Common Names
Common Names in English:
Fischer's Orchid, Fisher's Orchids, Key Flower, Keyflower, Kodiak Keyflower
Description
Tribe Orchideae
surgical removal, e.g. ovariectomy, removal of the ovaries.
Genus Dactylorhiza
Herbs, perennial
, terrestrial
, rather succulent, glabrous
. Roots
from base
of stem fascicled tuberoids
, usually palmately divided
with 2-5 lobes
, fleshy
. Stems leafy. Leaves several, ascending
to recurved, not enfolded around spike, with or without purplish spots; base sheathing in proximal
leaves, distal leaves bractlike, not sheathing
. Inflorescences terminal
, spikes; floral
bracts foliaceous
, prominent
. Flowers few to many, resupinate; dorsal sepal, sometimes lateral
sepals, and petals connivent, forming hood
distal to lip; petals ± obliquely dilated
basally; lip 3-lobed, base spurred, margins
occasionally entire
, nectarless; pollinaria 2, each with 1 pollen mass; viscidia within single 2-lobed bursicle
; stigma reniform
or obcordate
, concave
with median ridge
, hidden behind
bursicle. Fruits capsules, ascending, ellipsoid
.
Species ca.
75: Alaska, Canada, mostly Eurasian.
Dactylorhiza is a taxonomically complex
genus in which closely related species have been combined into species aggregates
(P. Vermeulen 1947; R. M.
Bateman et al.
1997). Recognition of the aggregate taxa alone reduces the number of species by more than half. Recent research synthesizing morphometric
and allozyme
data to circumscribe species (R. M. Bateman and I. Denholm 1983, 1985, 1989; M. Hédren 1996), and DNA sequences and chromosome studies to determine the relationships
of those species (R. M. Bateman et al. 1997; A. M. Pridgeon et al. 1997), is shedding much light on the evolution of the genus. The diploid lineage
(2n = 40) appears to have evolved in Asia, migrating
and speciating to both the west and northeast. Several alloploidy
events (hybridization followed by chromosome doubling) occurred recently in Europe, apparently between the distinct
diploids D. fuchsii and D. incarnata. That generated a highly complex suite of poorly distinguishable prospecies of 2n = 80, treated as a single species by some authorities and as many species by others (L. V. Averyanov 1990). Of the two North American species, D. aristata is a native
diploid originating during the northeasterly migration, and D. majalis is an allotetraploid that originated in Europe and presumably is naturalized
in North America (H. J. Clase and S. J. Meades 1996).
Despite an extensive literature, much taxonomic
work still remains to be done.Charles J. Sheviak, Paul M. Catling, Susan J. Meades & Richard M. Bateman "Dactylorhiza". in Flora of North America Vol. 26 Page 491, 496, 577, 579, 580. Oxford University Press. Online at EFloras.org.
Physical Description
Species Dactylorhiza aristata
Plants
5-40 cm. Leaves 2-5; blade
often purple-spotted or suffused
with purple, proximal
blade 3-18 × 1-6 cm. Inflorescences:
floral
bracts lanceolate, 15-50 × 5-15 mm, the larger often
suffused with purple, extending well beyond flowers. Flowers magenta,
pink, or white; sepals prominently oblique
and erect
at base
, upwardly
curving forward or downward, ovate-lanceolate, 9-11(-13) ×
3-6 mm, awned
; petals ovate-lanceolate, 6-9 × 3-4 mm, awned;
lip often magenta- to red-spotted, suborbiculate to obcordate
, 8-12
× 8-12 mm, shallowly 3-lobed or entire
, tip
of middle
lobe
apiculate
; spur 5.5-7.5 mm, 1/2-2/3 as long as ovary. 2n = 40, 42.
[source]
Dactylorhiza aristata var. kodiakensis has been separated primarily
on the basis of its being uniform
instead of variable like the typical
variety (C.
A. Luer 1975) ; supplementary characters all partially
overlap. While this variety is not recognized here, variability in
the Alaskan Fischer's orchids merits detailed taxonomic
study. [source]
Habit: Forb/herb
Flowers: Bloom Period: April, May, June.
