Overview
Interesting Facts
Common Names
Common Names in English:
Spreading Daisy, Spreading Fleabane
Description
Family Compositae
The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.
Genus Erigeron
Annuals
, biennials, or perennials [subshrubs
, shrubs
, trees
], (0.5-) 2-90(-100) cm (taprooted, fibrous-rooted, or rhizomatous
and fibrous-rooted, sometimes with simple
or branched caudices, sometimes stoloniferous
) . Stems erect
to ascending
, decumbent
, or prostrate
, simple or branched, glabrous
or hairy
, sometimes glandular
(hairs
2-seriate, minute, sometimes stipitate
) . Leaves basal and/or cauline (basal persistent or not to flowering) ; alternate; sessile or petiolate
; blades
1-nerved (3-nerved), linear
to lanceolate, oblanceolate
, or spatulate
(bases
sometimes clasping
), margins
entire or ± dentate
to pinnatifid
, faces
glabrous or hairy, sometimes glandular. Heads usually radiate
, sometimes discoid
or disciform
(erect, nodding
, or arching-pendent in bud), borne singly or in loose
, corymbiform
or paniculiform
arrays. Involucres turbinate
to hemispheric
, 5-35 mm diam. Phyllaries 30-125(-150) in 2-5 series, 1- or 3-nerved (nerves
golden-resinous; usually flat, rarely broadly keeled
to convex
), narrowly elliptic- to linear-lanceolate, unequal to equal, margins scarious
or not, faces hairy or glabrous, sometimes glandular. Receptacles flat to conic, pitted
, epaleate. Ray florets 0 or 12-350 in 1(-2+) series, pistillate
, fertile
; corollas usually white to bluish or purplish to pink, less commonly yellow (coiling
from apices, reflexing at tube/lamina junction, or remaining ± straight and spreading
) . Peripheral florets (disciform heads) 50-200 in 1-4 series, pistillate. Disc florets 25-450, bisexual
, fertile; corollas yellow (nerves orange-resinous), tubes
shorter than usually tubular
, sometimes strongly inflated
and indurate
throats
, lobes
5, erect to spreading, deltate; style-branch appendages
mostly deltate (papillate
) . Cypselae (tan) oblong
to oblong-obovoid, compressed
to flattened, 2(-4) -nerved, or subterete, 5-14-nerved (sect. Wyomingia and some other species), faces glabrous or strigose
or sericeous
, eglandular
; pappi persistent
or readily falling, usually of outer setae or scales
(0.1-0.4 mm), sometimes connate
, plus 5-40(-50), stramineous
, barbellate
bristles
, sometimes pappi only on ray or only on disc cypselae, or 0. x = 9.
Species ca. 390: nearly worldwide, mostly in temperate regions
.
The North American and Central American species of Erigeron have been divided
into sections
(G. L. Nesom 1989c, 1990g, 1994b; Nesom and R. D. Noyes 1999), emphasizing variation
in habit (especially taprooted versus rhizomatous and fibrous-rooted), vestiture
, arrangement
of heads
in arrays and orientation before flowering (erect, nodding, or arching-pendent), behavior of ray corolla laminae
(straight, reflexing, or coiling), cypsela and pappus morphology, and other morphologic features. The sequence and groupings of species treated here reflect significant modifications of earlier arrangements.
G. L. Nesom (1989d) hypothesized that Trimorpha [Erigeron sect. Trimorpha (Cassini) de Candolle] is separate from Erigeron, more closely related to Conyza. Studies by W. Huber and colleagues (e.g.
, Huber 1993; Huber and Ö. Nilsson 1995) and R. D. Noyes (2000) have shown that Trimorpha species are closely related to those of sect. Erigeron and that both sections are relatively recently derived within the genus. As suggested by Nesom (1994b) and by Huber and Nilsson, and as discussed in detail and experimentally confirmed by Noyes, autogamous
breeding systems apparently have arisen independently in groups of Astereae, including Trimorpha and Conyza, where the pistillate florets of a head are greatly increased in number (often outnumbering the bisexual florets), in multiple
series, the inner sometimes with filiform
, elaminate
corollas, and the outer with reduced laminae.
In the molecular analysis by R. D. Noyes (2000), Conyzinae comprises Erigeron, American Conyza, the four genera of the South American Leptostelma group, and the North American Aphanostephus; the cladistically basal and terminal
taxa of the subtribe
are members
of Erigeron. Noyes (p. 107) observed that "strictly speaking, although the Conyzinae form a monophyletic group [with caveats regarding Old World Conyza], Erigeron is paraphyletic, as five other genera are derived from within it." The molecular study included
46 of the 173 species treated here.
