Common Names in English:
Allegheny Hawkweed, Panicled Hawkweed
The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.
The Lactuceae are a tribe of closely related genera of the sunflower family that are easily recognized because the flowering heads are composed of wholly of ligulate florets that are usually 5-lobed. Another very distinguishing feature is the milky sap . Although not apparent without magnification, the pollen is distinctive in that the spines are more or less restricted to discrete ridges or flanges on the surface of the grain. In other members of the family the spines are distributed more or less evenly over the surface of the pollen grain . The pappus usually consists of scales or stiff hairs . -- Gerald D. Carr.
, (5 ) 20 150+ cm; taprooted (rootstocks
sometimes woody, branched; stolons produced
in some taxa) . Stems usually 1, usually erect
, usually branched distally, sometimes throughout, sometimes scapiform
, see discussion) . Leaves basal, basal and cauline, or cauline; petiolate
or sessile; blades
, lanceolate, oblanceolate
, or spatulate
, or dentate
glabrous or hairy, induments often complex, see discussion) . Heads borne singly or in corymbiform
, thyrsiform, umbelliform, or nearly racemiform
arrays. Peduncles (terminal
) not inflated
, often bracteate
. Calyculi 0 or of 3 13( 16+), deltate to lanceolate or linear
(in 1 2+ series; sometimes intergrading with phyllaries) . Involucres hemispheric
, 3 9[ 12+] mm diam. Phyllaries 5 21( 40+) in 2+ series, lanceolate to linear, subequal
to unequal (reflexed
in fruit), margins usually little, if at all, scarious
, apices obtuse
to acute or acuminate. Receptacles flat, pitted
, glabrous, epaleate. Florets 6 150+; corollas usually yellow, sometimes white or ochroleucous
, sometimes tinged with cyan
or red, rarely orange (then often drying scarlet or purplish) . Cypselae usually red-brown or black (tan in H. horridum), usually ± columnar
, sometimes ± urceolate
(slightly bulbous proximally and narrower distally) or nearly fusiform
, not distinctly beaked
) usually 10, faces glabrous; pappi persistent
(fragile), of 20 80+, distinct
, white, sordid
, or rufous
, ± equal or unequal, barbellulate
in 1 2+ series. x = 9.
Species 250 1000+: North America, Mexico, Central America, South America, Europe, Asia, Africa.
Most plants referable to Hieracium are apomictic (reproducing from asexually produced seeds) . Apomictic reproduction often results in perpetuation of morphologic variants at populational and regional levels. Temptation to name such variants as species has proven irresistible to some botanists; upward of 9,000 species names have been published in Hieracium. Circumscriptions of "species" in Hieracium seem to be more artificial than in most genera of composites . Under these circumstances, I feel it would be irresponsible to recognize any infraspecific taxa here. For the most part, the "species" recognized here are those that have been recognized in local and regional floras of the past 50 or so years. Accepted names and/or changes in synomymies for some species reflect changes suggested by J. H. Beaman (1990), A. Cronquist (1980), and E. G. Voss (1972 1996, vol. 3) . I have maintained a traditional generic circumscription of Hieracium. Some other botanists exclude some species (including numbers 1 6 here) from Hieracium and treat them in Pilosella Hill , which differs from Hieracium in having stolons often produced, ribs of cypselae slightly projecting distally, and bristles of the pappi in one series. For Greenland, T. W. Böcher et al. (1968) reported 18 species of Hieracium: 16 said to be endemic to Greenland plus H. alpinum Linnaeus (known from Greenland and Eurasia ) and H. groenlandicum Arvet-Touvet (= H. vulgatum Fries; known from Greenland and continental North America) . Following H. J. Scoggan (1978 1979, part 4), I have placed 15 names of Greenland "endemics" as synonyms of other names. The type of the 16th "endemic," H. trigonophorum Oskarsson, is probably conspecific with that of H. alpinum. In my key and descriptions , "piloso-hirsute" refers to surfaces with scattered to crowded, tapered, whiplike, straight or curly, smooth to ± barbellate hairs mostly (0.5 ) 2 8( 15+) mm (sometimes called "setae") ; "stellate-pubescent" refers to surfaces with scattered to crowded, ± dendritically branched (often called, but seldom truly, "stellate") hairs mostly 0.05 0.2+ mm (such surfaces are sometimes described as "floccose") ; and "stipitate-glandular" refers to surfaces with scattered to crowded gland-tipped hairs mostly 0.2 0.8( 1.2+) mm. Surfaces of stems, leaves, peduncles, and phyllaries may be glabrous or may bear one, two, or all three of the types of hairs mentioned here; other induments are seldom encountered in hieraciums of the flora area. Given the complexity of the reproductive modes among the plants and the likelihood of misidentifications of vouchers , I have not included chromosome numbers for species. Sexual hieraciums are usually diploids (2n = 18) and the apomictic hieraciums are usually triploids (2n = 27) .