Habitat
Moist open tundra
, meadows, bogs
; 0--800 m
(Ref. 53778).
Biology
Reproduction
Duration: Perennial
Taxonomy
- Kingdom:
Plantae
(
)
- Plants
- Phylum:
Magnoliophyta
(
)
- Flowering Plants
- Class:
Liliopsida
(
)
- Order:
Asparagales
(
)
- Family:
Noctuoidea
(
)
- Subfamily:
Orchidoideae
(
)
- Tribe:
Orchideae
(
)
- Subtribe:
Orchidinae
(
)
- Genus:
Dactylorhiza
(
)
- Necker ex Nevski, Trudy Bot. Inst. S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast.
- Orchis [Greek dactylos, finger, and rhiza, root, in reference to the fingerlike tuberoids of the more primitive species]
- Specific epithet:
aristata
- (Fischer ex Lindley) Soó, Nom. Nov. Gen. Dactylorhiza. 5. 1962.
- Botanical name: - Dactylorhiza aristata (Fisch. ex Lindl.) Soó
- Specific epithet:
aristata
- (Fischer ex Lindley) Soó, Nom. Nov. Gen. Dactylorhiza. 5. 1962.
- Genus:
Dactylorhiza
(
- Subtribe:
Orchidinae
(
- Tribe:
Orchideae
(
- Subfamily:
Orchidoideae
(
- Family:
Noctuoidea
(
- Order:
Asparagales
(
- Class:
Liliopsida
(
- Phylum:
Magnoliophyta
(
Unambiguous Synonyms
- Dactylorhiza aristata (Fisch. ex Lindl.) Soó
- Dactylorhiza aristata var. kodiakensis Luer & G. M. Luer
- Orchis aristata Fisch. ex Lindl.
- Orchis aristata Fischer ex Lindley, Gen. Sp. Orchid. Pl., 262. 1835
Notes
Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000
Similar Species
Members of the genus Dactylorhiza
There are approximately 501 species in this genus. Here are just 100 of them:
D. 'Ballerina' · D. 'Harold Esslemont' · D. 'Kilrymont' · D. 'Pink Leopard' · D. 'Tinney's Spotted' · D. affinis · D. alpestris · D. amblyoloba · D. angustata · D. aristata (Fisher's Orchids) · D. aristata f. alba · D. aristata f. perbracteata · D. aristata f. punctata · D. aristata f. rosea · D. aristata var. aristata (Keyflower) · D. aristata var. kodiakensis (Kodiak Keyflower) · D. aristata × fuchsii · D. armeniaca · D. aschersoniana · D. aschersoniana nothosubsp. wisniewskii · D. aschersoniana nothovar. wisniewskii · D. Atlanta · D. baldshuanica · D. baltica · D. bartonii · D. battandieri · D. baumanniana · D. baumanniana smolikana · D. Biskaya · D. bithynica · D. bohemica · D. bosniaca · D. braunii · D. braunii nm. monticola · D. braunii nothosubsp. lilacina · D. braunii nothosubsp. smitakii · D. brennensis · D. Calibra · D. cambrensis · D. cantabrica · D. caramulensis · D. carnea · D. carnea nothosubsp. maculatiformis · D. carpatica · D. cataonica · D. caucasica · D. chuhensis · D. cilicia · D. cilicica · D. coccinea · D. comosa · D. comosa cambrensis · D. comosa f. kerryensis · D. comosa f. occidentalis · D. comosa majalis · D. comosa occidentalis · D. comosa scotica · D. comosa turfosa · D. cordigera · D. cordigera f. albiflora · D. cordigera graeca · D. cordigera pindica · D. cordigera subsp. pindica · D. cordigera var. rhodopeia · D. cordigera var. vermionica · D. cruenta · D. cruenta f. ochrantha · D. cruenta lapponica · D. cruenta salina · D. cruenta subsp. lapponica · D. curenta · D. curvifolia · D. cyrenaica · D. czerniakowskae · D. delphinensis · D. dinglensis · D. dinglensis nothosubsp. robertsii · D. durandii · D. Dutch Angel · D. ebudensis · D. elata (Robust Marsh Orchid) · D. elata 'Alba' · D. elata 'Glasnevin' · D. elata 'Lydia' · D. elata ambigua · D. elata anatolica · D. elata brennensis · D. elata durandii · D. elata f. alba · D. elata f. cordata · D. elata f. leucantha · D. elata f. pallida · D. elata f. peltieri · D. elata iberica · D. elata sesquipedalis · D. elata var. brennensis · D. elata var. durandii · D. elata var. elongata · D. elata var. sesquipedalis · D. elata white-flowered
Bibliography
- Averyanov, L. V. 1990. A review of the genus Dactylorhiza. In: J. Arditti, ed. 1977+. Orchid Biology: Reviews and Perspectives. 7+ vols. Ithaca, N.Y. and New York. Vol. 5, pp. 159206.
