Common Names
Common Names in English:
Copper Beech, European Beech
Description
Family Fagaceae
Trees
or rarely shrubs
, monoecious, evergreen
or deciduous. Stipules usually early deciduous. Leaves alternate, sometimes false-whorled in Cyclobalanopsis. Inflorescences unisexual
or androgynous with female cupules at the base
of an otherwise male inflorescence. Male inflorescences a pendulous head
or erect
or pendulous catkin, sometimes branched; flowers in dense cymules
. Male flower: sepals 4-6(-9), scalelike, connate
or distinct
; petals absent; filaments
filiform
; anthers
dorsifixed
or versatile, opening by longitudinal
slits; with or without a rudimentary
pistil. Female inflorescences of 1-7 or more flowers subtended individually or collectively by a cupule formed from numerous
fused bracts, arranged individually or in small groups along an axis or at base of an androgynous inflorescence or on a separate axis. Female flower: perianth 1-7 or more; pistil 1; ovary inferior, 3-6(-9) -loculed; style
and carpels as many as locules; placentation axile
; ovules 2 per locule. Fruit a nut. Seed usually solitary by abortion
(but may be more than 1 in Castanea, Castanopsis, Fagus, and Formanodendron), without endosperm; embryo large.
Seven to 12 genera (depending on interpretation) and 900-1000 species: worldwide except for tropical
and S Africa; seven genera and 294 species (163 endemic, at least three introduced
) in China.
Many species are important timber trees. Nuts of Fagus, Castanea, and of most Castanopsis species are edible, and oil
is extracted from nuts of Fagus. Nuts of most species of this family
contain copious
amounts of water soluble tannin. Members
of the Fagaceae are the main element
of both broad-leaved evergreen and mixed mesophytic
forests
from 500-3200 m.
[1]
Genus Fagus
Trees
, winter-deciduous. Terminal
buds present, long, tapered in maturity, all scales
imbricate. Leaves: stipules prominent
on new growth, soon deciduous. Leaf blade
thin, secondary veins unbranched, ± parallel, extending to margin
, each vein
ending in acute or obscure
tooth
. Inflorescences unisexual
, axillary
in new growth leaves; staminate
inflorescence lax, loosely capitate cluster
of flowers; pistillate
inflorescence short, stiff, cupule 1, terminal. Staminate flowers
: sepals connate
; stamens 6-16; pistillode
typically absent. Pistillate flowers 2 per cupule; sepals distinct
; carpels and styles
3. Fruits: maturation
in 1st year following pollination; cupule 4-valved, valves
distinct, ±completely enclosing nuts until maturity, prickly, prickles stout, unbranched, short, not obscuring surface of cupule, internal valves absent; nuts 2 per cupule, sharply 3-angled, slightly winged
. x
= 12.
Species 8-10: temperate
, subtropical
, and montane
tropical
forests
, North America (e United
States), Mexico, Europe, Asia.[2]
Habitat
Typically found at an altitude of 0 to 2,500 meters (0 to 8,202 feet).[3]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Hamamelididae
(
)
- Takhtajan, 1967
- Superorder:
Faganae
(
)
- (Engler, 1892) Takhtajan, 1997
- Order:
Fagales
(
)
- Engler, 1892
- Family:
Fagaceae
(
)
- Dumortier, 1829
- Beech Family
- Subfamily:
Amygdaloideae
(
)
- Tribe:
Exochordeae
(
)
- Genus:
Fagus
(
)
- Linnaeus, Sp. Pl. 2: 997. 1753; Gen. Pl. ed. 5, 432, 1754.
- Beech [Classical Latin name, from Greek figos, an oak with edible acorns, probably from Greek fagein, to eat]
- Specific epithet:
sylvatica f. purpurea
- (Aiton) C.K.Schneid.
- Botanical name: - Fagus sylvatica f. purpurea (Aiton) C.K.Schneid.
- Specific epithet:
sylvatica f. purpurea
- (Aiton) C.K.Schneid.
