Overview
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Vulnerable |
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Common Names
Common Names in English:
Menzies´ Wallflower, Menzies' Wallflower, Yadon's Wallflower
Description
Family Brassicaceae
Herbs annual
, biennial, or perennial
, sometimes subshrubs
or shrubs
, with a pungent
, watery juice. Eglandular
trichomes
unicellular, simple
, stalked or sessile, 2- to many forked
, stellate
, dendritic
, or malpighiaceous (medifixed
, bifid, appressed
), rarely peltate and scalelike; glandular
trichomes multicellular, with uniseriate
or multiseriate stalk
. Stems erect
, ascending
, or prostrate
, sometimes absent. Leaves exstipulate
, simple, entire
or variously pinnately dissected
, rarely trifoliolate
or pinnately, palmately, or bipinnately compound
; basal leaf rosette present or absent; cauline leaves almost always alternate, rarely opposite or whorled
, petiolate
or sessile, sometimes absent. Inflorescence bracteate
or ebracteate
racemes
, corymbs, or panicles, sometimes flowers solitary on long pedicels originating from axils of rosette leaves
. Flowers hypogynous, mostly actinomorphic
. Sepals 4, in 2 decussate pairs, free
or rarely united
, not saccate
or lateral
(inner) pair saccate. Petals 4, alternate with sepals, arranged in the form of a cross
(cruciform
; hence the earlier family name
Cruciferae), rarely rudimentary
or absent. Stamens 6, in 2 whorls, tetradynamous
(lateral (outer) pair shorter than median
(inner) 2 pairs), rarely equal or in 3 pairs of unequal length, sometimes stamens 2 or 4, very rarely 8-24; filaments
slender, winged
, or appendaged, median pairs free or rarely united; anthers
dithecal
, dehiscing by longitudinal
slits. Pollen grains
3-colpate, trinucleate
. Nectar glands receptacular
, highly diversified in number, shape
, size, and disposition around base
of filaments, always present opposite bases of lateral filaments, median glands present or absent. Pistil 2-carpelled; ovary superior, sessile or borne on a distinct
gynophore
, mostly 2-locular and with a false septum
connecting 2 placentae; placentation parietal
, rarely apical; ovules anatropous
or campylotropous, bitegmic, 1 to many per locule. Fruit typically a 2-valved capsule, generally termed silique (siliqua) when length 3 × or more than width
, or silicle
(silicula) when length less than 3 × width, dehiscent
or indehiscent, sometimes schizocarpic, nutletlike, lomentaceous
, or samaroid
, segmented
or not, terete
, angled
, or flattened parallel to septum (latiseptate
) or at a right
angle
to septum (angustiseptate) ; valves
2(or 3-6) ; replum (persistent
placenta) rounded
, rarely flattened or winged; septum complete, perforated, reduced to a rim
, or lacking; style
1, distinct, obsolete
, or absent; stigma capitate or conical
, entire or 2-lobed, sometimes lobes
decurrent and free or connate
. Seeds without endosperm, uniseriately or biseriately arranged in each locule, aseriate when 1, winged or wingless, mucilaginous
or not when wetted; cotyledons incumbent
(embryo notorrhizal: radicle lying along back of 1 cotyledon), accumbent
(embryo pleurorrhizal: radicle applied to margins
of both cotyledons), or conduplicate
(embryo orthoplocal: cotyledons folded longitudinally around radicle), rarely spirally coiled (embryo spirolobal) . Germination epigeal.
About 330 genera and 3500 species: all continents except Antarctica, mainly in temperate
areas, with highest diversity
in Irano-Turanian, Mediterranean, and W North American regions; 102 genera (eight endemic) and 412 species (115 endemic) in China.
