Overview
Interesting Facts
Common Names
Click on the language to view common names.
Common Names in English:
Poorland Flat Sedge, Flat Sedge, Poorland Flatsedge
Common Names in Portuguese:
Junquinho, Tiririca
Description
Family Cyperaceae
Herbs, annual
or perennial
, cespitose or not, rhizomatous
or not, stoloniferous
or not. Roots
fibrous
, principally adventitious. Stems (culms
) usually trigonous
, occasionally terete
, rarely compressed
, usually solid, rarely hollow or septate
. Leaves basal and/or cauline, alternate, usually 3-ranked, rarely 2-ranked or multi-ranked, bases
forming cylindric
sheaths
enclosing stem, margins
usually fused; junction of sheaths and blades
often with adaxial
flaps of tissue
or fringes
of hair (ligules) ; blades frequently absent from some basal leaves
, rarely from cauline leaves, when present divergent or ascending
, flat, folded, plicate
, rolled, or terete, linear
, venation
parallel. Primary
inflorescences (spikelets
) a shortened axis; glumaceous
bracts (scales
) 1-many, spirally arranged
, sometimes 2-ranked, usually appressed
or ascending; scales usually all fertile
, each subtending
a single flower, sometimes proximal
and/or distal scales empty; lateral
spikes often with basal, usually empty, usually 2-keeled scale (prophyll) ; occasionally prophyll subtending and enclosing rachilla, bearing 1 pistillate
, sometimes (0-) 3 staminate flowers
and empty scales (Carex, Cymophyllus, and Kobresia) . Secondary inflorescences panicles, often modified to corymb, pseudoumbel, cyme (anthela), raceme
, spike, or capitulum (head
), rarely single spike, usually subtended by foliaceous
or, less frequently, glumaceous bracts; secondary inflorescences sometimes simulating spikelets (Carex, Cymophyllus, and Kobresia) . Flowers hypogynous, bisexual
in most genera, unisexual
in Scleria, Carex, Cymophyllus, and Kobresia; perianth absent or with (1-) 3-6(-30) bristles
and/or scales, usually falling off with fruit; stamens usually (1-) 3, rarely more, usually distinct
; anthers
basifixed
; pistils 1, 2-3(-4) -carpellate, fused, locule 1; style undivided or branches 2-3(-4) ; stigma sometimes papillate
. Fruits achenes, usually trigonous or biconvex
; pericarps thin (except in Scleria) . Seeds 1; testa thin, free
from pericarp; embryo basal; endosperm abundant. x
= 5-ca. 100.
Genera ca. 100, species ca.
5000 (27 genera, 843 species in the flora
) : worldwide.
No consensus exists regarding the number of genera and the overall relationships
of genera within Cyperaceae. The most recent account of the family
(P. Goetghebeur 1998) recognized 104 genera distributed among 4 subfamilies and 14 tribes
. That arrangement
differs somewhat from that of J. Bruhl (1995) . With one minor exception the arrangement of the family here follows that of Goetghebeur.
The family is characterized by the occurrence of a number of unusual cytological features including: (1) chromosomes with diffuse
centromeres
, (2) post-reductional meiosis, and (3) pollen grains
formed from tetrads
in which 3 of the 4 microspores fail to develop. The first two features are found in at least some Juncaceae and are unique to the two families. Juncaceae also have pollen in tetrads, but in that family all four microspores produce
pollen grains. Some species in some genera of Cyperaceae (particularly Eleocharis) possess chromosomes with localized centromeres (S. S. Bir et al.
1993) . The wide range
of chromosome numbers found in Cyperaceae is largely because of agmatoploidy; polyploidy has been hypothesized for some genera, especially Eleocharis, although polyploidy has not been demonstrated unequivocally.
Because of morphologic similarities in vegetative
and inflorescence characters, the family has commonly been associated with Poaceae. Cytological features discussed above clearly indicate that to be a superficial similarity
. Data from rbcL
studies also support
the view
that Cyperaceae and Poaceae are not closely related (M.
R. Duvall et al. 1993b; G. M. Plunkett et al. 1995) ; they do support the concept of close relationship between Cyperaceae and Juncaceae.
For most families of flowering plants
the phenological data given are flowering times. Because most Cyperaceae cannot be reliably identified when in flower, in this volume fruiting time is given for all species by season
, sometimes qualified by early, mid, or late, or by months. The fruiting time has been interpreted broadly to include the period when the fruit is more or less fully formed but not yet ripe
. The fruiting period provided covers
the entire range of the taxon
. Quite a difference between fruiting periods in different parts of the range of the species may well occur, especially for widespread species and species with extensive elevation
range.
For a recent, comprehensive review of the economic importance of Cyperaceae, see D. A. Simpson and C.
