Eriogonum corymbosum

• Interesting Facts
• Common Names
• Description
• Biology
• Taxonomy
• Similar Species
• More Info
• Further Reading
• Notes

Interesting Facts

Common Names

Description

Family Polygonaceae

Herbs, shrubs , or small trees , sometimes monoecious or dioecious. Stems erect , prostrate , twining , or scandent , often with swollen nodes, striate , grooved , or prickly. Leaves simple , alternate, rarely opposite or whorled , petiolate or subsessile ; stipules often united to a sheath (ocrea) . Inflorescence terminal or axillary , spicate , racemose, paniculate , or capitate. Pedicel occasionally articulate . Flowers small, actinomorphic , bisexual , rarely unisexual . Perianth 3-6-merous, in 1 or 2 series, herbaceous, often enlarged in fruit or inner tepals enlarged, with wings , tubercles , or spines. Stamens usually (3-) 6-9, rarely more; filaments free or united at base ; anthers 2-loculed, opening lengthwise; disk annular (often lobed ) . Ovary superior, 1-loculed; styles 2 or 3, rarely 4, free or connate at lower part. Fruit a trigonous , biconvex , or biconcave achene; seed with straight or curved embryo and copious endosperm.

About 50 genera and 1120 species: worldwide, but primarily N temperate with a few species in tropical regions ; 13 genera (two endemic) and 238 species (65 endemic) in China.[1]

Genus Eriogonum

Shrubs , subshrubs , or herbs, sometimes nearly arborescent , perennial , biennial, or annual , polycarpic or, rarely, monocarpic (subg. Pterogonum), synoecious (sometimes polygamodioecious in subg. Micrantha and Oligogonum, rarely dioecious in subg. Oligogonum) ; taproot slender to stout, solid, or rarely chambered (subg. Pterogonum). Stems prostrate or decumbent to erect , infrequently absent, glabrous or pubescent , sometimes glandular ; caudex stems absent or woody, tightly compact to spreading and at or just below surface, or spreading to erect and above surface; aerial flowering stems arising at nodes of caudex branches, at distal nodes of aerial branches, or directly from the root , prostrate or decumbent to erect, slender to stout, solid or slightly to distinctly hollow and fistulose, rarely disarticulating into ringlike segments (subg. Clastomyelon). Leaves usually persistent through anthesis , occasionally persistent through growing season or longer , sometimes marcescent or quickly deciduous, basal and sometimes sheathing up stems, cauline, or basal and cauline, alternate, opposite, or whorled , 1 per node or fasciculate; petiole usually present, sometimes obscure ; blade linear to orbiculate, entire apically. Inflorescences terminal or terminal and axillary , cymose and dichotomously or trichotomously branched, or racemose, simple or compound-umbellate, subcapitate , or capitate, occasionally distally uniparous due to suppression of secondary branches; branches mostly dichotomous except for initial trichotomous node, not brittle or disarticulating into segments, round and smooth , rarely grooved , angled or ridged , variously lanate , tomentose , floccose , sericeous , hispid , pilose-pubescent, or puberulent , occasionally glandular, rarely scabrellous; bracts 2-13 or more at proximal nodes, usually 3 distally, connate proximally, leaflike, semileaflike, or scalelike, not awn-tipped, glabrous or variously pubescent or glandular. Peduncles absent or erect to deflexed . Involucres 1-8 or more per cluster , smooth or ribbed , tubular , cylindric or narrowly turbinate to broadly campanulate or hemispheric ; teeth 5-10, sometimes lobelike, not awned . Flowers bisexual or, infrequently, unisexual , (2-) 6-100 per involucre at any single time during full anthesis, sometimes with stipelike base ; perianth usually white to red or variously yellow, broadly campanulate when open, cylindric to urceolate when closed , glabrous or pubescent or glandular abaxially; tepals 6, connate proximally to 2 their length , monomorphic or dimorphic , usually entire apically, rarely emarginate ; stamens 9; filaments adnate basally, glabrous or pubescent; anthers usually red to cream or yellow, oblong to ellipsoid or oval . Achenes included to exserted, various shades of brown, black, or occasionally yellow, rarely winged or ridged (subg. Pterogonum), lenticular or 3-gonous, glabrous or pubescent. Seeds: embryo curved or straight. x = 10.