Because of lack of a comprehensive classification based on phylogenetic
relationships
among genera, and in order
to facilitate direct comparison between the accounts in FOC and FRPS, the sequence of genera follows Schulz (Nat. Pflanzenfam. 17b: 227-658. 1936) . However, it should be emphasized that his system is largely artificial because he placed closely related genera in different tribes
and remotely related genera in the same tribe. For example, Arabis, Aubrieta, Draba, and Macropodium, which are very closely related on the basis of recent molecular studies, were placed by Schulz in four different tribes, whereas the unrelated Capsella, Lepidium, and Thlaspi were placed in one tribe. The delimitation
of genera is often difficult, and mature
fruits are needed for reliable identification.
The Brassicaceae include many important crop
plants
that are grown as vegetables (Brassica, Nasturtium, Raphanus) and sources of vegetable oils
(Brassica) and condiments (Armoracia, Brassica, Eutrema, Sinapis) . Oils of Brassica probably rank first in terms
of tonnage of the world's production
of edible oils. The family
includes many ornamentals
in the genera Erysimum, Iberis Linnaeus, Lobularia, Malcolmia, and Matthiola. Of these, only Lobularia maritima has become naturalized
in China. The family also includes more than 120 species of weeds
. Arabidopsis thaliana, which is naturalized in China, has become the model
organism
in many fields
of experimental biology
.
The delimitation of genera in the Brassicaceae is often difficult because of the frequent independent
evolution of what appear to be similar character states
, the variability of a given character in one genus and its fixture in another, and the inadequate sampling
of material
by most authors
. Fruit characters are essential in the identification of genera. However, a key
emphasizing flowering material is given in addition to the one emphasizing fruit. The most reliable determination of genera can be achieved when the material has both fruit and flowers and when both keys are successfully used to reach the same genus.
The types of cotyledonary position, which in reality is the position of the radicle in relation to the cotyledons (e.g.
, radicle accumbent, incumbent, or conduplicate, though radicle referred to as "cotyledon" throughout) are important in the separation
of many genera, and a cross section of the seed provides the easiest and simplest way of determining that position. The exact number of ovules per ovary can easily be determined from the fruit because aborted ovules persist through fruit dehiscence. However, it is more laborious to determine the ovule number from pistils and young fruit.Tai-yien Cheo, Lianli Lu, Guang Yang, Ihsan Al-Shehbaz & Vladimir Dorofeev "Brassicaceae". in Flora of China Vol. 8 Page 1. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
Genus Erysimum
Herbs annual
, biennial, or perennial
, rarely subshrubs
or shrubs
. Trichomes
sessile, medifixed
, appressed
, malpighiaceous or 3-5(-8) -rayed stellate
. Stems simple
or branched basally and/or apically. Basal leaves
petiolate
, rosulate, simple, entire
or dentate
, rarely pinnatifid
or pinnatisect
. Cauline leaves petiolate or sessile, cuneate or attenuate at base
, rarely auriculate
, entire or dentate. Racemes
ebracteate
or basally bracteate
, rarely bracteate throughout, corymbose
, elongated or not in fruit. Fruiting pedicels slender or thickened and nearly as wide as fruit, erect
, ascending
, divaricate
, or reflexed
. Sepals oblong
or linear
, erect, pubescent
, base of lateral
pair saccate
or not. Petals yellow or orange, rarely white, pink, purple, or violet; blade
suborbicular
, obovate
, spatulate
, or oblong, apex rounded
or emarginate
; claw
differentiated from blade, subequaling or longer
than sepals. Stamens 6, erect, tetradynamous
; anthers
oblong or linear. Nectar glands 1, 2, or 4, distinct
or confluent
and subtending
bases of all stamens; median
glands
present or absent. Ovules 15-100 per ovary. Fruit dehiscent siliques or rarely silicles
, linear or rarely oblong, terete
, 4-angled, latiseptate
, or angustiseptate, sessile or rarely shortly stipitate
; valves
with an obscure
to prominent
midvein
, pubescent on outside, rarely also on inside, keeled
or not, smooth
or torulose
; replum rounded; septum
complete
, membranous, translucent
or opaque
, veinless; style
obsolete
or short, rarely half as long as or subequaling fruit, often pubescent; stigma capitate, entire or 2-lobed. Seeds uniseriate
or rarely biseriate
, winged
, margined
, or wingless, oblong, plump or flattened; seed coat
minutely reticulate
, mucilaginous
when wetted; cotyledons incumbent
or rarely accumbent
.
