Overview
Interesting Facts
Common Names
Common Names in English:
Common Woolly Sunflower, Common Woolly-Sunflower, Oregon Sunshine, Woody Eriophyllum, Woolly Daisy, Woolly Eriophyllum, Woolly Sunflower, Wooly Daisy
Description
Family Compositae
The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.
Genus Eriophyllum
Annuals
, perennials
, subshrubs
, or shrubs
, 1-200 cm. Stems erect
or decumbent
, usually branched (proximally, distally, or ± throughout). Leaves mostly cauline; mostly alternate (proximal
sometimes opposite) ; petiolate
or sessile; blades
usually 1-2(-3) -pinnately lobed
, ultimate
margins
toothed
, serrate, or entire, faces
usually densely to sparsely woolly
(abaxial
or both, adaxial
sometimes glabrescent
). Heads radiate
or discoid
, borne singly or in loose
to tight, corymbiform
or compound-corymbiform arrays. Involucres campanulate
to hemispheric
, 3-12+ mm diam. Phyllaries persistent
, 4-13(-15) in 1+ series (± erect in fruit, distinct
or basally connate
, lanceolate to oblanceolate
, herbaceous or indurate
, slightly to deeply concave
, usually carinate
, margins sometimes scarious
, abaxial faces densely to sparsely woolly). Receptacles flat or convex
to conic, smooth
or pitted
, glabrous
, usually epaleate (with 1-6 hyaline
paleae in E. ambiguum, obscurely setose
in E. mohavense). Ray florets 0, or 4-13(-15), pistillate
, fertile
; corollas yellow or white (sometimes with reddish veins in E. lanosum). Disc florets (3-) 10-300, bisexual
, fertile; corollas yellow, tubes
shorter than or about equaling funnelform
throats
, lobes
5, deltate. Cypselae linear-clavate to prismatic
, flattened or 3-angled in rays, 4(-5) -angled in discs, hairy
or glabrous; pappi 0, or persistent, of 6-12+ (distinct) erose to laciniate
or aristate
scales
(in 1-2 similar or contrasting series), or ± coroniform
. x = 8.
Species 13: w North America, nw Mexico.
Eriophyllum encompasses taxa that occur in seashore, chaparral
, grassland, desert, forest
, and alpine
communities. Their disparateness encouraged taxonomic
multiplication. Between 1890 and 1937, about 157 designations
under the genus existed (L. Constance 1937). Constance reduced that taxonomic thicket to six annual and five perennial species; the perennial species E. lanatum consisted of ten varieties. Base
diploid chromosome numbers for Eriophyllum (in the sense of Constance) species are x
= 4, 5, and 7 for the annuals, and x = 8, 15, and 19 for the perennials (S. Carlquist 1956; J. S. Mooring 1997, 2001, 2002). Possibly, x = 15 and x = 19 represent paleopolyploidy. Only E. mohavense remains uncounted. B
. G. Baldwin (1999) linked Mooring's (1997) report of n = 19 in E. nevinii to chromosomal, morphologic, and rDNA evidence, and erected the genus Constancea on that species. Eriophyllum (in the sense of Constance) seems most closely related to the annuals Pseudobahia (x = 3, 4, 8) and Syntrichopappus (x = 6, 7). Eriophyllum (in the sense of Baldwin), Pseudobahia, and Syntrichopappus constitute a clade, and nomenclatural
changes are necessary for a monophyletic classification (Baldwin and B. L. Wessa 2000; Baldwin et al.