A. Inglis (2001) .[1]
Genus Cyperus
Herbs, perennial
or less often annual
, cespitose or not, rhizomatous
, stoloniferous
, rarely tuberous
. Culms
solitary or not, trigonous
or round
, glabrous
or scabridulous
with extrorse
or antrorse
(rarely retrorse
) prickles. Leaves usually basal; ligules absent; blades
keeled
abaxially, flat, V-, or inversely W-shaped in cross
section
. Inflorescences terminal
, rarely pseudolateral, 1st order
subumbellate to capitate, 2d order with spicate
or digitately arranged spikelets
, rarely a solitary spikelet; spikelets 1-150; 1st order rays unequal (rarely equal) in length
, produced
singly from the axils of inflorescence bracts
; involucral bracts
1-22, spirally arranged
at culm apex, spreading
to erect
, leaflike. Spikelets: scales
to 76, distichous, each subtending
flower, cylindric
to compressed
, borne spicately or digitately at ends of rays (occasionally proliferous) . Flowers bisexual
[rarely unisexual
], in axils of distichous floral scales, bases
often decurrent onto rachilla as ± hyaline
wings
; perianth absent; stamens 1-3; styles linear
, 2-3-fid, base deciduous or persistent
; stigmas 2-3. Achenes biconvex
, flattened, or trigonous.
Species ca.
600: pantemperate and tropical
.[2]
Physical Description
Species Cyperus compressus
Herbs, annual , cespitose with fibrous roots . Culms trigonous , (2-) 6-15(-35) cm × (0.3-) 1-1.8 mm, glabrous . Leaves 1-5, flat to V-shaped, (1-) 5-15(-22) cm × (0.5-) 1.5-3 mm. Inflorescences: spikes ± digitate, ± globose , 7-40 × 10-20 mm; rays (0-) 1-6, 2-12 cm; if absent, inflorescence a sessile cluster of (1-) 3-10 spikelets ; rachis 2-5 mm; bracts (1-) 4-5(-6), ascending , V-shaped, 1-15(-20) cm × 0.5-3.5 mm; rachilla persistent , wingless. Spikelets (1-) 3-12(-16), greenish white, linear to linear-lanceoloid, compressed-quadrangular, 6-34(-40) × 2-3(-3.6) mm; floral scales deciduous, (8-) 16-36(-42), spreading , laterally whitish, greenish, or pale brownish, medially green, weakly 7-9-ribbed, laterally weakly 1-2-ribbed, most ribs medial , ovate , (2.4-) 2.6-3 × (1.9-) 2-2.4(-2.5) mm, apex acute to ± acute, cuspidate tip (0.3-) 0.5-0.7(-0.8) mm. Flowers: anthers 0.6-0.7 mm; styles 0.9-1.3 mm; stigmas 0.5-0.8 mm. Achenes brown, stipitate to sessile, obovoid , (1-) 1.2-1.4(-1.5) × 0.9-1(-1.1) mm, apex obtuse to emarginate , surfaces very finely puncticulate . Fruiting summer-early fall . [source]
Habit: Graminoid • Growth Form: Bunch • Shape and Orientation: Semi-Erect
Flowers: Bloom Period: Late Summer • Flower Color: Green • Flower Conspicuous: No
Seeds: Seed Spread Rate: Moderate • Seedling Vigor: Medium • Fruit/Seed Abundance: Medium • Fruit/Seed Color: Brown • Fruit/Seed Conspicuous: No • Cold Stratification Required: No
Foliage: Foliage Color: Green • Foliage Porosity Summer: Moderate • Foliage Porosity Winter: Porous • Foliage Texture: Fine • Fall Conspicuous: No • Leaf Retention: No
Size/Age/Growth
Active Growth Period: Summer and Fall • Growth Rate: Rapid • After Harvest Regrowth Rate: Slow • Mature Height (feet): 1.0 • Vegetative Spread Rate: None • Lifespan: Lifespan
Habitat
Various disturbed soils; 0-900 m [3].
