3' (Heartnut) · J. cordiformis 'Campbell Cww' (Heartnut) · J. cordiformis 'Frank' (Heartnut) · J. cordiformis 'Imshu' (Heartnut) · J. cordiformis 'Locket' (Heartnut) · J. cordiformis 'Marvel' (Heartnut) · J. cordiformis 'Mitchell Hybrid' (Heartnut) · J. cordiformis 'Rhodes' (Heartnut) · J. cordiformis 'Simcoe' (Heartnut) · J. cordiformis 'Stealth' (Heartnut) · J. cordiformis 'Wright' (Heartnut) · J. hindsii (Northern California Black Walnut) · J. illinoinensis (Pecan) · J. intermedia (Intermediate Walnut) · J. jamaicensis (Walnut) · J. major (Arizona Black Walnut) · J. mandshurica (Manchurian Walnut) · J. microcarpa (Little Walnut) · J. microcarpa var. microcarpa (Little Walnut) · J. microcarpa var. stewartii (Stewarts Little Walnut) · J. microcarpa var. stewartii (I.M.Johnst.) W.Manning (Stewart's Little Walnut) · J. microcarpa subsp. major (Little Walnut) · J. neotropica (Andean Walnut) · J. nigra (Black Walnut) · J. nigra 'Beineke 11' (Black Walnut 'beineke 11') · J. nigra 'Black Gem' (Black Walnut) · J. nigra 'Daniels' (Black Walnut) · J. nigra 'Emma Kay' (Black Walnut) · J. nigra 'Football 2' (Black Walnut) · J. nigra 'Krause' (Black Walnut) · J. nigra 'Kwik Krop' (Black Walnut) · J. nigra 'Laciniata' (Black Walnut) · J. nigra 'Lamb's Curly' (Black Walnut) · J. nigra 'Ridgeway' (Black Walnut) · J. nigra 'Rowher' (Black Walnut) · J. nigra 'Sauber 1' (Black Walnut) · J. nigra 'Schrieber' (Black Walnut) · J. nigra 'Sparrow' (Black Walnut) · J. nigra 'Surprise' (Black Walnut) · J. nigra 'Thomas Black' (Black Walnut) · J. nigra 'Thomas Myers' (Black Walnut) · J. regia (Carpathian Walnut) · J. regia var. Carpathian (Carpathian Walnut) · J. regia 'Allegheny' (Carpathian Walnut) · J. regia 'Bedco 1' (Carpathian Walnut) · J. regia 'Broadview' (Dwarf English Walnut) · J. regia 'Cascade' (Carpathian Walnut) · J. regia 'Champion' (Carpathian Walnut) · J. regia 'China-B' (Carpathian Walnut) · J. regia 'Coble #2' (Carpathian Walnut) · J. regia 'Colby' (Carpathian Walnut) · J. regia 'Greenhaven' (Carpathian Walnut) · J. regia 'Hansen' (Carpathian Walnut) · J. regia 'Kaiser' (Carpathian Walnut) · J. regia 'Ky Giant' (Carpathian Walnut) · J. regia 'Lake' (Carpathian Walnut) · J. regia 'Looking Glass' (Carpathian Walnut) · J. regia 'Mckinster' (Carpathian Walnut) · J. regia 'Northern Prize' (Carpathian Walnut) · J. regia 'Perry' (Carpathian Walnut) · J. regia 'Purpurea' (Carpathian Walnut) · J. regia 'Reda' (Carpathian Walnut) · J. regia 'S-1' (Carpathian Walnut) · J. regia 'Shiawassee' (Carpathian Walnut) · J. regia 'Somers' (Carpathian Walnut) · J. regia 'Utah Giant' (Carpathian Walnut) · J. × bixbyi (Bixby Walnut) · J. x intermedia (Intermediate Walnut) · J. 'Royal' (Walnut)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 307.
- An introduction to California plant life Berkeley: University of California Press, [c1974] url p. 121.
- Botanical abstracts. Baltimore, Williams & Wilkins Co. url p. 61.
- California fish and game. [San Francisco, etc.]: State of California, Resources Agency, Dept. of Fish and Game. url p. 139, p. 140, p. 148, p. 266, p. 44.
- Genetics in relation to agriculture, New York, McGraw-Hill book company, inc.; [etc., etc.]1918. url p. 622.
- Great Basin naturalist memoirs. 1979 [Provo, Utah]Brigham Young University, 1976-1992. url p. 131.
- Microbiology for agricultural and domestic science students. Philadelphia, Blakiston[c1917] url .
- Microbiology, a text-book of microörganisms, general and applied. Contributors: F.T. Bioletti. .. R.E. Buchanan. .. [and others] ed. by Charles E. Marshall. .. Philadelphia, P. Blakiston's Son & Co.[c1921] url p. 965.
- Microbiology; a text-book of microörganisms, general and applied. Philadelphia, P. Blakiston's son & co.[c1921] url p. 965.
- Native woody plants of the United States, their erosion-control and wildlife values. Washington, U. S. Govt. print. off., 1938. url , , , p. 146.
- North American trees (exclusive of Mexico and tropical United States) A handbook designed for field use, with plates and distribution maps. Ames, Iowa State University Press[1961] url p. 134.
- Pamphlets on forestry in California. [1900?- url , p. 146, p. 147, p. 202, p. 357.
- Pamphlets on silviculture. [1899- url , p. 74, p. 75.
- Shade and ornamental trees of California, by Merritt B. Pratt, state forester. Sacramento?1922? url p. 110, p. 110.
- The Great Basin naturalist. 40 1980 Provo, Utah: M.L. Bean Life Science Museum, Brigham Young University, 1939-1999. url p. 398.
- The Journal of heredity. Washington, etc., American Genetic Association url p. 523, p. 523.
- The Monthly bulletin of the State Commission of Horticulture. Sacramento, Calif.: The Commission, 1911-1919. url p. 257.
- University of California publications in agricultural sciences. Berkeley, University of California Press. url , , , , , , , p. 75, p. 78.
- Kuang Ko-zen & Lu An-ming. 1979. Juglandaceae. In: Kuang Ko-zen & Li Pei-chun, eds., Fl. Reipubl. Popularis Sin. 21: 6-44.
- McGranahan, G. H. and P. B. Catlin. 1987. Juglans rootstocks. In: R. C. Rom and R. F. Carlson, eds. 1987. Rootstocks for Fruit Crops. New York. Pp. 411-450.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Data Sources
Accessed through GBIF Data Portal February 01, 2008:
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- Oregon State University, Vascular Plant Collection
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2645914
- Catalogue of Life Accepted Name Code: ITS-19251
- GRIN Nomen Number: 20747
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19251
- International Plant Names Index (IPNI) ID: 131754-2
- Natural Heritage Network Species Identifier: PDJUG02040
- U.S.D.A. Plant Symbol: JUCAH
- Zipcode Zoo Species Identifier: 44784
Footnotes
- Anmin Lu, Donald E. Stone & L. J. Grauke "Juglandaceae". in Flora of China Vol. 4 Page 277. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Alan T. Whittemore & Donald E. Stone "Juglans". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- "Juglans hindsii". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
