Overview
Interesting Facts
- Bonesets were once used in folk medicine for setting broken bones, thus the common name .
Common Names
Common Names in English:
Hyssop-Leaf Thoroughwort, Hyssop-Leaved Boneset, Hyssopleaf Thoroughwort
Description
Family Compositae
The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.
Genus Eupatorium
Perennials
, 30-200 cm. Stems erect
, usually not branched proximal
to arrays of heads
(from caudices or rhizomes) . Leaves mostly cauline; usually opposite (rarely whorled
, distal sometimes alternate) ; petiolate
or sessile; blades
usually 3-nerved from or distal to bases
, or pinnately nerved, mostly deltate or ovate
to lanceolate or linear
(and intermediate shapes
, sometimes elliptic
, oblong
, rhombic
, or suborbiculate, sometimes pinnatifid
, 1-2-pinnately, ternately, or palmately lobed
), ultimate
margins
entire or toothed
, faces
glabrous
or puberulent
, pubescent
, scabrous
, or setulose
, usually gland-dotted. Heads discoid
, in corymbiform
or diffuse
to dense, paniculiform
arrays. Involucres obconic to ellipsoid
, 1-3(-5+) mm diam. Phyllaries persistent
, 7-15+ in 2-3(-4+) series, (usually green) 2-3-nerved, or not notably nerved, or pinnately nerved, elliptic, lanceolate, oblong, or obovate
, usually unequal, sometimes ± equal (margins scarious
, hyaline
, apices rounded
to acute or acuminate sometimes mucronate
, faces usually puberulent or villous
, usually gland-dotted, rarely glabrous) . Receptacles flat or convex
, epaleate. Florets (3-) 5(-15+) ; corollas usually white, rarely pinkish, throats
funnelform
to campanulate
, lobes
5, triangular; styles: bases sometimes enlarged, usually puberulent (glabrous in E. capillifolium), branches mostly filiform
. Cypselae (brownish to black) prismatic
, 5-ribbed, usually glabrous, usually gland-dotted; pappi persistent, of 20-50 (whitish) barbellulate
bristles
in 1 series. x
= 10.
Species 41+ (24 species, including 2 hyb: e North America, Europe, e Asia.
Species identification within Eupatorium is sometimes complicated; polyploidy and apomixis have contributed to the complications. Some species include both sexual diploid and apomictic polyploid plants
or populations. V. I. Sullivan (1972) made important contributions to understanding Eupatorium in North America by showing that some fairly distinct
, sexual diploid species may include apomictic polyploid plants or populations that do not differ greatly from the diploids. Other apomictic polyploids appear to be intermediate morphologically between pairs of diploid or diploid/polyploid species and were proposed by Sullivan to have originated from interspecific
hybridization. Distinction and level of recognition of hybrid apomictic taxa have a large arbitrary
component
, in part because some apomicts appear to be ephemeral
and others may be relatively stable and in part because differences in the relative genomic contributions of the progenitors through dosage effects or backcrossing may affect whether an apomict is morphologically distinctive or part of a continuous series of variation
.[1]
Physical Description
Species Eupatorium hyssopifolium
Perennials
, 50-100+ cm. Stems (from short caudices
or rhizomes) single, sparsely branched distally, pubescent
throughout.
Leaves usually opposite or whorled
(distal sometimes alternate,
spreading
or horizontal) ; simple
, sessile; blades
3-nerved from
bases
(laterals
sometimes weak), lance-linear
, lance-oblong, or linear
,
20-60 × 2-15 mm (lengths
mostly 6-40 times widths
), bases cuneate,
margins
entire, laciniate-serrate, or serrate, apices acute, faces
scabrous
(at least abaxial
). Heads in corymbiform
arrays.
Phyllaries 8-10 in 2-3 series, elliptic
to oblong
, 1.5-5 ×
1-1.5 mm, apices obtuse to acute (not mucronate
), abaxial faces pubescent
throughout (more densely distally). Florets 5; corollas 3-3.5
mm.