Biology
Reproduction
Duration: Annual , Perennial • Coppice Potential: No • Progagated by Bulbs: No • Propagated by Bare Root: No • Propagated by Container: No • Propagated by Corms: No • Propagated by Cuttings: No • Propagated by Seed: No • Propagated by Sod: No • Propagated by Sprigs: Yes • Propagated by Tubers: No • Fruit/Seed Period Begin: Fall • Fruit/Seed Period End: Fall • Fruit/Seed Persistence: Yes
Growth
Soil: Adapted to Medium Textured: Adapted to Medium Textured Soils • Adapted to Coarse Textured Soils: Yes • Anaerobic Tolerance: Medium • Salinity Tolerance: None • CaCO3 Tolerance: Medium • Minimum pH: 5.0 • Maximum pH: 6.5 • Fertility Requirement: Low
Sunlight: Sun Exposure: Full Sun . • Shade Tolerance: Intolerant
Moisture: Drought Tolerance: Low • Minimum Precipitation: 30 • Maximum Precipitation: 50 • Moisture Use: Medium
Temperature: Minimum Temperature (F): 37 • Minimum Frost Free Days: 160 • Cold Hardiness: 5a, 5b, 6a, 6b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Commelinidae
(
)
- Takhtajan, 1967
- Superorder:
Juncanae
(
)
- Takhtajan, 1967
- Order:
Poales
(
)
- Small, 1903
- Family:
Cyperaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- Sedge Family
- Subfamily:
Cyperoideae
(
)
- Tribe:
Cypereae
(
)
- Genus:
Cyperus
(
)
- C. Linnaeus, 1753
- Umbrella-sedge, flatsedge [Greek kupeiros, name for Eurasian Cyperus longus Linnaeus]
- Subgenus:
intertidal
(
)
- Specific epithet:
compressus
- L.
- Botanical name: - Cyperus compressus L.
- Specific epithet:
compressus
- L.
- Subgenus:
intertidal
(
- Genus:
Cyperus
(
- Tribe:
Cypereae
(
- Subfamily:
Cyperoideae
(
- Family:
Cyperaceae
(
- Order:
Poales
(
- Superorder:
Juncanae
(
- Subclass:
Commelinidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
C. pectinatus Roxburgh • C. pectiniformis Roemer & Schultes • Cyperus compressus var. pectiniformis (Roemer & Schultes) C. b. Clarke
Notes
Name
Status: Accepted Name
.
Last scrutiny: 21-Jun-2005
Similar Species
Members of the genus Cyperus
ZipcodeZoo has pages for 172 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
C. acuminatus (Short-Point Flatsedge) · C. aggregatus (Inflated Scale Flatsedge) · C. albostriatus (Dwarf Umbrella Grass) · C. alopecuroides (Foxtail Flatsedge) · C. alternifolia (Umbrella Plant) · C. alternifolius var. Gracilis (Dwarf Umbrella Plant) · C. amabilis (Foothill Flatsedge) · C. amuricus (Asian Flatsedge) · C. aromaticus (Navua Sedge) · C. articulatus (Jointed Flatsedge) · C. auriculatus (Eared Flatsedge) · C. bipartitus (Brook Flatsedge) · C. calcicola (Caribbean Flatsedge) · C. cephalanthus (Buttonbush Flatsedge) · C. compressus (Poorland Flat Sedge) · C. confertus (West Indian Flatsedge) · C. congestus (Clustered Flatsedge) · C. croceus (Baldwin's Cyperus) · C. cuspidatus (Coastal Plain Flatsedge) · C. cyperinus (Old World Flatsedge) · C. cyperoides pseudoflavus (Pacific Island Flatsedge) · C. deamii (Deam's Flatsedge) · C. dentatus (Toothed Flatsedge) · C. dentoniae (Hairy Flatsedge) · C. diandrus (Umbrella Flat Sedge) · C. difformis (Smallflower Umbrella Sedge) · C. diffusus (Dwarf Umbrella Grass) · C. digitatus (Finger Flatsegde) · C. dioicus (Flatsedge) · C. dipsaceus (Wright Flatsedge) · C. distans (Piedmont Flatsedge) · C. distinctus (Marshland Flatsedge) · C. echinatus (Globe Flatsedge) · C. elegans (Royal Flatsedge) · C. entrerianus (Deeprooted Sedge) · C. eragrostis (Drain Flat-Sedge) · C. erythrorhizos (Red-Root Flat Sedge) · C. esculentus (Yellow Nutsedge) · C. esculentus L. var. esculentus (Chufa Flatsedge) · C. esculentus var. esculentus (Chufa Flatsedge) · C. esculentus var. hermannii (Yellow Nutsedge) · C. esculentus var. leptostachyus (Yellow Nutsedge) · C. esculentus var. macrostachyus (Yellow Nutsedge) · C. esculentus var. sativus (Chufa) · C. exaltatus (Tall Flat-Sedge) · C. fauriei (Alpine Flatsedge) · C. fendlerianus (Fendler Flatsedge) · C. filicinus (Fern Flatsedge) · C. filiformis (Wiry Flatsedge) · C. flavescens (Pale Flatsedge) · C. flavicomus (White-Edge Flatsedge) · C. flexuosus (Vahl's Flatsedge) · C. floribundus (Rio Grande Sedge) · C. floridanus (Florida Flatsedge) · C. fugax (Withering Flatsedge) · C. fuligineus (Limestone Flatsedge) · C. fuscus (Brown Flatsedge) · C. giganteus (Giant Flatsedge) · C. glaber (Galingale) · C. gracilis (Slimjim Flatsedge) · C. granitophilus (Granite Flatsedge) · C. grayi (Gray's Flatsedge) · C. grayoides (Illinois Flatsedge) · C. haspan (Haspan Flatsedge) · C. hermaphroditus (Hermaphrodite Flatsedge) · C. hillebrandii (Hillebrand's Flatsedge) · C. hillebrandii var. decipiens (Hillebrand's Flatsedge) · C. hillebrandii var. hillebrandii (Hillebrand's Flatsedge) · C. houghtonii (Houghton's Flatsedge) · C. hyalinus (Queensland Sedge) · C. hypochlorus (Oahu Flatsedge) · C. hypochlorus var. brevior (Oahu Flatsedge) · C. hypochlorus var. hypochlorus (Oahu Flatsedge) · C. hypochlorus var. kauaiensis (Oahu Flatsedge) · C. hypopitys (Pinewoods Sedge) · C. hystricinus (Bristly Flatsedge) · C. imbricatus (Shingle Flatsegde) · C. involucratus (Galingale) · C. iria (Iria Flatsedge) · C. isocladus (Dwarf Papyrus) · C. javanicus (Javanese Flatsedge) · C. kunthianus (Maui Flatsedge) · C. kyllinga (Spikesedge) · C. laevigatus (Smooth Flatsedge) · C. lancastriensis (Many-Flowered Umbrella-Sedge) · C. lanceolatus (Epiphytic Flatsedge) · C. lecontei (Le Conte's Flatsedge) · C. lentiginosus (Latin American Flatsedge) · C. ligularis (Alabama Swamp Flatsedge) · C. longus (Sweet Cyperus) · C. louisianensis (Louisiana Flatsedge) · C. lucidus (Leafy Flat-Sedge) · C. lupulinus lupulinus (Great Plains Flatsedge) · C. lupulinus subsp. macilentus (Great Plains Flatsedge) · C. manimae (Smoothstem Flatsedge) · C. manimae var. asperrimus (Spectacular Flatsedte) · C. meyenianus (Meyen's Flatsedge) · C. mutisii (Mutis Flatsedge) · C. nanus (Indian Flatsedge) · C. nanus var. nanus (Indian Flatsedge)
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Further Reading
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Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Data Sources
Accessed through GBIF Data Portal November 14, 2007:
- Australian National Herbarium
- , Australian National Herbarium
- Bernice Pauahi Bishop Museum, Bishop Museum Natural History Specimen Data
- Fairchild Tropical Botanic Garden, Fairchild Tropical Botanic Garden Virtual Herbarium Darwin Core format
- GBIF-Spain, Real Jardin Botanico
- , Vascular Plant Herbarium
- Herbarium of the University of Aarhus, The AAU Herbarium Database
- Herbier de la Guyane, Herbier de la Guyane
- Instituto Nacional de Biodiversidad
- , Biodiversidad de Costa Rica
- Missouri Botanical Garden, Missouri Botanical Garden
- National Herbarium of New South Wales, NSW herbarium collection
- National Herbarium of New South Wales, Plants of Papua New Guinea
- National Institute of Genetics, ROIS, Herbarium Specimens of Museum of Nature and Human Activities, Hyogo Pref., Japan
- Taiwan Biodiversity Information Facility, Magnoliophyta
- The Danish Biodiversity Information Facility, Galapagos grasses and sedges
- The New York Botanical Garden, Species of Eastern Brazil Vascular Plant Specimens
- USDA PLANTS, USDA PLANTS Database
- University of Alabama Biodiversity and Systematics, Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2660432
- Catalogue of Life Accepted Name Code: Kew-236092
- Global Biodiversity Information Facility Taxonkey: 13730640
- Globally Unique Identifier: urn:lsid:ipni.org:names:304158-1
- GRIN Nomen Number: 401630
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 39913
- Natural Heritage Network Species Identifier: PMCYP060E0
- U.S.D.A. Plant Symbol: CYCO
- Zipcode Zoo Species Identifier: 33894
Footnotes
- Peter W. Ball, A. A. Reznicek, David F. Murray "Cyperaceae". in Flora of North America Vol. 23 Page 3, 4, 192, 243, 252. Oxford University Press. Online at EFloras.org. [back]
- Gordon C. Tucker, Brian G. Marcks & J. Richard Carter "Cyperus". in Flora of North America Vol. 23 Page 7, 141, 154, 162, 163, 164, 168, 170, 184,. Oxford University Press. Online at EFloras.org. [back]
- "Cyperus". in Flora of North America Vol. 23 Page 144, 145, 171. Oxford University Press. Online at EFloras.org. [back]
