Interesting Facts
Common Names
Common Names in English:
Allegheny Hawkweed, Panicled Hawkweed
Description
Family Compositae
The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.
Tribe Lactuceae
The Lactuceae are a tribe of closely related genera of the sunflower family that are easily recognized because the flowering heads are composed of wholly of ligulate florets that are usually 5-lobed. Another very distinguishing feature is the milky sap . Although not apparent without magnification, the pollen is distinctive in that the spines are more or less restricted to discrete ridges or flanges on the surface of the grain. In other members of the family the spines are distributed more or less evenly over the surface of the pollen grain . The pappus usually consists of scales or stiff hairs . -- Gerald D. Carr.
Genus Hieracium
Perennials
, (5 ) 20 150+ cm; taprooted (rootstocks
sometimes woody, branched; stolons produced
in some taxa) . Stems usually 1, usually erect
, usually branched distally, sometimes throughout, sometimes scapiform
, glabrous
or hairy
(induments
often complex
, see discussion) . Leaves basal, basal and cauline, or cauline; petiolate
or sessile; blades
mostly elliptic
, lanceolate, oblanceolate
, oblong
, or spatulate
, margins
entire, denticulate
, or dentate
[laciniate
to pinnatifid
] (faces
glabrous or hairy, induments often complex, see discussion) . Heads borne singly or in corymbiform
, paniculiform
, thyrsiform, umbelliform, or nearly racemiform
arrays. Peduncles (terminal
and axillary
) not inflated
, often bracteate
. Calyculi 0 or of 3 13( 16+), deltate to lanceolate or linear
bractlets
(in 1 2+ series; sometimes intergrading with phyllaries) . Involucres hemispheric
or campanulate
to cylindric
, 3 9[ 12+] mm diam. Phyllaries 5 21( 40+) in 2+ series, lanceolate to linear, subequal
to unequal (reflexed
in fruit), margins usually little, if at all, scarious
, apices obtuse
to acute or acuminate. Receptacles flat, pitted
, glabrous, epaleate. Florets 6 150+; corollas usually yellow, sometimes white or ochroleucous
, sometimes tinged with cyan
or red, rarely orange (then often drying scarlet or purplish) . Cypselae usually red-brown or black (tan in H. horridum), usually ± columnar
or prismatic
, sometimes ± urceolate
(slightly bulbous proximally and narrower distally) or nearly fusiform
, not distinctly beaked
, ribs
(or grooves
) usually 10, faces glabrous; pappi persistent
(fragile), of 20 80+, distinct
, white, sordid
, stramineous
, or rufous
, ± equal or unequal, barbellulate
bristles
in 1 2+ series. x = 9.
Species 250 1000+: North America, Mexico, Central America, South America, Europe, Asia, Africa.
Most plants
referable to Hieracium are apomictic (reproducing
from asexually produced seeds) . Apomictic reproduction often results in perpetuation of morphologic variants
at populational and regional levels. Temptation to name
such variants as species has proven irresistible to some botanists; upward of 9,000 species names
have been published in Hieracium. Circumscriptions of "species" in Hieracium seem to be more artificial than in most genera of composites
. Under these circumstances, I feel it would be irresponsible to recognize any infraspecific
taxa here. For the most part, the "species" recognized here are those that have been recognized in local and regional floras
of the past 50 or so years. Accepted names
and/or changes in synomymies for some species reflect changes suggested by J. H. Beaman (1990), A. Cronquist (1980), and E. G. Voss (1972 1996, vol.
3) . I have maintained a traditional generic
circumscription of Hieracium. Some other botanists exclude some species (including numbers 1 6 here) from Hieracium and treat them in Pilosella Hill
, which differs from Hieracium in having stolons often produced, ribs of cypselae slightly projecting
distally, and bristles of the pappi in one series. For Greenland, T. W. Böcher et al.