Species ca. 250: North America (including n Mexico).

Eriogonum is the basal group of subfam. Eriogonoideae. Like all of its related genera, Eriogonum is a highly derived tetraploid taxon that has undergone rapid evolution in arid regions of western North America. The circumscription of the genera in the subfamily is now being studied molecularly and cladistically. The approach taken here is to divide the group into numerous genera, acknowledging that the resulting Eriogonum remains paraphyletic and that all genera of Eriogoneae are imbedded within Eriogonum as presently circumscribed. Resolution may well come with the reduction of the subfamily to two genera, Eriogonum and Pterostegia (including Harfordia Greene, a genus of Baja California, Mexico), or, at the other extreme, reducing Eriogonum to just two species. What the future will hold is difficult to ascertain at this time.

As presently circumscribed, Eriogonum is one of the larger genera in the flora area, being exceeded in numbers of species only by Carex (ca. 480), Astragalus (ca. 350), and Penstemon (ca. 250). As a native North American genus, Eriogonum (ca. 250) is second only to Penstemon. Ecologically, species of Eriogonum occur from the seashore to the highest mountains in the United States. They are among the last plants seen atop the Sierra Nevada and on the €œoutskirts€ of Badwater in Death Valley. About one-third of the species are uncommon to rare in their distribution. The United States Department of the Interior currently lists some as endangered or threatened species. Some species tend to be weedy, and some of the annual species are aggressively so.

Species of Eriogonum have long been regarded as among the most difficult in North America to distinguish. Regional treatments should be consulted before attempting to use this review, especially for plants found outside California or the Intermountain West. Geographic distribution is a useful character, and such information is given fully in keys and discussion here to aid with identification. In addition to regional keys noted below, keys exist for Texas (J. L. Reveal 1970b), the Pacific Northwest (J. L. Reveal 1973), and the Great Plains (R. Kaul 1986). In each instance the nomenclature should be compared with that presented here. To aid in the identification of species belonging to the largest subgenus , Eucycla, regional keys are given here, thereby avoiding a long and complex key to the more than 100 species.

In collecting specimens of Eriogonum, try to obtain leaves (especially for annuals), fruits (especially those belonging to subg. Pterogonum), and ample flowers (rarely difficult to accomplish). Field observations on flower color, pubescence , and overall size and habit are useful. Some species (especially those of subg. Oligogonum) are dioecious, with the mature staminate and pistillate plants occasionally markedly different in aspect . It is not uncommon for several annual species to grow intermixed in disturbed places, so care must be taken to prevent mixed collections . Finally, as in all cases, collectors should try to sample the range of variation rather than concentrate on extremes.

Eriogonum has a long history of aboriginal use. Today, several members of the genus are in cultivation, especially in the rock or alpine garden (G. Nicholls 2002).

