Hieracium atratum

• Interesting Facts
• Common Names
• Description
• Biology
• Taxonomy
• Similar Species
• More Info
• Further Reading
• Notes

Interesting Facts

Common Names

Description

Family Compositae

The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.

Tribe Lactuceae

The Lactuceae are a tribe of closely related genera of the sunflower family that are easily recognized because the flowering heads are composed of wholly of ligulate florets that are usually 5-lobed. Another very distinguishing feature is the milky sap . Although not apparent without magnification, the pollen is distinctive in that the spines are more or less restricted to discrete ridges or flanges on the surface of the grain. In other members of the family the spines are distributed more or less evenly over the surface of the pollen grain . The pappus usually consists of scales or stiff hairs . -- Gerald D. Carr.

Genus Hieracium

Perennials , (5 ) 20 150+ cm; taprooted (rootstocks sometimes woody, branched; stolons produced in some taxa) . Stems usually 1, usually erect , usually branched distally, sometimes throughout, sometimes scapiform , glabrous or hairy (induments often complex , see discussion) . Leaves basal, basal and cauline, or cauline; petiolate or sessile; blades mostly elliptic , lanceolate, oblanceolate , oblong , or spatulate , margins entire, denticulate , or dentate [laciniate to pinnatifid ] (faces glabrous or hairy, induments often complex, see discussion) . Heads borne singly or in corymbiform , paniculiform , thyrsiform, umbelliform, or nearly racemiform arrays. Peduncles (terminal and axillary ) not inflated , often bracteate . Calyculi 0 or of 3 13( 16+), deltate to lanceolate or linear bractlets (in 1 2+ series; sometimes intergrading with phyllaries) . Involucres hemispheric or campanulate to cylindric , 3 9[ 12+] mm diam. Phyllaries 5 21( 40+) in 2+ series, lanceolate to linear, subequal to unequal (reflexed in fruit), margins usually little, if at all, scarious , apices obtuse to acute or acuminate. Receptacles flat, pitted , glabrous, epaleate. Florets 6 150+; corollas usually yellow, sometimes white or ochroleucous , sometimes tinged with cyan or red, rarely orange (then often drying scarlet or purplish) . Cypselae usually red-brown or black (tan in H. horridum), usually ± columnar or prismatic , sometimes ± urceolate (slightly bulbous proximally and narrower distally) or nearly fusiform , not distinctly beaked , ribs (or grooves ) usually 10, faces glabrous; pappi persistent (fragile), of 20 80+, distinct , white, sordid , stramineous , or rufous , ± equal or unequal, barbellulate bristles in 1 2+ series. x = 9.

Species 250 1000+: North America, Mexico, Central America, South America, Europe, Asia, Africa.