Polyploidy is common among species of Erigeron, and agamospermy apparently is a common correlate of polyploidy, especially in odd-polyploid plants
. Molecular phylogenetic
data (R. D. Noyes 2000) indicate that agamospermy has arisen at least three times within the genus.
In the descriptions
and keys
, some characteristics are assumed constant unless otherwise indicated (usually in parentheses) ; particular application
of terms
is discussed here. The indumentum of erigerons is often complex
; in order
to simplify descriptions, glabrous applies here only to absence of non-glandular hairs, eglandular to the absence of glandular hairs; a totally glabrous plant (in the usual sense) would be glabrous and eglandular. Petiole
margins are eciliate
or sparsely ciliate
unless otherwise indicated. Leaf bases of most erigerons are broadened or not, not thickened and white-indurate. Margins of leaves in some erigerons are entire but for tiny callous
enations
that correspond to the callous tips of teeth on some leaves with serrate margins. Here, margins with such tiny enations are described as denticulate. Heads of some erigerons are "pseudodisciform" in the sense that the outer pistillate florets have relatively small, ± filiform laminae (such florets
are technically "ray florets" even though their "rays" are inconspicuous) and the inner pistillate florets have no laminae on their corollas. The distinction between corollas without and those with laminae is sometimes arbitrary
. Ray laminae are considered strap-shaped
and spreading unless otherwise indicated. Descriptions of ray color as "blue" should be read
as lavender-blue.[1]
Physical Description
Species Erigeron divergens
Annuals
or short-lived perennials
, (7-) 12-40(-70) cm; taprooted,
caudices simple
if perennial. Stems (single or multiple
from
bases
) erect
to ascending
, densely and evenly puberulous-hirsutulous
(hairs
spreading
to spreading-descending or -ascending, often crinkly,
bases not thickened), minutely glandular
at least distally, usually
throughout. Leaves basal (usually deciduous) and cauline;
basal blades obovate-spatulate, 10-70 × 4--14 mm, cauline gradually
reduced distally, margins
entire or with 2-3 pairs of teeth or lobes
,
faces
hirsute
to loosely strigoso-hirsute, sometimes sparsely glandular.
Heads (in early season
, 1-) 5-100+ (buds nodding
). Involucres
3-4 × (5-) 7-11 mm.
Phyllaries in 3-4 series, hirsute,
minutely glandular. Ray florets 75-150; corollas white, drying
lilac, without abaxial
midstripe, (2-) 4-9.5 mm, laminae
not coiling
or reflexing (remaining relatively straight). Disc corollas
1.8-2.3 mm (throats
indurate
and slightly inflated
). Cypselae
0.9-1.2 mm, 2-nerved (nerves
whitish), faces sparsely strigose
; pappi:
outer of setae or scales
, inner of 6-9(-12) bristles
. 2n
= 18, 27, 36. [source]
Polyploidy and agamospermy apparently are common in Erigeron divergens
and contribute to the variability and, probably to some extent, the
polymorphism
characteristic of this species. Diploids appear to be
scattered
through the range
of the species, at least in its southern
part. [source]
The form of Erigeron divergens with persistent
, deeply pinnatifid
basal leaves
with relatively long petioles
and tendency toward perenniality
(woody taproots
) has been named E. accedens [occurs at 200-1000(-1200)
m
and flowers Feb-Apr(-May) ]. Such plants
have sometimes been identified
as E. lobatus, and it is possible that they may have originated
as hybrids with it; in most ways they are similar to E. divergens
and apparently intergrade
with it. The E. accedens form occurs
in California, Arizona, New Mexico, southwest Texas, Baja California,
northern Sonora, Durango, and Chihuahua, apparently sympatrically
with more typical E. divergens, suggesting that the former
is stabilized and self-reproductive (probably agamospermic). Available
chromosome counts are triploid. Rare plants (e.g.
, E. incomptus)
may have filiform
rays barely extending past the involucre. [source]
Another common form of Erigeron divergens in the broad sense
(mostly in Chihuahua, also southern Texas, New Mexico, and Arizona)
has bowl-shaped plants with decumbent
stems, a discrete taproot,
persistent, lobed
basal leaves, and solitary heads
topping usually
simple stems, and usually is early-season in phenology. Further comments
related to variation
in E. divergens are found under 163.
Erigeron multiceps. [source]
Habit: Forb/herb
Flowers: Bloom Period: February, March, April, May, June, July, August, September, October. • Flower Color: near white, white
Size/Age/Growth
Size: 6-12" tall.