About 150 species: N hemisphere, primarily in Asia and Europe, with 14 species in Central and North America and eight species in N Africa and Macaronesia; 17 species (five endemic) in China.Zhou Taiyan (Cheo Tai-yien, Lu Lianli (Lou Lian-li, Yang Guang, Vladimir I. Dorofeyev, Ihsan A. Al-Shehbaz "Erysimum". in Flora of China Vol. 8 Page 163. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
Physical Description
Habit: Deciduous.
Flowers: Bloom Period: January, February, March. • Flower Color: gold, yellow-orange
Size/Age/Growth
Size: 6-12" tall.
Biology
Reproduction
Duration: Biennial, Perennial
Growth
Culture: Space 12-15" apart.
Soil: Minimum pH: 5.6 • Maximum pH: 7.5
Sunlight: Sun Exposure: Full Sun .
Moisture: Drought Tolerance: High
Temperature: Cold Hardiness: 9a, 9b, 10a, 10b, 11. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Dilleniidae
(
)
- Takhtajan, 1967
- Superorder:
Violanae
(
)
- R. Dahlgren Ex Reveal, 1992
- Order:
Capparales
(
)
- J. Hutchinson, 1924
- Suborder:
Capparineae
(
)
-
- Family:
Brassicaceae
(
)
- Burnett, 1835, Nom. Cons.
- Mustard Family
- Tribe:
Hesperideae
(
)
- Genus:
Erysimum
(
)
- Linnaeus, Sp. Pl. 2: 660. 1753.
- Wallflower
- Specific epithet:
menziesii
- Price
- Subspecies:
yadonii
- Botanical name: - Erysimum menziesii yadonii Price
- Subspecies:
yadonii
- Specific epithet:
menziesii
- Price
- Genus:
Erysimum
(
- Tribe:
Hesperideae
(
- Family:
Brassicaceae
(
- Suborder:
Capparineae
(
- Order:
Capparales
(
- Superorder:
Violanae
(
- Subclass:
Dilleniidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Erysimum menziesii (Hook.) Wettst. subsp. yadonii R.A. Price
- Erysimum menziesii ssp. yadonii Price
- Erysimum menziesii yadonii Price
Notes
An accepted name
in the RHS Horticultural Database.
Place of publication
: Oesterr. Bot. Z. 39:283. 1881
Name
verified on 08-Apr-1994 by ARS Systematic Botanists. Last updated: 26-Jan-1995
Similar Species
Members of the genus Erysimum
There are approximately 930 species in this genus. Here are just 100 of them:
E. 'Anne Marie' · E. 'Apricot Delight' · E. 'Apricot Twist' (Perennial Wallflower) · E. 'Aunt May' · E. 'Baby Purple' · E. 'Bowles' Mauve' (Bowles' Mauve Perennial Wallflower) · E. 'Bowles' Purple' · E. 'Bowles' Yellow' · E. 'Bredon' · E. 'Breedon' · E. 'Butterscotch Variegated' · E. 'Butterscotch' · E. 'Changeling' · E. 'Charity Rose Red' · E. 'Charity Scarlet' · E. 'Cheerfulness' · E. 'Chelsea Jacket' · E. 'Chequers' · E. 'Clent Calcutt' · E. 'Constant Cheer' · E. 'Cotswold Gem' (Perennial Wallflower) · E. 'Cream Delight' · E. 'Dawn Breaker' · E. 'Devon Cream' · E. 'Devon Gold' · E. 'Devon Sunset' · E. 'Dorothy Elmhirst' · E. 'Ellen Willmott' · E. 'Emm's Variety' · E. 'Eryzwer' · E. 'Fragrant Sunshine' (Perennial Wallflower) · E. 'Gingernut' · E. 'Glowing Embers' · E. 'Gold Flame' · E. 'Gold Shot' · E. 'Golden Gem' · E. 'Golden Jubilee' · E. 'Hareknoll' · E. 'Hector Harrison' · E. 'Hector's Gate Post' · E. 