Interesting Facts
Description
Family Cannabaceae
Herbs, annual
or perennial
, erect
or twining
, dioecious or sometimes monoecious, often with cystoliths
(a hard calcium carbonate
structure at base
of a hair) . Stems furrowed
or winged
. Stipules free
. Leaves alternate or opposite, palmately lobed
or compound
, sometimes simple
. Male inflorescences a bracteate
cymose
panicle. Male flowers: pedicellate
; sepals 5, free; petals absent; stamens 5, opposite sepals; filaments
short; anthers
2-loculed, dehiscent
by longitudinal
slits. Female inflorescences a bracteate spicate cyme much reduced in Cannabis, pendent or erect. Female flowers: sessile; calyx appressed
to ovary, membranous; petals absent; ovary 1-loculed; ovule solitary, pendulous from locule apex; style 2-parted, branches filiform
. Fruit an achene, covered by persistent
calyx; endosperm fleshy
; embryo curved
or spirally involute
.
Two genera and four species: N Africa, Asia, Europe, North America; two genera and four species (one endemic) in China.
Because all the Chinese species in this family
are cultivated and are often found naturalized
in disturbed
habitats
, it is difficult to know the true wild distributions.
Cannabaceae has sometimes been included
in Moraceae or Urticaceae but is now usually recognized as a distinct
family. The subfamily
Celtidoideae of Ulmaceae could possibly be included within Cannabaceae (see the discussion after the Ulmaceae family description
) .[1]
Genus Humulus
Herbs, taprooted annuals
or rhizomatous
perennials
, rightward-twining. Stems usually branched, armed
with rigid
2-branched, stalked
hairs
that facilitate climbing
. Leaves simple
; petioles
often twining
, with 2-branched hairs. Leaf blade
mostly cordate, palmately lobed
, sometimes unlobed; surfaces abaxially resin-dotted and×or gland-dotted. Inflorescences: staminate
inflorescences axillary
and terminal
, cymose
panicles, erect
to pendent, (10-) 20-100+-flowered, flowers small; pistillate
axillary, spikes or racemes
, flowers solitary or paired
, short-pedicellate, subtended by bracts and bracteoles. Staminate and pistillate flowers usually on different plants
. Achenes lenticular
or terete
, ensheathed by brownish or sometimes mottled
persistent
perianth; embryo coiled
. x
= 10.
Species 3: temperate
Northern Hemisphere.[2]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Urticanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Rosales
(
)
- Perleb, 1826
- Family:
Cannabaceae
(
)
- Augier, 1801 ex Martinov, 1820, nom. cons.
- hemp
- Subfamily:
Rhododendroideae
(
)
- Tribe:
Rhododendreae
(
)
- Genus:
Humulus
(
)
- C. Linnaeus, 1753
- Hop, houblon [Latin humulus, applied to hop plant]
- Specific epithet:
lupulus
- Botanical name: - Humulus lupulus 'Golden Showers'
- Specific epithet:
lupulus
- Genus:
Humulus
(
- Tribe:
Rhododendreae
(
- Subfamily:
Rhododendroideae
(
- Family:
Cannabaceae
(
- Order:
Rosales
(
- Superorder:
Urticanae
(
- Subclass:
Rosidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
A tentatively accepted name in the RHS Horticultural Database.
Similar Species
Members of the genus Humulus
ZipcodeZoo has pages for 19 species, subspecies, varieties, forms, and cultivars in this genus:
H. japonicus (Japanese Hop) · H. japonicus 'Variegatus' (Variegated Japanese Hops) · H. lupulus (Common Hop) · H. lupulus americanus (Common Hop) · H. lupulus L. var. lupuloides E.Small (Arizona Hops) · H. lupulus L. var. neomexicanus A.Nelson & Cockerell (Common Hop) · H. lupulus var. lupuloides (Arizona Hops) · H. lupulus var. lupulus (Common Hop) · H. lupulus var. neomexicanus (Common Hop) · H. lupulus var. pubescens (Common Hop) · H. lupulus 'Apollo' (Hop 'apollo') · H. lupulus 'Aureus' (Aureus Golden Hops) · H. lupulus 'Cascade' (Hops) · H. lupulus 'Fuggle' (Hops) · H. lupulus 'Hallertauer' (Hops) · H. lupulus 'Northern Brewer' (Hops) · H. lupulus 'Nugget' (Hops) · H. lupulus 'Sovereign' (Hop 'sovereign') · H. lupulus 'Sunbeam' (Golden Hops)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Chang Siushih. 1998. Cannaboideae. In: Chang Siushih & Wu Chengyih, eds., Fl. Reipubl. Popularis Sin. 23(1): 220224.
- Small, E. 1978. A numerical and nomenclatural analysis of morpho-geographic taxa of Humulus. Syst. Bot. 3: 37-76.
- Small, E. 1980. The relationships of hop cultivars and wild variants of Humulus lupulus. Canad. J. Bot. 58: 676-686.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 11, 2012.
Identifiers
- Zipcode Zoo Species Identifier: 4493120
Footnotes
- Zhengyi Wu, Zhe-Kun Zhou & Bruce Bartholomew "Cannabaceae". in Flora of China Vol. 5 Page 74. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Humulus". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