Members of Eriogonum are hosts for a number of butterfly species, including such endangered ones as the El Segundo dotted-blue (Euphilotes battoides allyni), Smith's dotted blue (Euphilotes enoptes smithi), and Lange's metalmark (Apodemia mormo Iangei). Species of the genus Euphilotes spend their entire life on particular species complexes. Other butterfly species found in association with Eriogonum and relatives (see P. A. Opler and A. B . Wright 1999) include the western green hairstreak (Callophrys affinis), desert green or Comstock's hairstreak (C. comstocki), bramble hairstreak (C. dumetorum), Lembert's hairstreak (C. lemberti), Sheridan's green hairstreak (C. sheridani), green hairstreak (C. viridis), varied blue (Chalceria heteronea), Rocky Mountain dotted-blue (Euphilotes ancilla), Bauer's dotted-blue (E. baueri), Bernardino dotted-blue (E. bernardino), Ellis's dotted-blue (E. ellisi), Pacific dotted-blue (E. enoptes), intermediate dotted-blue (E. intermedia), Mojave dotted-blue (E. mojave), pallid dotted-blue (E. pallescens), Rita dotted-blue (E. rita), Spalding's dotted-blue (E. spaldingi), Gorgon copper (Gaeides gorgon), gayas or Edward's blue (Hemiargus ceranus gyas), blue copper (Lycaena heteronea), small blue (Philotiella speciosa), Boisduval's blue (Plebeius icarioides), acmon blue (P. acmon), lupine blue (P. lupini), veined blue (P. neurona), California hairstreak (Satyrium californica), nut-brown hairstreak (S. saepium), Avalon scrub-hairstreak (Strymon avalona), and gray hairstreak (S. melinus). Flowering plants of Eriogonum are infrequently visited by the sooty hairstreak (Satyrium fulginosum), the flowers being a source of nectar for adults . According to Opler, several additional species and subspecies of these butterflies remain to be described.[2]

Physical Description

Habitat

Typically found at an altitude of 0 to 2,998 meters (0 to 9,836 feet).[3]

Biology

Reproduction

Duration: Perennial

Growth

Sunlight: Sun Exposure: Full sun .

Taxonomy

Synonyms

Eriogonum effusum Nutt. var. corymbosum S.Stokes • Eriogonum effusum Nuttall corymbosum (Bentham) S. Stokes

Notes

Publishing author : Benth. Publication : Prodr. (DC.) 14(1): 17 1856 [mid Oct 1856]

Similar Species

Members of the genus Eriogonum

ZipcodeZoo has pages for 521 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:

E. abertianum (Abert Buckwheat) · E. abertianum var. abertianum (Abert's Buckwheat) · E. abertianum var. cyclosepalum (Abert's Buckwheat) · E. acaule (Pointed Eriogonum) · E. alatum (Wind Wildbuckwheat) · E. alatum var. alatum (Winged Wild Buckwheat) · E. alatum var. glabriusculum (Winged Buckwheat) · E. alatum var. mogollense (Winged Buckwheat) · E. aliquantum (Cimarron Buckwheat) · E. allenii (Shalebarren Buckwheat) · E. alpinum (Trinity Buckwheat) · E. ammophilum (Ibex Buckwheat) · E. ampullaceum (Mono Buckwheat) · E. androsaceum (Rockjasmine Buckwheat) · E. anemophilum (West Humboldt Buckwheat) · E. angulosum (Angle-Stemmed Wild-Buckwheat) · E. annuum (Annual Buckwheat) · E. apiculatum (San Jacinto Buckwheat) · E. apricum (Ione Buckwheat) · E. apricum var. apricum (Ione Buckwheat) · E. apricum var. prostratum (Irish Hill Buckwheat) · E. arborescens (Santa Cruz Island Buckwheat) · E. arcuatum var. arcuatum (Baker's Wild Buckwheat) · E. aretioides (Red Canyon Buckwheat) · E. argillosum (Clay Buckwheat) · E. argophyllum (Ruby Valley Buckwheat) · E. arizonicum (Arizona Buckwheat) · E. artificis (Kaye's Wild Buckwheat) · E. atrorubens (Buckwheat) · E. atrorubens atrorubens (Buckwheat) · E. baileyi (Bailey Buckwheat) · E. baileyi var. baileyi (Bailey's Buckwheat) · E. baileyi var. praebens (Bailey's Buckwheat) · E. batemanii (Bateman Buckwheat) · E. beatleyae (Beatley Buckwheat) · E. bicolor (Pretty Buckwheat) · E. bifurcatum (Forked Buckwheat) · E. blissianum (Bliss' Buckwheat) · E. brachyanthum (Shortflower Buckwheat) · E. brachypodum (Parry's Buckwheat) · E. brandegeei (Brandegee Wild Buckwheat) · E. breedlovei (Pilute Buckwheat) · E. breedlovei var. breedlovei (Piute Buckwheat) · E. breedlovei var. shevockii (Needle's Buckwheat) · E. brevicaule (Short-Stemmed Wild-Buckwheat) · E. brevicaule var. bannockense (Bannock Wild Buckwheat) · E. brevicaule var. brevicaule (Shortstem Buckwheat) · E. brevicaule var. cottamii (Cottam's Buckwheat) · E. brevicaule var. laxifolium (Goldenball Eriogonum) · E. brevicaule var. micranthum (Shortstem Buckwheat) · E. butterworthianum (Butterworth's Buckwheat) · E. caespitosum (Matted Buckwheat) · E. caespitosum var. sublineare (Matted Wild Buckwheat) · E. capillare (San Carlos Buckwheat) · E. capistratum (Hidden Buckwheat) · E. capistratum var. capistratum (Hidden Buckwheat) · E. capistratum var. muhlickii (Muhlick's Buckwheat) · E. capistratum var. welshii (Welsh's Buckwheat) · E. cernuum (Nodding Buckwheat) · E. cernuum var. cernum (Nodding Wild Buckwheat) · E. cernuum var. cernuum (Nodding Buckwheat) · E. cernuum var. purpurascens (Nodding Wild Buckwheat) · E. cernuum var. viminale (Nodding Buckwheat) · E. cespitosum (Matted Buckwheat) · E. chrysops (Bitterroot Buckwheat) · E. cinereum (Coastal Buckwheat) · E. cithariforme (Cithara Buckwheat) · E. cithariforme var. agninum (Cithara Buckwheat) · E. cithariforme var. cithariforme (Cithara Buckwheat) · E. clavatum (Hoover's Desert Trumpet) · E. clavellatum (Clay-Loving Wild-Buckwheat) · E. codium (Basalt Desert Buckwheat) · E. collinum (Hill Buckwheat) · E. coloradense (Colorado Buckwheat) · E. compositum (Arrowleaf Buckwheat) · E. compositum var. compositum (Arrowleaf Buckwheat) · E. compositum var. lancifolium (Arrowleaf Buckwheat) · E. compositum var. leianthum (Arrowleaf Buckwheat) · E. compositum var. pilicaulis (Arrowleaf Wild Buckwheat) · E. concinnum (Darin Buckwheat) · E. congdonii (Congdon Buckwheat) · E. contiguum (Ash Meadows Buckwheat) · E. contortum (Grand Buckwheat) · E. correllii (Correll's Buckwheat) · E. corymbosum (Corymbed Buckwheat) · E. corymbosum var. aureum (Crispleaf Buckwheat) · E. corymbosum var. corymbosum (Crispleaf Buckwheat) · E. corymbosum var. glutinosum (Crispleaf Buckwheat) · E. corymbosum var. heilii (Crisp-Leaf Wild Buckwheat) · E. corymbosum var. nilesii (Crisp-Leaf Wild Buckwheat) · E. corymbosum var. orbiculatum (Crispleaf Buckwheat) · E. corymbosum var. revealianum (Crispleaf Buckwheat) · E. corymbosum var. thompsonae (Crisp-Leaf Wild Buckwheat) · E. corymbosum var. velutinum (Crispleaf Buckwheat) · E. covilleanum (Coville's Buckwheat) · E. crocatum (Conejo Buckwheat) · E. cronquistii (Bull Mountain Buckwheat) · E. crosbyae (Crosby's Buckwheat) · E. cusickii (Cusick's Buckwheat) · E. darrovii (Carrot Buckwheat)