Cypselae 2-3 mm; pappi of 20-30 bristles
3.5-4
mm. [source]
Eupatorium hyssopifolium includes both diploid and polyploid
cytotypes
and presents a complex
situation taxonomically. The diploids
are placed here in var. hyssopifolium, which also includes
polyploids
and is characterized by relatively narrow leaves. Variety
laciniatum has broader leaves and molecular data suggest that
it arose through hybridization between var. hyssopifolium and
E. serotinum; the name
E. torreyanum has been applied
to plants
of similar morphology, but these are hybrid derivatives
of E. serotinum and E. mohrii. It has also been suggested
that var. hyssopifolium has hybridized with E. album
var. album to form E. saltuense, and with diploids
of E. linearifolium to form polyploid populations included
here under E. linearifolium. Eupatorium hyssopifolium
is characterized by a tendency for the leaves to be whorled, in 3s
and 4s, and for axillary buds to produce
leaves without expansion
of shoots
; the combination
of whorled leaves and suppressed
axillary
shoots gives the nodes a somewhat tufted
or verticillate
appearance
.
[source]
Habit: Forb/herb
Flowers: Bloom Period: July, August, September, October, November. • Flower Color: near white, white
Size/Age/Growth
Size: 12-18" tall.
Landscaping
Landscape Uses: Back border . Cut flowers. • Care: Deer resistant. Tolerant of coastal conditions. Shear after blooming. Follow a regular watering schedule during the first growing season to establish a deep, extensive root system . Remove old flowers to prolong flowering. Cut back to the ground at the end of the season .
Habitat
Typically found at an altitude of 0 to 1,092 meters (0 to 3,583 feet).[2]
Biology
Reproduction
Duration: Perennial
Growth
Soil: Must have well-drained soil. Tolerates clay soil . Prefers humusy soil.
Sunlight: Sun Exposure: Full Sun .
Moisture: Water Requirements: Water regularly, when top 3 in. of soil is dry.
Temperature: Cold Hardiness: 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b, 9a, 9b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Campanulanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Asterales
(
)
- Lindley, 1833
- Family:
Compositae
(
)
- Giseke, 1792, nom. cons., nom. alt.
- Subfamily:
Asteroideae
(
)
- Tribe:
Eupatorieae
(
)
- Subtribe:
Eupatoriinae
(
)
- Genus:
Eupatorium
(
)
- C. Linnaeus, 1753
- [For Mithridates Eupator, King of Pontus, 132-63 B.C.]
- Specific epithet:
hyssopifolium
- L.
- Variety:
has broader leaves and molecular data suggest that
- Botanical name: - Eupatorium hyssopifolium L.
- Variety:
has broader leaves and molecular data suggest that
- Specific epithet:
hyssopifolium
- L.
- Genus:
Eupatorium
(
- Subtribe:
Eupatoriinae
(
- Tribe:
Eupatorieae
(
- Subfamily:
Asteroideae
(
- Family:
Compositae
(
- Order:
Asterales
(
- Superorder:
Campanulanae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Uncasia Hyssopifolia
Notes
Name
Status: Accepted Name
.
Comment: Data Providers: New Zealand Plant Name Database, IPNI, Tropicos.