Species Hieracium paniculatum
Plants 30-90 cm. Stems proximally usually glabrous
sometimes piloso-hirsute (hairs
3-8+ mm) and/or stellate-pubescent,
distally usually glabrous. Leaves: basal 0(-2), cauline 6-12+;
to lanceolate, 30-80(-150) ×
6-25(-35) mm, lengths
3-6+ times widths
, sometimes denticulate
or entire, apices acute to acuminate,
glabrous or piloso-hirsute (hairs 1-4+ mm). Heads (6-)
12-50+ in corymbiform
arrays. Peduncles usually
glabrous, sometimes stipitate-glandular
. Calyculi: bractlets
8-13+. Involucres campanulate
, 6-7 mm.
8-13+, apices acute to acuminate, abaxial
faces usually glabrous,
sometimes stipitate-glandular. Florets 8-20(-30) ; corollas
to yellow, 5-8 mm. Cypselae columnar
, 2-2.5 mm;
pappi of 35-40+, stramineous
in ± 2 series,
4-5 mm. [source]
The type of Hieracium scribneri Small may have resulted from a cross between plants of H. paniculatum and H. venosum (A. Cronquist 1980). [source]
Flowers: Bloom Period: July, August, September, October.
Openings in forests ; (10-)500-800 m .
Typically found at an altitude of 0 to 1,092 meters (0 to 3,583 feet).
- Whittaker & Margulis,1978
- Haeckel, 1866
- Cavalier-Smith, 1981
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Brongniart, 1843
- Takhtajan, 1967
- Takhtajan Ex Reveal, 1992
- Lindley, 1833
- Family: Compositae () - Giseke, 1792, nom. cons., nom. alt.
- Order: Asterales () - Lindley, 1833
- Superorder: Campanulanae () - Takhtajan Ex Reveal, 1992
- Subclass: Asteridae () - Takhtajan, 1967
- Class: Spermatopsida () - Brongniart, 1843
- Subphylum: Euphyllophytina ()
- Phylum: Tracheophyta () - Sinnott, 1935 ex Cavalier-Smith, 1998 - Vascular Plants
- Subkingdom: Viridaeplantae () - Cavalier-Smith, 1981
- Kingdom: Plantae () - Haeckel, 1866 - Plants
Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000
Members of the genus Hieracium
ZipcodeZoo has pages for 163 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
H. acranthophorum (Tunugdliarfik Hawkweed) · H. acranthophorum var. acranthophorum (Tunugdliarfik Hawkweed) · H. acranthophorum var. isortoquense (Tunugdliarfik Hawkweed) · H. albiflorum (White Hawkweed) · H. alleghaniense (Alleghany Hawkweed) · H. alpinum (Alpine Hawkweed) · H. amitsokense (Royal Hawkweed) · H. angmagssalikense (Blackstyle Hawkweed) · H. argutum (Southern Hawkweed) · H. argutum var. argutum (Southern Hawkweed) · H. arvense (Perennial Sowthistle) · H. atramentarium (Hawkweed) · H. atratum (Polar Hawkweed) · H. aurantiacum (Orange Hawkweed) · H. bolanderi (Bolander Hawkweed) · H. caespitosum (Field Hawkweed) · H. canadense (Canada Hawkweed) · H. canadense var. canadense (Canadian Hawkweed) · H. canadense var. canadense Michx. (Canadian Hawkweed) · H. canadense var. divaricatum (Canadian Hawkweed) · H. canadense var. divaricatum Lepage (Canadian Hawkweed) · H. canadense var. fasciculatum (Canadian Hawkweed) · H. canadense var. subintegrum (Canadian Hawkweed) · H. canadense var. subintegrum Lepage (Canadian Hawkweed) · H. carneum (Huachuca Hawkweed) · H. cernuiforme (Hawkweed) · H. cynoglossoides (Hounds-Tongue Hawkweed) · H. devoldii (Devold's Hawkweed) · H. dutillyanum (Hawkweed) · H. eugenii (Neria Hawkweed) · H. fassettii (Fassett's Hawkweed) · H. fassettii Lepage var. wisconsinense Lepage (pro nm.) (Wisconsin Hawkweed) · H. fassettii var. fassettii (Fassett's Hawkweed) · H. fassettii var. mendicum (Fassett's Hawkweed) · H. fassettii var. wisconsinense (Wisconsin Hawkweed) · H. fendleri (Yellow Hawksbeard) · H. fendleri var. discolor (Yellow Hawkweed) · H. fendleri var. discolor Gray (Yellow Hawkweed) · H. fendleri var. fendleri (Fendler's Hawkweed) · H. fendleri var. fendleri Sch.Bip. (Fendler Hawkweed) · H. fendleri