2002). Mooring (1997) hypothesized a descending
dysploidy
phylogeny in Eriophyllum (in the sense of Constance) from E. nevinii. Baldwin et al. (2002, p. 174) stated that E. nevinii "is an evolutionary outlier (although probably not ancestral) to" Eriophyllum in the sense of Constance.Natural intertaxon
hybrids have been reported for the perennial species (L. Constance 1937; J. S. Mooring 1994) but not for the annuals. Experimental hybridizations have produced
sterile
hybrids between the annual E. congdonii and the perennial E. lanatum. Experimental crosses among seven of the annual species
produced fertile hybrids between two morphologically similar species; the other combinations
either failed or produced sterile hybrids (Mooring 2002).D. P. Tibor (2001) cited nine taxa of Eriophyllum as rare or endangered
: the perennials E. confertiflorum var. tanacetiflorum, E. jepsonii, E. latilobum, E. (Constancea) nevinii, and E. lanatum vars. hallii and obovatum, and the annuals E. congdonii, E. mohavense, and E. nubigenum.[1]
Physical Description
Species Eriophyllum lanatum
Perennials
or subshrubs
, 10-100 cm (sometimes flowering first
year). Stems erect
to decumbent
(usually woolly
). Leaves
(proximal
usually alternate) : blades
mostly lanceolate to oblanceolate
,
1-8 cm, often 1-2(-3) -pinnately lobed
, ultimate
margins
toothed
,
serrate, or entire, revolute
or plane
, faces
hairy
, often woolly
(more densely abaxially, sometimes glabrate
adaxially; distal leaves
reduced in size and lobing). Heads borne singly or (2-5+)
in corymbiform
arrays. Peduncles mostly 3-30 cm. Involucres
campanulate
to hemispheric
, 6-15 mm diam. Phyllaries 5-13(-15),
distinct
or connate
at bases
(lanceolate to ovate
, carinate
or plane).
Ray florets 0 or 5-13(-15) ; laminae
golden yellow to yellow,
6-20 (× 2-7) mm.
Disc florets 20-300; corollas 2.5-5
mm (tubes
usually glandular
or glandular-hairy, glabrous
in var.
hallii). Cypselae 2-5 mm; pappi usually of 6-12
ovate or cuneate to lanceolate or lance-linear
(often unequal), erose
or lacerate
scales
0.3-2 mm, sometimes coroniform
, rarely 0. [source]
Eriophyllum lanatum is a polyploid complex
of intergrading
regional facies treated here as varieties. Artificial hybridization
studies show that strong
barriers
to interbreeding
exist among the
varieties at the diploid level (J. S. Mooring 2001). In nature, morphologically
intermediate polyploid populations often occur in regions where the
ranges
of the varieties approach one another. Edaphic factors
and
light intensity
also make identification more difficult by strongly
influencing leaf morphology and sizes of structures. For example,
cultivated individuals of var. achilleoides may have laciniately
toothed rather than pinnatifid
leaves. Rarely, plants
of different
varieties maintain their identity while growing side by side. In
some instances, one is diploid and the other tetraploid
; in others
both are diploid. Varieties arachnoideum, croceum, grandiflorum,
and obovatum apparently form natural hybrids with E. confertiflorum
var. confertiflorum; past hybridizations may have resulted
in the origin
of E. latilobum and E. jepsonii (L. Constance
1937; P. A. Munz 1959; Mooring 1994) and E. confertiflorum
var. tanacetiflorum (Mooring 1994). [source]
Our treatment of Eriophyllum lanatum closely follows that
of L. Constance (1937), which was done without benefit of cytogeographic
studies. The key
is to modal populations of the varieties, usually
based on living plants. [source]
Some varieties have been introduced
into cultivation as ornamentals
.
[source]
Habit: Subshrub , Shrub , Forb/herb
Flowers: Bloom Period: January, February, March, April, May. • Flower Color: yellow
Size/Age/Growth
Size: 6-12" tall.
Habitat
Typically found at an altitude of 0 to 2,770 meters (0 to 9,088 feet).[2]
Biology
Reproduction
Duration: Annual , Perennial
Growth
Culture: Space 12-15" apart.
Soil: Minimum pH: 6.1 • Maximum pH: 7.8
Sunlight: Sun Exposure: Full Sun .
Temperature: Cold Hardiness: 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b, 9a, 9b, 10a, 10b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Campanulanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Asterales
(
)
- Lindley, 1833
- Family:
Compositae
(
)
- Giseke, 1792, nom. cons., nom. alt.
- Subfamily:
Asteroideae
(
)
- Tribe:
Helenieae
(
)
- Subtribe:
Madiinae
(
)
- Genus:
Eriophyllum
(
)
- Lagasca, 1816
- Woolly sunflower [Greek erion, wool, and phyllon, leaf]
- Specific epithet:
lanatum
- (Pursh) J.Forbes
- Botanical name: - Eriophyllum lanatum (Pursh) J.Forbes
- Specific epithet:
lanatum
- (Pursh) J.Forbes
- Genus:
Eriophyllum
(
- Subtribe:
Madiinae
(
- Tribe:
Helenieae
(
- Subfamily:
Asteroideae
(
- Family:
Compositae
(
- Order:
Asterales
(
- Superorder:
Campanulanae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Actinella lanata Pursh
Notes
Name
Status: Accepted Name
.