GCC LSID: urn
:lsid:compositae.org:names:E365EAD7-D75E-4A72-84C1-E7D0EA64C236
Last scrutiny: 17-Aug-09
Similar Species
Members of the genus Eupatorium
ZipcodeZoo has pages for 93 species, subspecies, varieties, forms, and cultivars in this genus:
E. album (White Thoroughwort) · E. album var. album (White Thoroughwort) · E. album var. album L. (White Thoroughwort) · E. album var. glandulosum (White Thoroughwort) · E. album var. subvenosum (White Thoroughwort) · E. altissimum (Tall Boneset) · E. anomalum (Florida Thoroughwort) · E. aromaticum 'Joicus Variegated' (Boneset) · E. cannabinum (Ague Weed) · E. cannabinum 'Flore Pleno' (Hemp Agrimony) · E. cannabinum 'Plenum' (Hemp Agrimony) · E. capillifolium (Dog Fennel) · E. chapmanii (Chapman's Thorough-Wort) · E. chinense (Boneset) · E. coelestinum 'Cori' (Cori Hardy Ageratum) · E. coelestinum 'Wayside' (Blue Mistflower) · E. compositifolium (Dogfennel Eupatorium) · E. dubium (Coastal Plain Joe-Pye Weed) · E. dubium 'Little Joe' (Coastal Plain Joe Pye Weed) · E. fistulosum (Hollow Joe-Pye Weed) · E. fistulosum 'Atropurpureum' (Joe Pye Weed) · E. fortunei (Pei Lan) · E. glaucescens (Small-Flower Thorough-Wort) · E. glechonophyllum (Eupatorium) · E. godfreyanum (Godfrey's Boneset) · E. grandiflorum (Tasselflower Brickellbush) · E. hyssopifolium (Hyssop-Leaf Thoroughwort) · E. hyssopifolium var. angustissimum (Hyssop-Leaf Thoroughwort) · E. hyssopifolium var. calcaratum (Hyssopleaf Thoroughwort) · E. hyssopifolium var. hyssopifolium (Hyssopleaf Thoroughwort) · E. hyssopifolium var. laciniatum (Hyssopleaf Thoroughwort) · E. hyssopifolium var. laciniatum Gray (Hyssopleaf Thoroughwort) · E. laciniatum (Hyssopleaf Thoroughwort) · E. lancifolium (Lance-Leaf Thorough-Wort) · E. leptophyllum (False Fennel) · E. leucolepis (Justiceweed) · E. leucolepis (DC.) Torr. and Gray var. leucolepis (DC.) Torr. and Gray (Justiceweed) · E. leucolepis var. leucolepis (Justiceweed) · E. leucolepis var. novae-angliae (Justiceweed) · E. maculatum (Spotted Joe Pye Weed) · E. maculatum Atropurpureum Group (Joe-Pye Weed Atropurpureum Group) · E. maculatum bruneri var. bruneri (Spotted Joe-Pyeweed) · E. maculatum var. bruneri (Spotted Joe-Pye-Weed) · E. maculatum var. bruneri (Gray) Breitung (Spotted Joe-Pye-Weed) · E. maculatum var. foliosum (Spotted Joepyeweed) · E. maculatum var. foliosum (Fern.) Wieg. (Spotted Joepyeweed) · E. maculatum var. maculatum (Spotted Joepyeweed) · E. maculatum 'Bartered Bride' (Boneset) · E. maculatum 'Carin' (Spotted Joe Pye Weed) · E. maculatum'Little Joe' (Little Joe Dwarf Spotted Joe Pie Weed) · E. maculatum 'Purple Bush' (Spotted Joe Pye Weed) · E. mikanioides (Semaphore Thorough-Wort) · E. mohrii (Mohr's Thorough-Wort) · E. perfoliatum (Agueweed) · E. perfoliatum f. trifolium (Thoroughwort) · E. perfoliatum L. var. colpophilum Fern. & Grisc. (Common Boneset) · E. perfoliatum var. colpophilum (Common Boneset) · E. perfoliatum var. perfoliatum (Common Boneset) · E. pilosum (Rough Boneset) · E. pinnatifidum (Eupatorium) · E. purpureum (Sweet Joe Pye Weed) · E. purpureum maculatum (Green-Stem Joe-Pye-Weed) · E. purpureum maculatum 'Gateway' (Gateway Joe Pye Weed) · E. purpureum var. holzingeri (Holzinger's Eupatorium) · E. purpureum var. holzingeri (Rydb.) E.Lamont (Holzinger's Eupatorium) · E. purpureum var. purpureum (Sweet-Scented Joe-Pye Weed) · E. purpureum var. ternifolium (Green-Stem Joe-Pye-Weed) · E. purpureum 'Joe White' (Joe Pye Weed) · E. rebaudianum (Sweet Honey Leaf) · E. resinosum (Pine Barren Thoroughwort) · E. rotundifolium (Round-Leaf Thorough-Wort) · E. rotundifolium