Description
Family Arecaceae
Trees
or shrubs
[lianas], perennial
, branched or unbranched, solitary or clustered. Roots
adventitious, thick. Stems woody, subterranean
or terrestrial
, creeping
or erect
[climbing
], slender or massive, sometimes conspicuously enlarged and storing starch
and water, smooth
or covered with fibrous
or prickly remains of leaf bases
. Leaves spirally arranged
; sheaths
tubular
, often forming crownshaft
, sometimes with ligular appendages
; petioles
terete
, channeled
, or ridged
, unarmed
or bearing prickles or marginal
teeth; hastula (flap of tissue
from petiole apex at junction with surface of blade
) absent or present adaxially, rarely present abaxially. Leaf blade palmate, costapalmate
(intermediate between palmate and pinnate), pinnate, or 2-pinnate [undivided]; plication
(folding lengthwise into pleats or furrows
) ^ - or tent-shaped (reduplicate
, splitting
along abaxial
ridges
) or V-shaped (induplicate
, splitting along adaxial
ridges) ; segments lanceolate, linear
, or cuneate [rhombic
], glabrous
or variously scaly
, unarmed or bearing prickles (proximal
segments modified into spines in Phoenix) . Inflorescences from solitary [clustered] axillary buds, borne within, below, or above crown of leaves, paniculate
, rarely spicate
, usually branched to 1--5 orders
; prophyll (1st bract on main inflorescence axis
) 2-keeled; peduncular bract(s) (empty bract[s] between 1st prophyll and 1st bract subtending branch
) present [absent]; flowers bisexual
, unisexual
with staminate
and pistillate
on same plants
or on different plants, or both bisexual and unisexual on same plant. Flowers solitary or variously clustered along rachillae of inflorescence, radially symmetric
; perianth 1--2-seriate; sepals [2--]3[--4], distinct
or connate
; petals [2--]3[--4], distinct or variously connate; androecium: stamens [3--]6--34[--1000]; filaments
distinct or connate or basally adnate
to petals; anthers
basifixed
or dorsifixed
, dehiscing latrorsely or introrsely; staminodes in pistillate flowers distinct or variously connate or adnate to pistil or petals; pistils 1 or 3, distinct or partially connate, each bearing 1 ovule and 1 stigma, or 1 pistil bearing 1--3 ovules and 3 stigmas; styles
distinct or connate, short; stigmas dry; pistillode
in staminate flower
present or absent. Fruits drupaceous
or berrylike; stigmatic
remains basal or apical; exocarp
smooth, warty, prickly, or hirsute
[corky or scaly]; mesocarp
fleshy
or dry and fibrous; endocarp papery
, leathery, or bony, sometimes with 3 germination pores
. Seeds 1(--2+), free
or adhering to endocarp; seed coat
thin; endosperm homogeneous
or ruminate
, sometimes penetrated by seed coat; embryo basal, lateral
, or apical, peglike, minute; eophyll
(1st seedling leaf with blade) undivided and lanceolate or 2-cleft [pinnate].
Genera 1914, species ca.
2500 (19 genera, 29 species in the flora
) : worldwide, especially abundant in Central America, South America, se Asia.
Although palms appeared in various taxonomic
schemes since the time of Linnaeus, the first attempt at a modern phylogenetic
classification of the palms was published by H. E. Moore Jr. (1973) . Moore left his "major groups" unranked, and his untimely death
in 1980 prevented his completing a formal synthesis. J. Dransfield and N. W. Uhl (1986) gave formal ranks
to Moore€™s groups and divided
the family
into six subfamilies and numerous
tribes
and subtribes
. Their Genera Palmarum (N. W. Uhl and J. Dransfield 1987, 1999) is a model
of accuracy and completeness and will long serve the needs of the scientific, horticultural, and resource-management communities. With the advent of molecular techniques and a resurgence in palm research, however, realignments in the classification may be expected, and indeed additional data already require some changes in the current
scheme (A. Barford 1991; R. G. Bernal et al.
1991; J. L. Dowe and N. W. Uhl 1989; J. Dransfield 1989, 1991; J. Dransfield and H. J. Beentje 1995, 1995b; A. Henderson and M.
J. Balick 1991; N. W. Uhl and J. Dransfield 1999; N. W. Uhl et al. 1990, 1995.)
Modern cladistic analyses place the palms as the sister group
to the Commelinanae
clade (M. W. Chase et al. 1993; J. I. Davis 1995; M. R. Duvall et al. 1993b), with which they share ultraviolet-fluorescent phenolic compounds
in their cell walls
and Strelitzia-type epicuticular wax
morphology (W. Barthlott and D. Frölich 1983; P. J. Harris and R. D. Hartley 1980) . Palms are currently treated as the sole
representative of the superorder Arecanae
, order Arecales (R. M. T. Dahlgren et al. 1985; R. F. Thorne 1992b) .
Morphologically the family is diverse
and complex
(see especially P. B
. Tomlinson 1990) . The majority of palms produce
a single indeterminate stem with axillary
inflorescences; several noteworthy departures, however, also occur in numbers of vegetative
and floral
axes, position of inflorescence, and displacement of terminal
bud. Stems may be solitary (monopodial) or clustered (sympodial), erect, prostrate
, or lianoid. A majority of palms have unbranched vegetative axes, although aerial
branching, sometimes dichotomous, is known in a variety of unrelated genera (e.g.