Comment: Data Providers: CONABIO, Govaerts World Compositae Checklist
A-G, IPNI, Tropicos, LCR Editor. GCC LSID: urn
:lsid:compositae.org:names:1C730207-83B9-4A38-9533-4619FC9BD884
Last scrutiny: 13-Aug-09
Similar Species
Members of the genus Eriophyllum
ZipcodeZoo has pages for 32 species, subspecies, varieties, forms, and cultivars in this genus:
E. ambiguum (Beautiful Woolly Sunflower) · E. ambiguum var. ambiguum (Beautiful Woolly Sunflower) · E. ambiguum var. paleaceum (Beautiful Woolly Sunflower) · E. confertiflorum (Golden-Yarrow) · E. confertiflorum (DC.) Gray var. confertiflorum (DC.) Gray (Golden-Yarrow) · E. confertiflorum var. confertiflorum (Golden-Yarrow) · E. confertiflorum var. tanacetiflorum (Golden-Yarrow) · E. congdonii (Congdon's Eriophyllum) · E. jepsonii (Jepson Woolly-Sunflower) · E. lanatum (Common Woolly Sunflower) · E. lanatum (Pursh) Forb. var. integrifolium (Hook.) Smiley (Common Woolly Sunflower) · E. lanatum (Pursh) Forb. var. leucophyllum (DC.) W. R. Carter (Common Woolly Sunflower) · E. lanatum var. achillaeoides (Common Woolly Sunflower) · E. lanatum var. aphanactis (Common Woolly Sunflower) · E. lanatum var. arachnoideum (Common Woolly Sunflower) · E. lanatum var. croceum (Common Woolly Sunflower) · E. lanatum var. cuneatum (Common Woolly Sunflower) · E. lanatum var. grandiflorum (Common Woolly Sunflower) · E. lanatum var. hallii (Hall's Woolly Sunflower) · E. lanatum var. integrifolium (Common Woolly Sunflower) · E. lanatum var. lanatum (Common Woolly Sunflower) · E. lanatum var. lanceolatum (Common Woolly Sunflower) · E. lanatum var. leucophyllum (Common Woolly Sunflower) · E. lanatum var. obovatum (Common Woolly Sunflower) · E. lanatum 'Pointe' (Oregon Sunshine) · E. latilobum (San Mateo Woolly Sunflower) · E. mohavense (Barstow Wooly-Sunflower) · E. multicaule (Many-Stem Woolly-Sunflower) · E. nevinii (Nevin's Woolly Sunflower) · E. nubigenum (Yosemite Woolly Sunflower) · E. pringlei (Pringle's Eriophyllum) · E. stoechadifolium (Seaside Woolly Sunflower)
More Info
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Further Reading
- 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 170.
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- A manual of the flora of northern Idaho /Carl Epling and Joe Ewan. 4 1941 1941. url p. 828.
- A report upon the boreal flora of the Sierra Nevada of California / by Frank Jason Smiley. Berkeley, Calif.: University of California Press, 1921. url p. 377, p. 378.
- A study of the vegetation of southeastern Washington and adjacent Idaho. .. Lincoln, Neb.[1917] url p. 61.
- An annotated list of the Diptera (flies) of Oregon, San Francisco, 1921 url .
- Bartonia;proceedings of the Philadelphia botanical club. .. 62 2004 Philadelphia, Philadelphia Botanical Club, Academy of Natural Sciences. url p. 75.
- Birds of Oregon, by Ira N. Gabrielson. .. and Stanley G. Jewett. .. Corvallis, Or., Pub. by Oregon State College through the O.S.C. Cooperative Association[c1940] url p. 36.
- Brigham Young University science bulletin. 11 1970 Provo, Utah: Brigham Young University, [1955-1976] url p. 21.
- Britton, N. L. (ed.). North American flora. 7 1922 [New York]New York Botanical Garden. url p. 603, p. 894, p. 92.
- Bulletin of the Torrey Botanical Club. 1914 41 New York: Torrey Botanical Club, 1870-1996 url p. 341, p. 342, p. 646.
- California fish and game. [San Francisco, etc.]: State of California, Resources Agency, Dept. of Fish and Game. url p. 172, p. 264.
- Carnegie Institution of Washington publication. Washington, Carnegie Institution of Washington, 1902- url p. 152, p. 193.
