Interesting Facts
Description
Family Malvaceae
Herbs, shrubs
, or less often trees
; indumentum usually with peltate scales
or stellate
hairs
. Leaves alternate, stipulate
, petiolate
; leaf blade
usually palmately veined, entire or various lobed
. Flowers solitary, less often in small cymes or clusters
, axillary
or subterminal
, often aggregated into terminal
racemes
or panicles, usually conspicuous
, actinomorphic
, usually bisexual
(unisexual
in Kydia) . Epicalyx
often present, forming an involucre around calyx, 3- to many lobed. Sepals 5, valvate
, free
or connate
. Petals 5, free, contorted, or imbricate, basally adnate
to base
of filament
tube
. Stamens usually very many, filaments connate into tube; anthers
1-celled. Pollen spiny
. Ovary superior, with 2-25 carpels, often separating from one another and from axis; ovules 1 to many per locule; style as many or 2 Ã as many as pistils, apex branched or capitate. Fruit a loculicidal capsule or a schizocarp, separating into individual mericarps, rarely berrylike when mature
(Malvaviscus) ; carpels sometimes with an endoglossum (a crosswise projection from back wall of carpel to make it almost completely septate
. Seeds often reniform
, glabrous
or hairy
, sometimes conspicuously so.
About 100 genera and ca.
1000 species: tropical
and temperate regions
of N and S Hemisphere; 19 genera (four introduced
) and 81 species (24 endemic, 16 introduced) in China.
Molecular studies have shown that the members
of the Bombacaceae, Malvaceae, Sterculiaceae, and Tiliaceae form a very well-defined monophyletic group that is divided
into ten also rather well-defined clades, only two of which correspond to the traditional families Bombacaceae and Malvaceae. Some of the remaining groups are included
entirely within either of the remaining families but others cut
across the traditional divide between the Sterculiaceae and Tiliaceae. A majority of authors
, most notably Bayer and Kubitzki (Fam. Gen. Vasc. Pl. 5: 225-311. 2003), has favored including everything within a greatly enlarged Malvaceae, and treating the individual clades as subfamilies. The alternative view
is that the individual clades should be treated as a series of ten families: Bombacaceae (Bombacoideae), Brownlowiaceae (Brownlowioideae), Byttneriaceae (Byttnerioideae), Durionaceae (Durionoideae), Helicteraceae (Helicteroideae), Malvaceae (Malvoideae), Pentapetaceae (Dombeyoideae), Sparrmanniaceae (Grewioideae), Sterculiaceae (Sterculioideae), and Tiliaceae (Tilioideae) (Cheek in Heywood et al.
, Fl.
Pl. Fam. World. 201-202. 2007) . For the present treatment, we prefer to retain the familiar, traditional four families, so as to maintain continuity
with the treatments in FRPS, and to await a consensus on the two alternative strategies for dealing with the very widely accepted clades.
The traditional Malvaceae coincides exactly with one of the major clades. The only possible problem is the relationship
with the Bombacaceae, which also has primarily 1-loculed anthers, and some authorities have suggested that the Bombacaceae should be included within the Malvaceae.
Members of the Malvaceae are important as fiber crops
(particularly cotton, Gossypium) . Young leaves of many species can be used as vegetables, and species of Abelmoschus and Hibiscus are grown as minor food crops. Many species have attractive flowers and an ever-increasing selection is grown as ornamentals
. Several have been cultivated for a very long time, particularly species of Hibiscus, and some of these are not known in the wild.[1]
Genus Gossypium
Herbs annual
or perennial
, sometimes shrubs
; all parts irregularly dotted
with dark oil glands
. Leaf blades
usually palmately 3-9-lobed, rarely entire. Flowers solitary, subterminal
. Pedicel not articulated, usually with glands
below insertion
of epicalyx
lobes
. Epicalyx lobes 3(-7), foliaceous
, glandular
, free
or connate
at base
, entire or toothed
to deeply laciniate
. Calyx cup-shaped, nearly truncate
to 5-lobed. Corolla white or yellow, sometimes with purple center, large; petals 5, apex rounded
. Staminal
column with many anthers
along entire length, apex truncate. Ovary 3-5-loculed; ovules 2 to many per locule; style short, rod-shaped, stigma clavate
, 5-grooved. Capsule globose
or ellipsoid
, loculicidally dehiscent
. Seeds globose, densely white long woolly
, mixed with short hairs
or without short hairs.