Most plants referable to Hieracium are apomictic (reproducing from asexually produced seeds) . Apomictic reproduction often results in perpetuation of morphologic variants at populational and regional levels. Temptation to name such variants as species has proven irresistible to some botanists; upward of 9,000 species names have been published in Hieracium. Circumscriptions of "species" in Hieracium seem to be more artificial than in most genera of composites . Under these circumstances, I feel it would be irresponsible to recognize any infraspecific taxa here. For the most part, the "species" recognized here are those that have been recognized in local and regional floras of the past 50 or so years. Accepted names and/or changes in synomymies for some species reflect changes suggested by J. H. Beaman (1990), A. Cronquist (1980), and E. G. Voss (1972 1996, vol. 3) . I have maintained a traditional generic circumscription of Hieracium. Some other botanists exclude some species (including numbers 1 6 here) from Hieracium and treat them in Pilosella Hill , which differs from Hieracium in having stolons often produced, ribs of cypselae slightly projecting distally, and bristles of the pappi in one series. For Greenland, T. W. Böcher et al. (1968) reported 18 species of Hieracium: 16 said to be endemic to Greenland plus H. alpinum Linnaeus (known from Greenland and Eurasia ) and H. groenlandicum Arvet-Touvet (= H. vulgatum Fries; known from Greenland and continental North America) . Following H. J. Scoggan (1978 1979, part 4), I have placed 15 names of Greenland "endemics" as synonyms of other names. The type of the 16th "endemic," H. trigonophorum Oskarsson, is probably conspecific with that of H. alpinum. In my key and descriptions , "piloso-hirsute" refers to surfaces with scattered to crowded, tapered, whiplike, straight or curly, smooth to ± barbellate hairs mostly (0.5 ) 2 8( 15+) mm (sometimes called "setae") ; "stellate-pubescent" refers to surfaces with scattered to crowded, ± dendritically branched (often called, but seldom truly, "stellate") hairs mostly 0.05 0.2+ mm (such surfaces are sometimes described as "floccose") ; and "stipitate-glandular" refers to surfaces with scattered to crowded gland-tipped hairs mostly 0.2 0.8( 1.2+) mm. Surfaces of stems, leaves, peduncles, and phyllaries may be glabrous or may bear one, two, or all three of the types of hairs mentioned here; other induments are seldom encountered in hieraciums of the flora area. Given the complexity of the reproductive modes among the plants and the likelihood of misidentifications of vouchers , I have not included chromosome numbers for species. Sexual hieraciums are usually diploids (2n = 18) and the apomictic hieraciums are usually triploids (2n = 27) .

[1]

Physical Description

Habitat

Typically found at an altitude of 0 to 2,500 meters (0 to 8,202 feet).[2]

Biology

Reproduction

Duration: Perennial

Growth

Culture: Space 12-15" apart.

Sunlight: Sun Exposure: Full Sun .

Taxonomy

Notes

Name Status: Accepted Name . Latest taxonomic scrutiny: 17-Oct-2001

Place of publication : Nova Acta Regiae Soc. Sci. Upsal. 14:105. 1848

Name verified on 29-Apr-2003 by ARS Systematic Botanists. Last updated: 20-Jul-2004

Similar Species

Members of the genus Hieracium

ZipcodeZoo has pages for 163 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:

H. acranthophorum (Tunugdliarfik Hawkweed) · H. acranthophorum var. acranthophorum (Tunugdliarfik Hawkweed) · H. acranthophorum var. isortoquense (Tunugdliarfik Hawkweed) · H. albiflorum (White Hawkweed) · H. alleghaniense (Alleghany Hawkweed) · H. alpinum (Alpine Hawkweed) · H. amitsokense (Royal Hawkweed) · H. angmagssalikense (Blackstyle Hawkweed) · H. argutum (Southern Hawkweed) · H. argutum var. argutum (Southern Hawkweed) · H. arvense (Perennial Sowthistle) · H. atramentarium (Hawkweed) · H. atratum (Polar Hawkweed) · H. aurantiacum (Orange Hawkweed) · H. bolanderi (Bolander Hawkweed) · H. caespitosum (Field Hawkweed) · H. canadense (Canada Hawkweed) · H. canadense var. canadense (Canadian Hawkweed) · H. canadense var. canadense Michx. (Canadian Hawkweed) · H. canadense var. divaricatum (Canadian Hawkweed) · H. canadense var. divaricatum Lepage (Canadian Hawkweed) · H. canadense var. fasciculatum (Canadian Hawkweed) · H. canadense var. subintegrum (Canadian Hawkweed) · H. canadense var. subintegrum Lepage (Canadian Hawkweed) · H. carneum (Huachuca Hawkweed) · H. cernuiforme (Hawkweed) · H. cynoglossoides (Hounds-Tongue Hawkweed) · H. devoldii (Devold's Hawkweed) · H. dutillyanum (Hawkweed) · H. eugenii (Neria Hawkweed) · H. fassettii (Fassett's Hawkweed) · H. fassettii Lepage var. wisconsinense Lepage (pro nm.) (Wisconsin Hawkweed) · H. fassettii var. fassettii (Fassett's Hawkweed) · H. fassettii var. mendicum (Fassett's Hawkweed) · H. fassettii var. wisconsinense (Wisconsin Hawkweed) · H. fendleri (Yellow Hawksbeard) · H. fendleri var. discolor (Yellow Hawkweed) · H. fendleri var. discolor Gray (Yellow Hawkweed) · H. fendleri var. fendleri (Fendler's Hawkweed) · H. fendleri var. fendleri Sch.Bip. (Fendler Hawkweed) · H. fendleri var. mogollense (Mogollon Hawkweed) · H. fendleri var. mogollense Gray (Mogollon Hawkweed) · H. fernaldii (Fernald's Hawkweed) · H. flagellare (Hawkweed) · H. flagellare var. amauracron (Hawkweed) · H. flagellare var. cernuiforme (Hawkweed) · H. flagellare var. flagellare (Hawkweed) · H. flagellare var. glatzense (Hawkweed) · H. flagellare var. pilosius (Hawkweed) · H. flagellare Willd. (pro sp.) var. flagellare Willd. (pro sp.) (Hawkweed) · H. floribundum (Hawkweed) · H. lactucella (Lindenleaf Rosemallow) · H. fuscatrum (Hawkweed) · H. gracile (Alpine Hawkweed) · H. gracile var. alaskanum (Alaska Hawkweed) · H. gracile var. alaskanum Zahn (Alaska Hawkweed) · H. gracile var. detonsum (Low Alpine Hawkweed) · H. gracile var. gracile (Slender Hawkweed) · H. gracile var. yukonense (Yukon Hawkweed) · H. gracile var. yukonense Porsild (Yukon Hawkweed) · H. greenei (Greene's Hawkweed) · H. greenii (Maryland Hawkweed) · H. groenlandicum (Greenland Hawkweed) · H. grohii (Groh's Hawkweed) · H. gronovii (Gronovis Hawkweed) · H. horridum (Prickly Hawkweed) · H. hyparcticum (Arctic Hawkweed) · H. inuloides (Butterfly Hawkweed) · H. ivigtutense (Holsteinsborg Hawkweed) · H. kalmii (Kalm's Hawkweed) · H. kalmii var. kalmii (Kalm's Hawkweed) · H. lachenalii (Common Hawkweed) · H. lactucella (Hawkweed) · H. laevigatum (Smooth Hawkweed) · H. lanatum (Hairy Hawkweed) · H. lemmonii (Lemmon's Hawkweed) · H. lividorubens (Cow Hawkweed) · H. lividorubens var. lividorubens (Cow Hawkweed) · H. lividorubens var. pseudostylum (Cow Hawkweed) · H. lividorubens var. subnudulum (Cow Hawkweed) · H. longiberbe (Long-Bearded Hawkweed) · H. longipilum (Hairy Hawkweed) · H. maculatum (Spotted Hawkweed) · H. marianum (Hawkweed) · H. megacephalon (Coastal Plain Hawkweed) · H. lactucella (Crevice Alumroot) · H. lactucella (Showy Goldeneye) · H. murorum (Wall Hawkweed) · H. musartutense (Musartut Hawkweed) · H. nepiocratum (Lindenows Fjord Hawkweed) · H. paniculatum (Allegheny Hawkweed) · H. lactucella (Littleflower Alumroot) · H. lactucella (Littleleaf Alumroot) · H. perpropinquum (Hawkweed) · H. pilosella (Mouse-Ear Hawkweed) · H. pilosella pseudosedunense (Mouse-Ear Hawkweed) · H. pilosella var. niveum (Mouseear Hawkweed) · H. pilosella var. pilosella (Mouseear Hawkweed) · H. piloselloides (King-Devil Hawkweed) · H. pilosius (Hawkweed)