- Catalog of hymenoptera in America north of Mexico / prepared cooperatively by specialists on the various groups of Hymenoptera under the direction of Karl V. Krombein. .. [et al.]. Washington: Smithsonian Institution Press, 1979- url , p. 1938, p. 2008, p. 2009, p. 2035.
- Compositae newsletter. Columbus, Ohio: Dept. of Botany, Ohio State University, 1975- url p. 32, p. 33, p. 46.
- Compositae of southern California / by Harvey Monroe Hall. Berkeley, [Calif.]: The University Press, 1907. url p. 180.
- Contributions from the United States National Herbarium 48 2003 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 18, p. 583.
- Contributions from the University of Michigan Herbarium. 11 1973 Ann Arbor: University Herbarium, University of Michigan, 1939- url p. 137.
- Dedication papers: scientific papers presented at the dedication of the laboratory building and plant houses, April 19-21, 1917. Brooklyn, N.Y.: Brooklyn Botanic Garden, 1918. url p. 246, p. 293.
- Elementary flora of the Northwest, Cincinnati [etc.]American Book Company[c1914] url .
- Field book of western wild flowers, by Margaret Armstrong in collaboration with J.J. Thornber. .. with five hundred illustrations in black and white, and forty-eight plates in color drawn from nature by the author. 1915 London, C. [sic] P. Putnam's Sons, 1915. url p. 586, p. 586.
- Flora of southeastern Washington and adjacent Idaho / by Charles V. Piper and R. Kent Beattie. Lancaster, Pa.: New Era, 1914. url p. 278, p. 279.
- Flora of southern British Columbia and Vancouver Island: with many references to Alaska and northern species / by Joseph Kaye Henry. Toronto: W.J. Gage, c1915. url p. 306.
- Flora of the Rocky Mountains and adjacent plains, Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and neighboring parts of Nebraska, South Dakota, North Dakota, and British Columbia / by P.A. Rydberg. New York: The author, 1922. url p. 947.
- Flora of the northwest coast, including the area west of the summit of the Cascade Mountains, from the forty-ninth parallel south to the Calapooia Mountains on the south border of Lane County, Oregon. Lancaster, Pa., Press of the New Era Printing Company, 1915. url .
- Flora of the northwest coast: including the area west of the summit of the Cascade Mountains, from the forty-ninth parallel south to the Calapooia Mountains on the south border of Lane County, Oregon / by Charles V. Piper and R. Kent Beattie. Lancaster, Pa.: Press of the New era printing company, 1915. url p. 378.
- Flora of the southeastern Washington and adjacent Idaho, Lancaster, Pa., Press of the New Era Printing Company, 1914. url , .
- Flora of the state of Washington / by Charles V. Piper. Washington: G.P.O., 1906 url p. 583.
- Flora of the state of Washington. By Charles V. Piper. Washington, Govt. Print. Off., 1906. url p. 583.
- Great Basin naturalist memoirs. 1987 [Provo, Utah]Brigham Young University, 1976-1992. url p. 192, p. 864.
- Journal of ethnobiology. 8-9 1988-1989 Flagstaff, Ariz.: Center for Western Studies, 1981- url p. 110.
- Key to the Rocky Mountain flora. Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and parts of Nebraska, South Dakota, North Dakota, and British Columbia. New York, [c1919] url .
- Key to the Rocky Mountain flora; Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and parts of Nebraska, South Dakota, North Dakota, and British Columbia. New York, 1919. url p. 257.
- Leaflets of western botany. San Fransisco:[J. T. Howell], 1932-1966. url , , p. 121, p. 124, p. 126, p. 203, p. 271, p. 272, p. 42.
- Memoirs / Brooklyn Botanic Garden. Brooklyn, N.Y.: The Garden, 1918-1936. url p. 246, p. 293.
- Memoirs of the California Academy of Sciences. San Francisco: The Academy, 1868- url p. 131.
- Muhlenbergia. Chico, Calif. [etc] url p. 131, p. 176.
- Novon a journal of botanical nomenclature from the Missouri Botanical Garden. 1 1991 St. Louis, MO: Missouri Botanical Garden, url p. 119, p. 121.
- Plant indicators: the relation of plant communities to process and practice. WashingtonCarnegie Institution of Washington1920 url p. 152, p. 193.
- Proceedings of the California Academy of Sciences, 4th series. San Francisco, California Academy of Sciences. url p. 326.
- Proceedings of the Entomological Society of Washington. Washington, etc.: Entomological Society of Washington url p. 283, p. 554, p. 612, p. 620, p. 695.