- Bateman, R. M. and I. Denholm. 1983. A reappraisal of the British and Irish dactylorchids: 1. The tetraploid marsh orchids. Watsonia 14: 347376.
- Bateman, R. M. and I. Denholm. 1985. A reappraisal of the British and Irish dactylorchids: 2. The diploid marsh orchids. Watsonia 15: 321355.
- Bateman, R. M. and I. Denholm. 1989. A reappraisal of the British and Irish dactylorchids: 3. The spotted orchids. Watsonia 17: 319349.
- Bateman, R. M., A. M. Pridgeon, and M. W. Chase. 1997. Phylogenetics of subtribe Orchidinae (Orchidoideae, Orchidaceae) based on nuclear ITS sequences: 2. Infrageneric relationships and taxonomic revision to achieve monophyly of Orchis sensu stricto. Lindleyana 12: 113141.
- Hédren, M. 1996. Genetic differentiation, polyploidization and hybridization in northern European Dactylorhiza (Orchidaceae): Evidence from allozyme markers. Pl. Syst. Evol. 201: 3155.
- Pridgeon, A. M. et al. 1997. Phylogenetics of subtribe Orchidinae (Orchidoideae, Orchidaceae) based on nuclear ITS sequences: 1. Intergeneric relationships and polyphyly of Orchis sensu lato. Lindleyana 12: 89109.
- Stace, C. A. 1991. New Flora of the British Isles. Cambridge and New York. Pp. 11661173.
- Tyteca, D. and J.-L. Gathoye. 1993. On the morphological variability of Dactylorhiza praetermissa (Druce) Soó (Orchidaceae). Belg. J. Bot. 126: 8199.
- Vermeulen, P. 1947. Studies on Dactylorchids. Utrecht.
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Notes
Contributors
- "Dactylorhiza aristata". in Flora of North America Vol. 26 Page 578. Published by Oxford University Press. Online at EFloras.org.
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-2006. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed April 19, 2007.
- Global Biodiversity Information Facility. Accessed February 28, 2008. http://www.gbif.org Mediated distribution data from 2 providers.
- MBLWHOI Library: Universal Biological Index and Organizer. uBio.org accessed July 17, 2008.
- USDA, NRCS. 2005. The PLANTS Database, Version 3.5 (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA.
- World Checklist of Selected Plant Families
Data Sources
Accessed through GBIF Data Portal February 28, 2008:
- Burke Museum: Vascular Plant Collection - University of Washington Herbarium (WTU)
- University of Alaska Museum of the North: University of Alaska Museum of the North Herbarium (ALA)
Identifiers
- Biodiversity Heritage Library NamebankID: 2663044
- Catalogue of Life Accepted Name Code: Kew-55162
- Global Biodiversity Information Facility Taxonkey: 249393
- Globally Unique Identifier: urn:lsid:ipni.org:names:626576-1
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 193461
- Natural Heritage Network Species Identifier: PMORC0S010
- U.S.D.A. Plant Symbol: DAARK DAAR
- Zipcode Zoo Species Identifier: 34302