(1968) reported 18 species of Hieracium: 16 said to be endemic to Greenland plus H. alpinum Linnaeus (known from Greenland and Eurasia
) and H. groenlandicum Arvet-Touvet (= H. vulgatum Fries; known from Greenland and continental North America) . Following H. J. Scoggan (1978 1979, part 4), I have placed 15 names of Greenland "endemics" as synonyms of other names. The type of the 16th "endemic," H. trigonophorum Oskarsson, is probably conspecific
with that of H. alpinum. In my key
and descriptions
, "piloso-hirsute" refers to surfaces with scattered
to crowded, tapered, whiplike, straight or curly, smooth
to ± barbellate
hairs
mostly (0.5 ) 2 8( 15+) mm (sometimes called "setae") ; "stellate-pubescent" refers to surfaces with scattered to crowded, ± dendritically branched (often called, but seldom truly, "stellate") hairs mostly 0.05 0.2+ mm (such surfaces are sometimes described as "floccose") ; and "stipitate-glandular" refers to surfaces with scattered to crowded gland-tipped hairs mostly 0.2 0.8( 1.2+) mm.
Surfaces of stems, leaves, peduncles, and phyllaries may be glabrous or may bear one, two, or all three of the types of hairs mentioned here; other induments are seldom encountered in hieraciums of the flora area. Given the complexity of the reproductive modes
among the plants and the likelihood of misidentifications
of vouchers
, I have not included
chromosome numbers for species. Sexual hieraciums are usually diploids (2n = 18) and the apomictic hieraciums are usually triploids (2n = 27) .
[1]
Physical Description
Species Hieracium paniculatum
Plants 30-90 cm. Stems proximally usually glabrous
,
sometimes piloso-hirsute (hairs
3-8+ mm) and/or stellate-pubescent,
distally usually glabrous. Leaves: basal 0(-2), cauline 6-12+;
blades
elliptic
or oblanceolate
to lanceolate, 30-80(-150) ×
6-25(-35) mm, lengths
3-6+ times widths
, bases
cuneate, margins
usually
toothed
, sometimes denticulate
or entire, apices acute to acuminate,
faces
glabrous or piloso-hirsute (hairs 1-4+ mm). Heads (6-)
12-50+ in corymbiform
to paniculiform
arrays. Peduncles usually
glabrous, sometimes stipitate-glandular
. Calyculi: bractlets
8-13+. Involucres campanulate
to cylindric
, 6-7 mm.
Phyllaries
8-13+, apices acute to acuminate, abaxial
faces usually glabrous,
sometimes stipitate-glandular. Florets 8-20(-30) ; corollas
ochroleucous
to yellow, 5-8 mm. Cypselae columnar
, 2-2.5 mm;
pappi of 35-40+, stramineous
bristles
in ± 2 series,
4-5 mm. [source]
The type of Hieracium scribneri Small may have resulted from
a cross
between plants
of H. paniculatum and H. venosum
(A. Cronquist 1980). [source]
Habit: Forb/herb
Flowers: Bloom Period: July, August, September, October.
Habitat
Openings in forests ; (10-)500-800 m [2].
Typically found at an altitude of 0 to 1,092 meters (0 to 3,583 feet).[3]
Biology
Reproduction
Duration: Perennial
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Campanulanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Asterales
(
)
- Lindley, 1833
- Family:
Compositae
(
)
- Giseke, 1792, nom. cons., nom. alt.
- Subfamily:
Cichorioideae
(
)
- Tribe:
Lactuceae
(
)
- Subtribe:
Hieraciinae
(
)
- Genus:
Hieracium
(
)
- C. Linnaeus, 1753
- Hawkweed [said to be from Greek hierax, hawk]
- Specific epithet:
paniculatum
- Linnaeus, Sp. Pl. 2: 802. 1753.
- Botanical name: - Hieracium paniculatum Linnaeus, Sp. Pl. 2: 802. 1753.
- Specific epithet:
paniculatum
- Linnaeus, Sp. Pl. 2: 802. 1753.