- Report on the conservation status of Penstemon lemhiensis, a candidate threatened species: Montana / by J. Stephen Shelly. Helena, Mont.: Montana Natural Heritage Program, [1990] url p. 88.
- Sensitive plant surveys in the Gallatin National Forest, Montana / prepared by Jim Vanderhorst; prepared for U.S. Forest Service, Gallatin National Forest. Helena, MT: Montana Natural Heritage Program, [1994]. url p. 71.
- Status review update and establishment of demographic monitoring studies: Penstemon lemhiensis / prepared for U.S.D.A. Forest Service, Region 1, Beaverhead and Bitterroot National Forest, Montana; prepared by J. Stephen Shelly. Helena, Mont.: Montana Natural Heritage Program, [1990] url p. 75.
- Syllogeus. Ottawa, National Museum of Natural Sciences, 1972-1995. url p. 115.
- The Canadian field-naturalist. Ottawa, Ottawa Field-Naturalists' Club. url p. 241, p. 242, p. 604.
- The Great Basin naturalist. 43 1983 Provo, Utah: M.L. Bean Life Science Museum, Brigham Young University, 1939-1999. url p. 103, p. 233, p. 281, p. 32.
- The Mountaineer. Seattle, The Mountaineers, 1907-1921. url p. 115.
- The Plant world. Baltimore [etc.]Plant World Association [etc.] url p. 258.
- The University of Missouri studies. Columbia, Mo.: The University, 1905-[1918] url p. 283, p. 393, p. 431.
- The flora of Boulder, Colorado, and vicinity, by Francis Potter Daniels. .. [Columbia, Mo.]The University of Missouri, 1911. url p. 245, p. 283.
- Torreya. Burlington, Vt., Torrey Botanical Club, 1901-1945. url p. 225, p. 251.
- University of California publications in botany. Berkeley, Calif., University of California Press, 1902-2001. url p. 378, p. 408.
- University studies of the University of Nebraska. Lincoln. url p. 61.
- Constance, L. 1937. A systematic study of the genus Eriophyllum Lag. Univ. Calif. Publ. Bot. 18: 69136.
- Mooring, J. S. 1997. A new base chromosome number and phylogeny for Eriophyllum (Asteraceae, Helenieae). Madroño 44: 364373.
- Mooring, J. S. 2002. Experimental hybridizations of Eriophyllum annuals (Asteraceae, Helenieae). Amer. J. Bot. 89: 19731983.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 11, 2012.
- "Eriophyllum lanatum". in Flora of North America Vol. 21 Page 352, 353, 354, 357, 361, 362. Published by Oxford University Press. Online at EFloras.org.
- GCC: Global Compositae Checklist. Release date: November 18, 2009
- Global Biodiversity Information Facility. Accessed November 25, 2007. http://www.gbif.org Mediated distribution data from 13 providers.
- Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 27, 2008)
Data Sources
Accessed through GBIF Data Portal November 25, 2007:
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- Bundesamt für Naturschutz / Zentralstelle für Phytodiversität Deutschland, Bundesamt fuer Naturschutz / Zentralstelle fuer Phytodiversitaet Deutschland
- Missouri Botanical Garden, Missouri Botanical Garden
- Oregon State University, Vascular Plant Collection
- School of Life Sciences, Arizona State University, The Deaver Herbarium, Northern Arizona University
- USDA PLANTS, USDA PLANTS Database
- University of Washington Burke Museum, Vascular Plant Collection - University of Washington Herbarium
- Utah State University, USU-UTC Specimen Database
- Utah Valley State College
- , Utah Valley State College Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2658530
- Catalogue of Life Accepted Name Code: Ast-18506
- Global Biodiversity Information Facility Taxonkey: 13749080
- Globally Unique Identifier: urn:lsid:ipni.org:names:204863-1
- GRIN Nomen Number: 454786
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 37334
- Natural Heritage Network Species Identifier: PDAST3N050
- U.S.D.A. Plant Symbol: ERLAT ERLA
- Zipcode Zoo Species Identifier: 38349
Footnotes
- Dale E. Johnson, John S. Mooring "Eriophyllum". in Flora of North America Vol. 21 Page 255, 335, 336, 352, 353, 354, 363, 380. Oxford University Press. Online at EFloras.org. [back]
- Mean = 989.620 meters (3,246.785 feet), Standard Deviation = 618.360 based on 1,338 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