Habitat
Gravelly or sandy flats, riverbanks, meadows, disturbed sites, desert scrubland to grasslands, saltbush, blackbrush, sagebrush, pinyon-juniper, juniper-oak, oak, and ponderosa pine woodlands; 60-2000(-2700) m
Typically found at an altitude of 0 to 3,180 meters (0 to 10,433 feet).[2]
Biology
Reproduction
Duration: Biennial
Growth
Culture: Space 12-15" apart.
Soil: Minimum pH: 6.6 • Maximum pH: 7.8
Sunlight: Sun Exposure: Full Sun .
Moisture: Drought Tolerance: High
Temperature: Cold Hardiness: 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b, 9a, 9b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Campanulanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Asterales
(
)
- Lindley, 1833
- Family:
Compositae
(
)
- Giseke, 1792, nom. cons., nom. alt.
- Subfamily:
Asteroideae
(
)
- Tribe:
Astereae
(
)
- Subtribe:
Conyzinae
(
)
- Genus:
Erigeron
(
)
- C. Linnaeus, 1753
- Fleabane [Greek eri, early, or erio, woolly, and geron, old man, perhaps alluding to pappus, which becomes gray and accrescent in some species, or to solitary, woolly heads of some of species]
- Specific epithet:
divergens
- Torr. & A.Gray
- Botanical name: - Erigeron divergens Torr. & A.Gray
- Specific epithet:
divergens
- Torr. & A.Gray
- Genus:
Erigeron
(
- Subtribe:
Conyzinae
(
- Tribe:
Astereae
(
- Subfamily:
Asteroideae
(
- Family:
Compositae
(
- Order:
Asterales
(
- Superorder:
Campanulanae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
E. accedens Greene • E. incomptus A. Gray • E. solisaltator G. L. Nesom • Erigeron divaricatus Nuttall • Erigeron divergens var. typicus Cronq. • Trans. Amer. Philos. Soc.
Notes
Name
Status: Accepted Name
.
Comment: Data Providers: CONABIO, Govaerts World Compositae Checklist
A-G, IPNI, Tropicos. GCC LSID: urn
:lsid:compositae.org:names:03250E34-FCA7-4516-80A8-73D5E7D800CB
Last scrutiny: 13-Aug-09
Similar Species
Members of the genus Erigeron
ZipcodeZoo has pages for 358 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
E. abajoensis (Abajo Daisy) · E. acer (Blue Fleabane) · E. acer acer (Bitter Fleabane) · E. acomanus (Acoma Fleabane) · E. acris (Bitter Boreal Daisy) · E. acris debilis (Bitter Fleabane) · E. acris politus (Bitter Fleabane) · E. aequifolius (Hall's Daisy) · E. algidus (Sierra Daisy) · E. aliceae (Alice Eastwood's Fleabane) · E. allocotus (Big Horn Fleabane) · E. alpiniformis (Alpine Fleabane) · E. anchana (Sierra Ancha Fleabane) · E. angustatus (Serpentine Fleabane) · E. annuus (Annual Fleabane) · E. annuus annuus (Eastern Daisy Fleabane) · E. aphanactis (Beach Daisy) · E. aphanactis var. aphanactis (Rayless Shaggy Fleabane) · E. aphanactis var. congestus (Rayless Shaggy Fleabane) · E. arenarioides (Sand Fleabane) · E. argentatus (Silver Fleabane) · E. arisolius (Arid Throne Fleabane) · E. arizonicus (Arizona Fleabane) · E. asper (Rough Fleabane) · E. asperugineus (Idaho Daisy) · E. aurantiacus (Double Orange Daisy) · E. aureus (Alpine Yellow Fleabane) · E. aureus Greene var. acutifolius Raup (Alpine Yellow Fleabane) · E. aureus var. acutifolius (Alpine Yellow Fleabane) · E. aureus var. aureus (Alpine Yellow Fleabane) · E. austiniae (Dwarf Yellow Daisy) · E. awapensis (Awapa Daisy) · E. 'Azure Fairy' (Azure Fairy Fleabane) · E. barbellulatus (Shining Fleabane) · E. basalticus (Basalt Daisy) · E. bellidiastrum (Western Daisy Fleabane) · E. bellidiastrum Nutt. var. arenarius (Greene) Nesom (Sandwort Daisy Fleabane) · E. bellidiastrum var. arenarius (Sandwort Daisy Fleabane) · E. bellidiastrum var. bellidiastrum (Western Daisy Fleabane) · E. bellidiastrum var. robustus (Western Daisy Fleabane) · E. bellioides (Bellorita) · E. bigelovii (Bigelow's Fleabane) · E. biolettii (Biolett's Erigeron) · E. bistiensis (Bisti Fleabane) · E. blochmaniae (Blochman Leafy Daisy) · E. bloomeri (Bloomer Fleabane) · E. bloomeri var. bloomeri (Scabland Fleabane) · E. bloomeri var. nudatus (Scabland Fleabane) · E. bloomeri var. nudatus (Gray) Cronq. (Scabland Fleabane) · E. bloomeri var. pubens (Scabland Fleabane) · E. borealis (Boreal Fleabane) · E. breweri (Brewer's Daisy) · E. breweri var. bisanctus (Brewer's Erigeron) · E. breweri var. breweri (Brewer's Fleabane) · E. breweri var. covillei (Coville's Erigeron) · E. breweri var. covillei (Greene) Nesom (Coville's Erigeron) · E. breweri var. jacinteus (Brewer's Fleabane) · E. breweri var. jacinteus (Hall) Cronq. (Brewer's Fleabane) · E. breweri var. klamathensis (Klamath Erigeron) · E. breweri var. klamathensis Nesom (Klamath Erigeron) · E. breweri var. porphyreticus (Brewer's Fleabane) · E. caespitosus (Caespitose Fleabane) · E. caespitosus capillaris (Tufted Fleabane) · E. caespitosus uncialis (Tufted Fleabane) · E. caespitosus var. caninum (Tufted Fleabane) · E. caespitosus var. pectinacea (Tufted Fleabane) · E. calvus (Bald Daisy) · E. canus (Hoary Fleabane) · E. cascadensis (Cascade Fleabane) · E. cervinus (Siskiyou Daisy) · E. chrysopsidis (Dwarf Yellow Fleabane) · E. clokeyi (Clokey's Daisy) · E. colomexicanus (Running Daisy) · E. compactus (Cushion Daisy) · E. compactus var. cognatum (Cushion Daisy) · E. compactus var. coloradoensis (Cushion Daisy) · E. compactus var. covillei (Cushion Daisy) · E. compactus var. praebens (Cushion Daisy) · E. compositus (Cutleaf Daisy) · E. compositus var. compositus (Dwarf Mountain Fleabane) · E. concinnus (Hairy Daisy) · E. concinnus var. concinnus (Navajo Fleabane) · E. concinnus var. condensatus (Navajo Fleabane) · E. concinnus var. subglaber (Navajo Fleabane) · E. consimilis (Fern-Leaf Fleabane) · E. corymbosus (Long-Leaf Fleabane) · E. coulteri (Coulter Fleabane) · E. covillei (Coville's Erigeron) · E. cronquistii (Cronquist Daisy) · E. cuneifolius (Wedgeleaf Fleabane) · E. 'Darkest of All' (Fleabane) · E. decumbens (Meadow Fleabane) · E. decumbens var. decumbens (Willamette Fleabane) · E. decumbens var. decumbens Nutt. (Willamette Fleabane) · E. decumbens var. robustior (Robust Daisy) · E. disparipilus (White Cushion Fleabane) · E. divergens (Spreading Daisy) · E. eatonii (Eaton Fleabane) · E. eatonii Gray var. eatonii Gray (Eaton Fleabane) · E. eatonii Gray var. lavandulus Strother & Ferlatte (Eaton's Fleabane)
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Further Reading
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Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Data Sources
Accessed through GBIF Data Portal February 16, 2008:
- Australian National Herbarium
- , Australian National Herbarium
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- Comisión nacional para el conocimiento y uso de la biodiversidad, Herbario del Instituto de Ecología, A.C., México
- Missouri Botanical Garden, Missouri Botanical Garden
- School of Life Sciences, Arizona State University, Arizona State University Vascular Plant Herbarium
- School of Life Sciences, Arizona State University, The Deaver Herbarium, Northern Arizona University
- USDA PLANTS, USDA PLANTS Database
- University of Colorado Museum, Zoological specimens
- Utah State University, USU-UTC Specimen Database
- Utah Valley State College
- , Utah Valley State College Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2657472
- Catalogue of Life Accepted Name Code: Ast-18809
- Global Biodiversity Information Facility Taxonkey: 13748571
- Globally Unique Identifier: urn:lsid:ipni.org:names:203788-1
- GRIN Nomen Number: 448141
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 35852
- MoBot NameID: 2717984
- Natural Heritage Network Species Identifier: PDAST3M160
- U.S.D.A. Plant Symbol: ERDIT
- Zipcode Zoo Species Identifier: 37802
Footnotes
- Guy L. Nesom "Erigeron". in Flora of North America Vol. 20 Page 3,9, 11, 12, 14, 17, 36, 204, 256, 257, 334. Oxford University Press. Online at EFloras.org. [back]
- Mean = 1,424.300 meters (4,672.900 feet), Standard Deviation = 657.680 based on 785 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