'Hector's Gatepost' · E. 'Jacob's Jacket' · E. 'Jaunty Joyce' · E. 'Jenny Brook' · E. 'Joan Adams' · E. 'John Codrington' (Perennial Wallflower) · E. 'Joseph's Coat' · E. 'Jubilee Gold' · E. 'Julian Orchard' · E. 'Lady Roborough' · E. 'Lewis Hart' · E. 'Little Kiss' · E. 'Mayflower' · E. 'Mill Cottage Dawn' · E. 'Mill Cottage Dusk' · E. 'Miss Hopton' · E. 'Miss Massey' · E. 'Moonlight' · E. 'Mrs L.K. Elmhirst' · E. 'Multicolor' · E. 'New Zealand Limelight' · E. 'Newark Park' · E. 'Onslow Seedling' · E. 'Orange Flame' · E. 'Orange Queen' · E. 'Orange' · E. 'Parish's' · E. 'Pastel Patchwork' (Perennial Wallflower) · E. 'Perry's Peculiar' · E. 'Perry's Pumpkin' · E. 'Perry's Variegated' · E. 'Plant World Antique' · E. 'Plant World Gold' (Perennial Wallflower) · E. 'Plant World Lemon' (Perennial Wallflower) · E. 'Primrose' · E. 'Purple' · E. 'Rosemoor' · E. 'Rufus' · E. 'Rushfield Sunrise' · E. 'Rushfield Sunset' · E. 'Rushfield Surprise' · E. 'Ruston Royal' · E. 'Scorpio' · E. 'Sissinghurst Variegated' · E. 'Sprite' · E. 'Starbright' · E. 'Stars and Stripes' · E. 'Sunbright' · E. 'Sunshine' · E. 'Sweet Sorbet' · E. 'Tricolor' · E. 'Turkish Bazaar' · E. 'Valerie Finnis' · E. 'Variegated Butterscotch' · E. 'Walfrasun' · E. 'Wembdon Bravery' · E. 'Wenlock Beauty Variegated' · E. 'Wenlock Beauty' · E. 'Winter Joy' · E. 'Yellow Bird'
Bibliography
- Hickman, J. C., ed. 1993. The Jepson manual: Higher plants of California. (F CalifJep)
- Kartesz, J. T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. (L US Can ed2)
- United States Fish and Wildlife Service. U.S. Fish & Wildlife Service. Species Information: threatened and endangered animals and plants - on-line resource. (US Fish & Wildl List)
- Cheo Tai-yien, Guo Rong-lin, Lan Young-zhen, Lou Lian-li, Kuan Ke-chien & An Zheng-xi. 1987. Cruciferae. In: Cheo Tai-yien, ed., Fl. Reipubl. Popularis Sin. 33: 1-483.
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Notes
Contributors
- Bisby FA, Roskov YR, Orrell TM, Nicolson D, Paglinawan LE, Bailly N, Kirk PM, Bourgoin T, van Hertum J, eds (2008). Species 2000 & ITIS Catalogue of Life: 2008 Annual Checklist Taxonomic Classification. CD-ROM; Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-2006. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed October 5, 2006.
- MBLWHOI Library: Universal Biological Index and Organizer. uBio.org accessed July 17, 2008.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 27, 2008)
- USDA, NRCS. 2005. The PLANTS Database, Version 3.5 (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA.
Identifiers
- Biodiversity Heritage Library NamebankID: 8289324
- Catalogue of Life Accepted Name Code: ITS-524934
- Global Biodiversity Information Facility Taxonkey: 13902120
- GRIN Nomen Number: 319879
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 524934
- Natural Heritage Network Species Identifier: PDBRA160E4
- U.S.D.A. Plant Symbol: ERMEY
- Zipcode Zoo Species Identifier: 38188