- Genus:
Hieracium
(
- Subtribe:
Hieraciinae
(
- Tribe:
Lactuceae
(
- Subfamily:
Cichorioideae
(
- Family:
Compositae
(
- Order:
Asterales
(
- Superorder:
Campanulanae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000
Similar Species
Members of the genus Hieracium
ZipcodeZoo has pages for 163 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
H. acranthophorum (Tunugdliarfik Hawkweed) · H. acranthophorum var. acranthophorum (Tunugdliarfik Hawkweed) · H. acranthophorum var. isortoquense (Tunugdliarfik Hawkweed) · H. albiflorum (White Hawkweed) · H. alleghaniense (Alleghany Hawkweed) · H. alpinum (Alpine Hawkweed) · H. amitsokense (Royal Hawkweed) · H. angmagssalikense (Blackstyle Hawkweed) · H. argutum (Southern Hawkweed) · H. argutum var. argutum (Southern Hawkweed) · H. arvense (Perennial Sowthistle) · H. atramentarium (Hawkweed) · H. atratum (Polar Hawkweed) · H. aurantiacum (Orange Hawkweed) · H. bolanderi (Bolander Hawkweed) · H. caespitosum (Field Hawkweed) · H. canadense (Canada Hawkweed) · H. canadense var. canadense (Canadian Hawkweed) · H. canadense var. canadense Michx. (Canadian Hawkweed) · H. canadense var. divaricatum (Canadian Hawkweed) · H. canadense var. divaricatum Lepage (Canadian Hawkweed) · H. canadense var. fasciculatum (Canadian Hawkweed) · H. canadense var. subintegrum (Canadian Hawkweed) · H. canadense var. subintegrum Lepage (Canadian Hawkweed) · H. carneum (Huachuca Hawkweed) · H. cernuiforme (Hawkweed) · H. cynoglossoides (Hounds-Tongue Hawkweed) · H. devoldii (Devold's Hawkweed) · H. dutillyanum (Hawkweed) · H. eugenii (Neria Hawkweed) · H. fassettii (Fassett's Hawkweed) · H. fassettii Lepage var. wisconsinense Lepage (pro nm.) (Wisconsin Hawkweed) · H. fassettii var. fassettii (Fassett's Hawkweed) · H. fassettii var. mendicum (Fassett's Hawkweed) · H. fassettii var. wisconsinense (Wisconsin Hawkweed) · H. fendleri (Yellow Hawksbeard) · H. fendleri var. discolor (Yellow Hawkweed) · H. fendleri var. discolor Gray (Yellow Hawkweed) · H. fendleri var. fendleri (Fendler's Hawkweed) · H. fendleri var. fendleri Sch.Bip. (Fendler Hawkweed) · H. fendleri var. mogollense (Mogollon Hawkweed) · H. fendleri var. mogollense Gray (Mogollon Hawkweed) · H. fernaldii (Fernald's Hawkweed) · H. flagellare (Hawkweed) · H. flagellare var. amauracron (Hawkweed) · H. flagellare var. cernuiforme (Hawkweed) · H. flagellare var. flagellare (Hawkweed) · H. flagellare var. glatzense (Hawkweed) · H. flagellare var. pilosius (Hawkweed) · H. flagellare Willd. (pro sp.) var. flagellare Willd. (pro sp.) (Hawkweed) · H. floribundum (Hawkweed) · H. lactucella (Lindenleaf Rosemallow) · H. fuscatrum (Hawkweed) · H. gracile (Alpine Hawkweed) · H. gracile var. alaskanum (Alaska Hawkweed) · H. gracile var. alaskanum Zahn (Alaska Hawkweed) · H. gracile var. detonsum (Low Alpine Hawkweed) · H. gracile var. gracile (Slender Hawkweed) · H. gracile var. yukonense (Yukon Hawkweed) · H. gracile var. yukonense Porsild (Yukon Hawkweed) · H. greenei (Greene's Hawkweed) · H. greenii (Maryland Hawkweed) · H. groenlandicum (Greenland Hawkweed) · H. grohii (Groh's Hawkweed) · H. gronovii (Gronovis Hawkweed) · H. horridum (Prickly Hawkweed) · H. hyparcticum (Arctic Hawkweed) · H. inuloides (Butterfly Hawkweed) · H. ivigtutense (Holsteinsborg Hawkweed) · H. kalmii (Kalm's Hawkweed) · H. kalmii var. kalmii (Kalm's Hawkweed) · H. lachenalii (Common Hawkweed) · H. lactucella (Hawkweed) · H. laevigatum (Smooth Hawkweed) · H. lanatum (Hairy Hawkweed) · H. lemmonii (Lemmon's Hawkweed) · H. lividorubens (Cow Hawkweed) · H. lividorubens var. lividorubens (Cow Hawkweed) · H. lividorubens var. pseudostylum (Cow Hawkweed) · H. lividorubens var. subnudulum (Cow Hawkweed) · H. longiberbe (Long-Bearded Hawkweed) · H. longipilum (Hairy Hawkweed) · H. maculatum (Spotted Hawkweed) · H. marianum (Hawkweed) · H. megacephalon (Coastal Plain Hawkweed) · H. lactucella (Crevice Alumroot) · H. lactucella (Showy Goldeneye) · H. murorum (Wall Hawkweed) · H. musartutense (Musartut Hawkweed) · H. nepiocratum (Lindenows Fjord Hawkweed) · H. paniculatum (Allegheny Hawkweed) · H. lactucella (Littleflower Alumroot) · H. lactucella (Littleleaf Alumroot) · H. perpropinquum (Hawkweed) · H. pilosella (Mouse-Ear Hawkweed) · H. pilosella pseudosedunense (Mouse-Ear Hawkweed) · H. pilosella var. niveum (Mouseear Hawkweed) · H. pilosella var. pilosella (Mouseear Hawkweed) · H. piloselloides (King-Devil Hawkweed) · H. pilosius (Hawkweed)
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Further Reading
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- A preliminary list of the vascular flora of Allegheny County, Pennsylvania / by John A. Shaffer. Pittsburgh?: Carnegie Institute Museum?, 1901? url p. 71.
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- An illustrated flora of the northern United States, Canada and the British possessions: from Newfoundland to the parallel of the southern boundary of Virginia and from the Atlantic Ocean westward to the 102nd meridian / by Nathaniel Lord Britton and Hon. Addison Brown. New York: Scribner, 1913. url p. 328, p. 330.
- An illustrated guide to the flowering plants of the middle Atlantic and New England states (excepting the grasses and sedges) the descriptive text written in familiar language, by George T. Stevens. .. with more than 1800 illustrations from drawings by the author. New York, Dodd, Mead and company, 1910. url p. 621.
- Annals of the Carnegie Museum. [Pittsburgh]: Published by authority of the Board of Trustees of the Carnegie Institute, 1901- url p. 420, p. 71.
- Annotated list of the ferns and flowering plants of New York state, by Homer D. House. Albany, The University of the state of New York, 1924. url p. 670.
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Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Data Sources
Accessed through GBIF Data Portal November 23, 2007:
- Canadian Museum of Nature, Canadian Museum of Nature Herbarium
- Missouri Botanical Garden, Missouri Botanical Garden
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2658744
- Catalogue of Life Accepted Name Code: ITS-37718
- Global Biodiversity Information Facility Taxonkey: 13749891
- Globally Unique Identifier: urn:lsid:ipni.org:names:220886-1
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 37718
- Natural Heritage Network Species Identifier: PDAST4W120
- U.S.D.A. Plant Symbol: HIPA2
- Zipcode Zoo Species Identifier: 42986
Footnotes
- John L. Strother "Hieracium". in Flora of North America Vol. 19, 20 and 21 Page 219, 278, 279. Oxford University Press. Online at EFloras.org. [back]
- "Hieracium paniculatum". in Flora of North America Vol. 19, 20 and 21 Page 281, 287. Oxford University Press. Online at EFloras.org. [back]
- Mean = 339.290 meters (1,113.156 feet), Standard Deviation = 246.430 based on 356 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