var. mogollense (Mogollon Hawkweed) · H. fendleri var. mogollense Gray (Mogollon Hawkweed) · H. fernaldii (Fernald's Hawkweed) · H. flagellare (Hawkweed) · H. flagellare var. amauracron (Hawkweed) · H. flagellare var. cernuiforme (Hawkweed) · H. flagellare var. flagellare (Hawkweed) · H. flagellare var. glatzense (Hawkweed) · H. flagellare var. pilosius (Hawkweed) · H. flagellare Willd. (pro sp.) var. flagellare Willd. (pro sp.) (Hawkweed) · H. floribundum (Hawkweed) · H. lactucella (Lindenleaf Rosemallow) · H. fuscatrum (Hawkweed) · H. gracile (Alpine Hawkweed) · H. gracile var. alaskanum (Alaska Hawkweed) · H. gracile var. alaskanum Zahn (Alaska Hawkweed) · H. gracile var. detonsum (Low Alpine Hawkweed) · H. gracile var. gracile (Slender Hawkweed) · H. gracile var. yukonense (Yukon Hawkweed) · H. gracile var. yukonense Porsild (Yukon Hawkweed) · H. greenei (Greene's Hawkweed) · H. greenii (Maryland Hawkweed) · H. groenlandicum (Greenland Hawkweed) · H. grohii (Groh's Hawkweed) · H. gronovii (Gronovis Hawkweed) · H. horridum (Prickly Hawkweed) · H. hyparcticum (Arctic Hawkweed) · H. inuloides (Butterfly Hawkweed) · H. ivigtutense (Holsteinsborg Hawkweed) · H. kalmii (Kalm's Hawkweed) · H. kalmii var. kalmii (Kalm's Hawkweed) · H. lachenalii (Common Hawkweed) · H. lactucella (Hawkweed) · H. laevigatum (Smooth Hawkweed) · H. lanatum (Hairy Hawkweed) · H. lemmonii (Lemmon's Hawkweed) · H. lividorubens (Cow Hawkweed) · H. lividorubens var. lividorubens (Cow Hawkweed) · H. lividorubens var. pseudostylum (Cow Hawkweed) · H. lividorubens var. subnudulum (Cow Hawkweed) · H. longiberbe (Long-Bearded Hawkweed) · H. longipilum (Hairy Hawkweed) · H. maculatum (Spotted Hawkweed) · H. marianum (Hawkweed) · H. megacephalon (Coastal Plain Hawkweed) · H. lactucella (Crevice Alumroot) · H. lactucella (Showy Goldeneye) · H. murorum (Wall Hawkweed) · H. musartutense (Musartut Hawkweed) · H. nepiocratum (Lindenows Fjord Hawkweed) · H. paniculatum (Allegheny Hawkweed) · H. lactucella (Littleflower Alumroot) · H. lactucella (Littleleaf Alumroot) · H. perpropinquum (Hawkweed) · H. pilosella (Mouse-Ear Hawkweed) · H. pilosella pseudosedunense (Mouse-Ear Hawkweed) · H. pilosella var. niveum (Mouseear Hawkweed) · H. pilosella var. pilosella (Mouseear Hawkweed) · H. piloselloides (King-Devil Hawkweed) · H. pilosius (Hawkweed)
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- Beaman, J. H. 1990. Revision of Hieracium (Asteraceae) in Mexico and Central America. Syst. Bot. Monogr. 29: 177.
- Fernald, M. L. 1943c. Notes on Hieracium. Rhodora 45: 317325.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Accessed through GBIF Data Portal November 23, 2007:
- Canadian Museum of Nature, Canadian Museum of Nature Herbarium
- Missouri Botanical Garden, Missouri Botanical Garden
- USDA PLANTS, USDA PLANTS Database
- Biodiversity Heritage Library NamebankID: 2658744
- Catalogue of Life Accepted Name Code: ITS-37718
- Global Biodiversity Information Facility Taxonkey: 13749891
- Globally Unique Identifier: urn:lsid:ipni.org:names:220886-1
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 37718
- Natural Heritage Network Species Identifier: PDAST4W120
- U.S.D.A. Plant Symbol: HIPA2
- Zipcode Zoo Species Identifier: 42986
- John L. Strother "Hieracium". in Flora of North America Vol. 19, 20 and 21 Page 219, 278, 279. Oxford University Press. Online at EFloras.org. [back]
- "Hieracium paniculatum". in Flora of North America Vol. 19, 20 and 21 Page 281, 287. Oxford University Press. Online at EFloras.org. [back]
- Mean = 339.290 meters (1,113.156 feet), Standard Deviation = 246.430 based on 356 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]