More Info

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Further Reading

• 1997 IUCN red list of threatened plants Cambridge: IUCN, World Conservation Union, 1998 url p. 462.
• An illustrated flora of the northern United States, Canada and the British possessions: from Newfoundland to the parallel of the southern boundary of Virginia and from the Atlantic Ocean westward to the 102nd meridian / by Nathaniel Lord Britton and Hon. Addison Brown. New York: Scribner, 1913. url p. 648, p. 651.
• Botanical Museum leaflets, Harvard University. 8 1940 Cambridge, Mass.: Botanical Museum, Harvard University, 1932- url p. 159.
• Botany Cambridge, Mass., John Wilson and Son, 1880 url p. 28.
• Botany. By W. H. Brewer, Sereno Watson, and Asa Gray. Boston, Little, Brown, 1880. url p. 28.
• Botany. Washington, Govt. Print. Off., 1871. url .
• Brigham Young University science bulletin. 13 1971 Provo, Utah: Brigham Young University, [1955-1976] url p. 1, p. 3, fig. 22, page 36, p. 44.
• Bulletin of the Torrey Botanical Club. 17 1890 New York: Torrey Botanical Club, 1870-1996 url p. 324, p. 55.
• Bulletin of the United States Geological and Geographical Survey of the Territories / Department of the Interior. Washington: Government Printing Office, 1874- url p. 242.
• Contributions from the New York Botanical Garden. New York: The Garden, 1899- url .
• Contributions from the United States National Herbarium 25 1925 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 155, p. 185, p. 186, p. 190, p. 235.
• Flora of Colorado, Fort Collins, Col., Experiment Station, 1906. url .
• Flora of Colorado, by P. A. Rydberg, PH. D. Fort Collins, Col., Experiment Station, 1906. url p. 106.
• Flora of Colorado, by P.A. Rydberg. Fort Collins, Col., Experiment Station, 1906. url p. 106, p. 106.
• Flora of Nebraska; a list of the conifers and flowering plants of the state with keys for their determination. Plainview, Neb.[c1912] url .
• Flora of Nebraska; a list of the ferns, conifers and flowering plants of the state with keys for their determination. Plainview, Neb., 1923 url p. 85.
• Flora of New Mexico / by E.O. Wooton and Paul C. Standley. Washington: G.P.O., 1915. url p. 190.
• Flora of the state of Washington / by Charles V. Piper. Washington: G.P.O., 1906 url p. 235.
• Great Basin naturalist memoirs. 1987 [Provo, Utah]Brigham Young University, 1976-1992. url p. 477.
• IX.
• List of Pteridophyta and Spermatophyta growing without cultivation in northeastern North America. Prepared by a Committee of the Botanical Club, American Association for the Advancement of Science. New York, 1894. url p. 137.
• Manual of the flora of the northern states and Canada / by Nathaniel Lord Britton. New York: Holt, 1905. url p. 351, p. 353.
• National list of scientific plant names. [Washington, D.C.]: U.S. Dept. of Agriculture, Soil Conservation Service, 1982- url p. 159, p. 160, p. 161.
• Native woody plants of the United States, their erosion-control and wildlife values. Washington, U. S. Govt. print. off., 1938. url p. 122.
• New manual of botany of the central Rocky mountains (vascular plants) Cincinnati [etc.]American Book Company[c1909] url .
• North American fauna. Washington: Fish and Wildlife Service; for sale by the Supt. of Docs., U. S. Govt. Print. Off. url p. 28.
• Phytogeography of Nebraska. 1. General survey by Roscoe Pound and Frederic E. Clements. Lincoln, Neb.Published by the Seminar1900 url p. 385, p. 425, p. 84.
• Phytologia memoirs. Plainfield, N.J.: H.N. Moldenke and A.L. Moldenke, 1980- url p. 228.
• Phytologia. Bronx Park, New York, H.A. Gleason and H.N. Moldenke, url p. 125, p. 208, p. 216, p. 252, p. 293, p. 483.
• Report of the geological exploration of the fortieth parallel / Washington, D.C.: G.P.O., 1870-80 url , p. 303.
• The Gardeners' chronicle: a weekly illustrated journal of horticulture and allied subjects. London: [Gardeners Chronicle], 1874-1955. url p. 525.
• The Great Basin naturalist. 41 1981 Provo, Utah: M.L. Bean Life Science Museum, Brigham Young University, 1939-1999. url p. 114, p. 145, p. 16, p. 168, p. 168, brotherson & brotherson, page 177, reveal, page 183, p. 185, p. 189, p. 190, p. 190, p. 191, p. 191, p. 191, p. 198, p. 199, p. 202, p. 203, p. 205, p. 206, map 3, page 209, fig. 8, page 211, fig. 9, page 214, fig. 10, page 215, p. 22, fig. 15, page 227, p. 228, p. 241, p. 242, p. 27, p. 29, p. 293, p. 30, p. 31, p. 362, p. 375, p. 4, p. 433, p. 5, p. 502, p. 533, p. 540, p. 77, p. 86, p. 89, p. 91.
• The flora of southwestern Colorado / by T.S. Brandegee. Washington: Dept. of the Interior, U.S. Geological and Geographical Survey of the Territories, 1876. url p. 242.
• Watson, S. Botany /by Sereno Watson, aided by Daniel C. Eaton, and others. 5 1871 Washington, D.C.: G.P.O., 1871. url p. 303, p.
• Zoe:a biological journal. 4 1893 San Diego, Calif. [etc.]Zoe Publishing Co. url p. 11, p. 12, p. 12.
• Li Anjen, Kao Tsoching, Mao Zumei & Liu Yulan. 1998. Polygonaceae. In: Li Anjen, ed., Fl. Reipubl. Popularis Sin. 25(1): 1209.
• Reveal, J. L. 1973b. Eriogonum (Polygonaceae) of Utah. Phytologia 25: 169-217.
• Reveal, J. L. 1976. Eriogonum (Polygonaceae) of Arizona and New Mexico. Phytologia 34: 409-484.
• Reveal, J. L. 1985. An annotated key to Eriogonum (Polygonaceae) of Nevada. Great Basin Naturalist 45: 493-519.
• Stokes, S. G. 1936. The Genus Eriogonum, a Preliminary Study Based on Geographical Distribution. San Francisco.