More Info

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Further Reading

• A new British flora: British wild flowers in their natural haunts / described by A. R. Horwood; with sixty-four plates in colour representing 350 different plants, from drawings by J. N. Fitch and many illustrations from photographs. London: Gresham, 1919. url p. 182.
• A new British flora; British wild flowers in their natural haunts, described by A. R. Horwood, with 64 plates in col. representing 350 different plants, from drawings by J. N. Fitch and many illus. from photo. London, Gresham Pub. Co., 1919. url p. 182.
• Botaniska notiser. Upsala: Wahlstro?m, 1857-c1980. url p. 182.
• Cybele Britannica: or British Plants and their geographical relations. London, 1847-59. url p. 453.
• English botany, or, Coloured figures of British plants / edited by John T. Boswell Syme; the popular portion by Mrs. Lankester; the figures by J. Sowerby, J. de C. Sowerby, J.W. Salter, and John Edward Sowerby. London: R. Hardwicke, 1863-1886. url p. 935.
• English botany, or, coloured figures of British plants / edited by John T. Boswell Syme; the popular portion by Mrs. Lankester; the figures by J. Sowerby, J. de C. Sowerby, J.W. Salter, and John Edward Sowerby. London: G. Bell, 1873. url p. 174, p. 176.
• Journal of botany, British and foreign. 42 1904 London: Robert Hardwicke, 1863-1942. url p. 273.
• Manual of British botany, containing the flowering plants and ferns. Arranged according to the natural orders. London, J. Van Voorst, 1856. url .
• Manual of British botany: containing the flowering plants and ferns arranged according to the natural orders / by Charles Cardale Babington. London: John Van Voorst, 1856. url p. 201.
• Meddelelser om Grønland. København, C. A. Reitzels Forlag, 1879-1979. url p. 201, p. 259, p. 35.
• Notes from the Royal Botanic Garden, Edinburgh. 2 1902 Edinburgh: H. M. Stationery Off. url p. 202.
• The Annals of Scottish natural history. Edinburgh: David Douglas, 1892-1911. url p. 245.
• The Eskimo tribes; their distribution and characteristics, especially as regards language, with a comparative vocabulary and sketch-map. Copenhagen1887-91. url p. 35.
• The student's flora of the British Islands / by J.D. Hooker. London: Macmillan, 1884. url p. 234.
• Transactions and proceedings of the Botanical Society of Edinburgh. [Edinburgh]: The Society, 1891-1970. url p. 73.
• Beaman, J. H. 1990. Revision of Hieracium (Asteraceae) in Mexico and Central America. Syst. Bot. Monogr. 29: 177.
• Fernald, M. L. 1943c. Notes on Hieracium. Rhodora 45: 317325.

Notes

Contributors

• Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.

Data Sources

Accessed through GBIF Data Portal December 02, 2007:

• Biologiezentrum der Oberoesterreichischen Landesmuseen, Biologiezentrum Linz
• Bundesamt für Naturschutz / Zentralstelle für Phytodiversität Deutschland, Bundesamt fuer Naturschutz / Zentralstelle fuer Phytodiversitaet Deutschland
• Natural History Museum, University of Oslo, Vascular Plant Herbarium, Oslo
• Natural History Museum, University of Oslo, Vascular Plants, Field notes, Oslo
• Tiroler Landesmuseum Ferdinandeum, Tiroler Landesmuseum Ferdinandeum
• inatura - Erlebnis Naturschau Dornbirn, inatura - Erlebnis Naturschau Dornbirn

Identifiers

• Biodiversity Heritage Library NamebankID: 2667468
• Catalogue of Life Accepted Name Code: ITS-503008
• Global Biodiversity Information Facility Taxonkey: 13741354
• Globally Unique Identifier: urn:lsid:ipni.org:names:215116-1
• GRIN Nomen Number: 433133
• Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 503008
• U.S.D.A. Plant Symbol: HIAT2
• Zipcode Zoo Species Identifier: 41967

Footnotes

1. John L. Strother "Hieracium". in Flora of North America Vol. 19, 20 and 21 Page 219, 278, 279. Oxford University Press. Online at EFloras.org. [back]
2. Mean = 1,219.380 meters (4,000.591 feet), Standard Deviation = 485.400 based on 50 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]

©2004-2012 David Stang. All rights reserved.