- Genus:
Fagus
(
- Tribe:
Exochordeae
(
- Subfamily:
Amygdaloideae
(
- Family:
Fagaceae
(
- Order:
Fagales
(
- Superorder:
Faganae
(
- Subclass:
Hamamelididae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Fagus sylvatica var. atropunicea Weston
Similar Species
Members of the genus Fagus
There are approximately 202 species in this genus. Here are just 100 of them:
F. alpina · F. americana · F. antarctica · F. antarctica var. bicrenata · F. antarctica var. subalpina · F. antarctica var. uliginosa · F. antipofi · F. antipofii · F. apiculista · F. aspleniifolia · F. attenuata · F. beluloides · F. betuloides · F. blairii · F. carroni · F. carronii · F. castanea L. var. dentata Marshall · F. chienii · F. cliffortioides · F. cochinchinensis · F. crenata (Japanese Beech) · F. crenata 'Mount Fuji' · F. cunninghamii · F. decurrens · F. deucalionis · F. dombeyi · F. dubia · F. engleriana · F. ferruginea · F. ferruginea var. caroliniana · F. forsteri · F. fusca · F. glandulosa · F. glauca · F. glutinosa · F. grandiflora · F. grandifolia (American Beech) · F. grandifolia Ehrh. var. caroliniana (Loudon) Fernald · F. grandifolia f. pubescens · F. grandifolia grandifolia · F. grandifolia mexicana · F. grandifolia Ehrh. forma pubescens Fernald & Rehder · F. grandifolia subsp. mexicana · F. gunnii · F. hayatae · F. hohenackerana · F. japonica · F. japonica var. japonica · F. japonica var. multinervis · F. japoniciformis · F. longepetiolata · F. longipetiolata · F. longipetiolata f. clavata · F. longipetiolata f. longipetiolata · F. longipetiolata forma clavata · F. longipetiolata forma longipetiolata · F. lucida · F. lutea · F. menziesii · F. minima · F. moorei · F. multinervis · F. nervosa · F. nitida · F. obligua · F. obliqua · F. obliqua var. dentatosquamata · F. obliqua var. macrocarpa · F. obliqua var. valdiviana · F. orientale · F. orientalis (Oriental Beech) · F. pacifica · F. palaeojaponica · F. pliocaenica · F. pumilio · F. pumilo · F. quercifolia · F. silesiaca · F. silvatica · F. solandri · F. stuxbergi · F. stuxbergii · F. sylvatica (Fern-Leaf Beech) · F. sylvatica 'Albomarginata' · F. sylvatica 'Albovariegata' · F. sylvatica 'Ansorgei' (Ansorge European Beech) · F. sylvatica 'Argenteomarmorata' · F. sylvatica 'Asplenifolia' (Fernleaf European Beech) · F. sylvatica 'Atropunicea' (Purple European Beech) · F. sylvatica L. 'Atropunicea Macrophylla' · F. sylvatica Atropurpurea Group (European Beech) · F. sylvatica L. 'Atropurpurea Pendula' · F. sylvatica 'Aurea Pendula' (Weeping European Beech) · F. sylvatica 'Aureomarginata' · F. sylvatica var. Beth Dwarf (Beth Dwarf European Beech) · F. sylvatica 'Bicolor Sartini' · F. sylvatica 'Birr Zebra' · F. sylvatica 'Black Swan' (European Beech) · F. sylvatica 'Bornyensis' · F. sylvatica 'Brathay Purple'
More Info
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- Search for Scholarly Articles: Google Scholar
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- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Judd, W.S., Campbell, C.S., Kellog, E.A. and Donoghue, M.J. (2002): Plant Systematics: a phylogenetic approach, Sinauer, Sunderland, Mass.
- Huang Chengchiu, Chang Yongtian, Hsu Yongchun & Jen Hsienwei. 1998. Fagaceae. In: Chun Woonyong & Huang Chengchiu, eds. Fl. Reipubl. Popularis Sin. 22: 1-332.
- Cooper, A. W. and E. P. Mercer. 1977. Morphological variation in Fagus grandifolia Ehrh. in North Carolina. J. Elisha Mitchell Sci. Soc. 93: 136-149.
- Hardin, J. W. and G. P. Johnson. 1985. Atlas of foliar surface features in woody plants, VIII. Fagus and Castanea (Fagaceae) of eastern North America. Bull. Torrey Bot. Club 112: 11-20.
- Rehder, A. J. 1907. Some new or little known forms of New England trees. Rhodora 9: 109-116.
Notes
Contributors
- Brands, S.J. (comp.) 1989-2006. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed April 19, 2007.
- Global Biodiversity Information Facility. Accessed July 21, 2007. http://www.gbif.org Mediated distribution data from provider.
Identifiers
- Zipcode Zoo Species Identifier: 646284
Footnotes
- Chengjiu Huang, Yongtian Zhang & Bruce Bartholomew "Fagaceae". in Flora of China Vol. 4 Page 314. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Haining Qin & Peter Fritsch "Fagus". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- Mean = 361.000 meters (1,184.383 feet), Standard Deviation = 473.220 based on 3,697 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