, Korthalsia Blume, Nannorrhops H. Wendland) . Branching may also be nonaxiallary in some genera (J. B. Fisher
et al. 1989) .
Studies of pollination (F. Borchsenius 1997; F. Ervik and J. P. Feil 1997; A. Henderson 1986; C.
Listabarth 1992, 1993, 1993b, 1994; A. O. Scariot et al. 1991) indicate that insect pollination, especially by beetles (Coleoptera), bees and wasps (Hymenoptera), and flies (Diptera), is apparently more common than wind pollination. Bats (Chiroptera) play a role in the pollination of some species (S. A. Cunningham 1995) .
Dispersal
of seeds is generally by means of animals for fleshy-fruited palms (S. Zona and A. Henderson 1989) . Many species of mammals include palm fruits in their diets
(S. H. Bullock 1980; R. F. Harlow 1961; W. D. Klimstra and A. L. Dooley 1990; D. S. Maehr 1984; D. S. Maehr and J. R. Brady 1984), but birds also play a significant role. In the Eastern Hemisphere, Cocos Linnaeus and Nypa Steck have achieved a wide distribution as the result of dispersal by water. For the relationship
between palms and seed-eating bruchid beetles (Bruchidae: Pachymerinae: Pachmerini), see C. D. Johnson et al. (1995) .Scott Zona "Arecaceae". in Flora of North America Vol. 22 Page 95. Oxford University Press. Online at EFloras.org.
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Liliopsida
(
)
- Scopoli, 1760
- Subclass:
Arecidae
(
)
- Takhtajan, 1967
- Superorder:
Arecanae
(
)
- Takhtajan, 1967
- Order:
Arecales
(
)
- Bromhead, 1840
- Family:
Arecaceae
(
)
- Schultz-Schultzenstein, 1832
- Palm Family
- Genus:
Daemonorops
(
)
- Specific epithet:
asteracanthus
- Becc.
- Botanical name: - Daemonorops asteracanthus Becc.
- Specific epithet:
asteracanthus
- Becc.
- Genus:
Daemonorops
(
- Family:
Arecaceae
(
- Order:
Arecales
(
- Superorder:
Arecanae
(
- Subclass:
Arecidae
(
- Class:
Liliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Similar Species
Members of the genus Daemonorops
There are approximately 189 species in this genus. Here are just 100 of them:
D. acamptostachys · D. acantholobus · D. accedens · D. acehensis · D. aciculatus · D. adspersus · D. affinis · D. angustifolia (Water Rattan) · D. angustifolius · D. angustispathus · D. annulatus · D. aruensis · D. asteracantha · D. asteracanthus · D. atra · D. aurea · D. bakauensis · D. banggiensis · D. barbatus · D. beguinii · D. binnendijkii · D. brachystachys · D. calapparius · D. calicarpa · D. calicarpus · D. calospatha · D. calothyrsus · D. carcharodon · D. clemensianus · D. cochleatus · D. collarifera · D. collariferus · D. confusus · D. congesta · D. crinita · D. crinitus · D. cristatus · D. curranii · D. curtisii · D. didymophylla · D. didymorphylla · D. dissitophyllus · D. diversispinus · D. draco · D. draconcellus · D. dracunculus · D. dransfieldii · D. elongata · D. elongatus · D. erinaceus · D. fasciculatus · D. fissa · D. fissus · D. floridus · D. forbesii · D. formicarius · D. fuscus · D. gaudichaudii · D. geniculata · D. geniculatus · D. gracilipes · D. gracilis · D. grandis · D. guruba · D. hallierianus · D. heteracanthus · D. hirsutus · D. horridus · D. hygrophilus · D. hypoleucus · D. hystrix · D. imbellis · D. ingens · D. intermedius · D. intumescens · D. javanicus · D. jenkinsiana · D. jenkinsianus · D. kiahii · D. kirtong · D. korthalsii · D. kunstleri · D. kunstlerii · D. kurzianus · D. laciniatus · D. lamprolepis · D. lasiospathus · D. latispinus · D. leptopus · D. lewisiana (Daemonorops) · D. lewisianus · D. loberianus · D. loherianus · D. longipedunculatus · D. longipes (Daemonorops) · D. longispatha · D. longispathus · D. longispinosus · D. longistipes · D. macrophylla
Bibliography
- Dransfield, J. and N. W. Uhl. 1986. An outline of a classification of palms. Principes 30: 3--11.
- Henderson, A. 1986. A review of pollination studies in the Palmae. Bot. Rev. 52: 221--259.
- Henderson, A., G. Galeano, and R. G. Bernal. 1995. Field Guide to the Palms of the Americas. Princeton.
- McClintock, E. 1993. Arecaceae [Palmae]. In: J. C. Hickman, ed. 1993. The Jepson Manual. Higher Plants of California. Berkeley, Los Angeles, and London. P. 1105.
- Moore, H. E. Jr. 1973. The major groups of palms and their distribution. Gentes Herb. 11: 27--141.
- Tomlinson, P. B. 1990. The Structural Biology of Palms. Oxford.
- Uhl, N. W. and J. Dransfield. 1987. Genera Palmarum. Lawrence, Kans.
- Zona, S. 1997. The genera of Palmae (Arecaceae) in the southeastern United States. Harvard Pap. Bot. 2: 71--107.
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Notes
Identifiers
- International Plant Names Index (IPNI) ID: 666423-1
- Zipcode Zoo Species Identifier: 3805178