var. cordigerum (Roundleaf Thoroughwort) · E. rotundifolium var. ovatum (Roundleaf Thoroughwort) · E. rotundifolium var. rotundifolium (Roundleaf Thoroughwort) · E. rotundifolium var. scabridum (Roundleaf Thoroughwort) · E. rugosum album (Richweed) · E. rugosum 'Chocolate' (Chocolate White Snakeroot) · E. semiserratum (Small-Flower Thorough-Wort) · E. serotinum (Late Eupatorium) · E. sessilifolium (Upland Boneset) · E. sessilifolium L. var. brittonianum Porter (Upland Boneset) · E. sessilifolium L. var. sessilifolium L. (Upland Boneset) · E. sessilifolium L. var. vaseyi (Porter) Fern. & Grisc. (An Upland Boneset) · E. sessilifolium var. brittonianum (Upland Boneset) · E. sessilifolium var. sessilifolium (Upland Boneset) · E. sessilifolium var. vaseyi (Upland Boneset) · E. sordidum (Dirty Thoroughwort) · E. steelei (Steele's Eupatorium) · E. triplinerve (Yapana) · E. truncatum (Eupatorium) · E. variabile 'Variegatum' (Joe Pye Weed) · E. x pinnatifidum (Eupatorium) · E. x truncatum (Eupatorium)
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Further Reading
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- American bee journal. Hamilton, Ill., etc., Dadant & Sons url p. 539.
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- An illustrated guide to the flowering plants of the middle Atlantic and New England states (excepting the grasses and sedges) the descriptive text written in familiar language, by George T. Stevens. .. with more than 1800 illustrations from drawings by the author. New York, Dodd, Mead and company, 1910. url p. 637.
- Annotated list of the ferns and flowering plants of New York state, by Homer D. House. Albany, The University of the state of New York, 1924. url p. 680.
- Annual report / Albany: University of the State of New York, 1902-1918. url p. 12.
- Annual report / New York State Museum. Albany, N.Y.: University of the State of New York, c1904-1920. url p. 14.
- Annual report of the New Jersey State Museum. Trenton, N.J.: MacCrellish & Quigley, url p. 735.
- Annual report of the regents of the university of the state of New York on the condition of the State Cabinet of Natural History and the historical and antiquarian collection annexed thereto. Albany, Regents of the University of the State of New York. url p. 16, p. 204.
- Bartonia;proceedings of the Philadelphia botanical club. .. 57 Supplement 1993-1996 Philadelphia, Philadelphia Botanical Club, Academy of Natural Sciences. url p. 10, p. 120, p. 123, p. 133, p. 27, p. 40, p. 40, p. 42.
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Notes
Contributors
Data Sources
Accessed through GBIF Data Portal November 24, 2007:
- Missouri Botanical Garden, Missouri Botanical Garden
- USDA PLANTS, USDA PLANTS Database
- University of Alabama Biodiversity and Systematics, Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 3875110
- Catalogue of Life Accepted Name Code: Ast-8177
- Global Biodiversity Information Facility Taxonkey: 13748668
- Globally Unique Identifier: urn:lsid:ipni.org:names:206119-1
- GRIN Nomen Number: 454513
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 35979
- Natural Heritage Network Species Identifier: PDAST3P0U0
- U.S.D.A. Plant Symbol: EUHYL
- Zipcode Zoo Species Identifier: 38204
Footnotes
- Kunsiri Chaw Siripun, Edward E. Schilling "Eupatorium". in Flora of North America Vol. 21 Page 459, 461, 462, 470, 471. Oxford University Press. Online at EFloras.org. [back]
- Mean = 154.690 meters (507.513 feet), Standard Deviation = 169.690 based on 395 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