About 20 species: tropical
and subtropical
regions; four species (all introduced
) in China.[2]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Malvanae
(
)
- Takhtajan, 1967
- Order:
Malvales
(
)
- Dumortier, 1829
- Family:
Malvaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- mallows, mauves
- Subfamily:
Malvoideae
(
)
- Tribe:
Gossypieae
(
)
- Genus:
Gossypium
(
)
- C. Linnaeus, 1753
- Cotton
- Specific epithet:
turneri
- Fryxell
- Botanical name: - Gossypium turneri Fryxell
- Specific epithet:
turneri
- Fryxell
- Genus:
Gossypium
(
- Tribe:
Gossypieae
(
- Subfamily:
Malvoideae
(
- Family:
Malvaceae
(
- Order:
Malvales
(
- Superorder:
Malvanae
(
- Subclass:
Rosidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author
: Fryxell Publication
: MadroƱo 25: 155 (-156) 1978
Place of publication: Madroño 25:155, t. 1. 1978
Name
verified on 10-Aug-1999 by ARS Systematic Botanists. Last
updated: 05-Nov-2001
Similar Species
Members of the genus Gossypium
ZipcodeZoo has pages for 23 species, subspecies, varieties, forms, and cultivars in this genus:
G. arboreum (China Cotton) · G. australe (Australian Native Cotton) · G. barbadense (Cotton) · G. barbadense var. breve (American-Egyptian Cotton) · G. barbadense var. deltaicum (American-Egyptian Cotton) · G. barbadense var. nubarense (American-Egyptian Cotton) · G. barbadense var. peruvianum (Peru Tree Cotton) · G. barbadense var. strictiflorum (American-Egyptian Cotton) · G. capitis-viridis (Cape Verde Cotton) · G. darwinii (Darwins Cotton) · G. herbaceum (Cotton) · G. herbaceum 'Erlene's Green' (Cotton) · G. hirsutum (American Cotton) · G. hirsutum f. rubrocoloratum (American Upland Cotton) · G. hirsutum L. var. hirsutum OFF. (Upland Cotton) · G. hirsutum var. harlandianum (American Upland Cotton) · G. hirsutum var. hirsutum (Upland Cotton) · G. hirsutum var. marie-galante (Marie Galante Cotton) · G. hirsutum var. praticolum (American Upland Cotton) · G. hirsutum 'Betica' (Upland Cotton 'betica') · G. sturtianum (Sturt's Desert Rose) · G. thurberi (Thurber's Cotton) · G. tomentosum (Hawai'i Cotton)
More Info
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Further Reading
- Fryxell, P. A. 1979. The natural history of the cotton tribe. (Nat Hist Cotton) 58.
- Fryxell, P. A. 1992. A revised taxonomic interpretation of Gossypium L. (Malvaceae). Rheedea 2:140.
- Feng Kuo-mei. 1984. Malvaceae. In: Feng Kuo-mei, ed., Fl. Reipubl. Popularis Sin. 49(2): 1-102.
- Feng Kuo-mei. 1984. Malvaceae. In: Feng Kuo-mei, ed., Fl. Reipubl. Popularis Sin. 49(2): 1-102.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 11, 2012.
Data Sources
Accessed through GBIF Data Portal November 20, 2007:
- The New York Botanical Garden, Vascular Plant Type Specimens
- UNIBIO, IBUNAM, Tipos de plantas vasculares
Identifiers
- Biodiversity Heritage Library NamebankID: 1
- Global Biodiversity Information Facility Taxonkey: 15602035
- Globally Unique Identifier: urn:lsid:ipni.org:names:559855-1
- GRIN Nomen Number: 310815
- International Plant Names Index (IPNI) ID: 559855-1
- Zipcode Zoo Species Identifier: 1082477
Footnotes
- Ya Tang, Michael G. Gilbert & Laurence J. Dorr "Malvaceae". in Flora of China Vol. 12 Page 240, 264,299, 302. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Gossypium". in Flora of China Vol. 12 Page 264, 296. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