Notes

Contributors

Data Sources

Accessed through GBIF Data Portal February 02, 2008:

• Harvard University Herbaria, Harvard University Herbaria
• Missouri Botanical Garden, Missouri Botanical Garden
• School of Life Sciences, Arizona State University, Arizona State University Vascular Plant Herbarium
• School of Life Sciences, Arizona State University, The Deaver Herbarium, Northern Arizona University
• The New York Botanical Garden, Vascular Plant Type Specimens
• USDA PLANTS, USDA PLANTS Database
• University of Washington Burke Museum, Vascular Plant Collection - University of Washington Herbarium
• Utah State University, USU-UTC Specimen Database
• Utah Valley State College
• , Utah Valley State College Herbarium

Identifiers

• Biodiversity Heritage Library NamebankID: 2647204
• Catalogue of Life Accepted Name Code: ITS-21103
• Global Biodiversity Information Facility Taxonkey: 13741778
• Globally Unique Identifier: urn:lsid:ipni.org:names:694087-1
• GRIN Nomen Number: 316668
• Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 21103
• International Plant Names Index (IPNI) ID: 94825-2
• Natural Heritage Network Species Identifier: PDPGN081E0
• U.S.D.A. Plant Symbol: ERMIC3 ERC
• Zipcode Zoo Species Identifier: 38062

Footnotes

1. Anjen Li, Bojian Bao, Alisa E. Grabovskaya-Borodina, Suk-pyo Hong, John McNeill, Sergei L. Mosyakin, Hideaki Ohba & Chong-wook Park "Polygonaceae". in Flora of China Vol. 5 Page 277. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
2. James L. Reveal "Eriogonum". in Flora of North America Vol. 5. Oxford University Press. Online at EFloras.org. [back]
3. Mean = 1,522.770 meters (4,995.965 feet), Standard Deviation = 1,159.550 based on 146 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]

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