Interesting Facts
Common Names
Common Names in English:
Dwarf Alpine Hawksbeard, Small Hawksbeard, Tiny Hawksbeard
Description
Family Compositae
The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.
Tribe Lactuceae
The Lactuceae are a tribe of closely related genera of the sunflower family that are easily recognized because the flowering heads are composed of wholly of ligulate florets that are usually 5-lobed. Another very distinguishing feature is the milky sap . Although not apparent without magnification, the pollen is distinctive in that the spines are more or less restricted to discrete ridges or flanges on the surface of the grain. In other members of the family the spines are distributed more or less evenly over the surface of the pollen grain . The pappus usually consists of scales or stiff hairs . -- Gerald D. Carr.
Genus Crepis
Annuals
, biennials, or perennials
, 3-120 cm; usually taprooted, sometimes rhizomatous
(roots
deep or shallow, woody or fibrous
, caudices often woody). Stems 1-20+. erect
to decumbent
. simple
(sometimes scapiform
) or branched, usually striate
, glabrous
or hairy
, often densely hispid
or setose
(hairs
often stipitate-glandular
). Leaves basal (often in rosettes) and cauline; petiolate
(at least basal, petioles
± winged
) ; basal blades mostly elliptic
, ovate
, or lanceolate to linear
, or spatulate
to oblanceolate
, often lyrate or runcinate, margins
entire, dentate
, serrate, toothed
, or pinnately lobed
, lobes
sometimes toothed; cauline usually present, lobed or entire, usually reduced in size and lobing distally. Heads (erect) usually in cymiform, corymbiform
, or paniculiform
arrays, sometimes borne singly. Peduncles not inflated
distally, not bracteate
. Calyculi of 5-12, reduced, subulate
to lanceolate or deltate bractlets
in ± 1 series, mostly unequal, glabrous, tomentulose
, or setose. Involucres cylindric
to campanulate
(sometimes becoming turbinate
in fruit), 4-15 mm diam. Phyllaries 5-18 in 1-2 series. lanceolate, equal or subequal
, (bases
becoming thickened and keeled
, keels sometimes pronounced in fruit) margins green to yellowish, often scarious
, apices acute to acuminate, abaxial
faces
glabrous, tomentose
, or setose, sometimes stipitate-glandular, adaxial
glabrous or with appressed
hairs. Receptacles flat or convex
, usually pitted
, glabrous or hairy, epaleate [paleate, paleae narrow, thin]. Florets 5-100+; corollas usually yellow or orange, sometimes white, pink, or reddish. Cypselae monomorphic
or dimorphic
. yellow, brown, green, red, and/or black, subcylindric
or fusiform
, terete
or subterete, usually curved
, apices tapered or beaked
, ribs
10-20, sometimes spiculate-roughened, faces glabrous or hispidulous
; pappi persistent
or falling, of 80-150, usually distinct
, sometimes basally connate
, white to tawny
, coarse
to fine, ± equal (or outer shorter), barbellulate
bristles
in 1-2 series. x = 3, 4, 5, 6, 11.
Species ca.
200: North America, Eurasia
, Africa; introduced
nearly worldwide.
Crepis is generally recognized by the rosettes of coarse, often pinnately lobed leaves, erect heads, epaleate receptacles, calyculate involucres. yellow corollas, subcylindric or fusiform, ribbed
cypselae, and pappi of barbellulate bristles. The taxonomy and evolutionary relationships
of Crepis were studied by E. B
. Babcock (1947) and his associates. Their work was thorough and important because of the effort
to incorporate cytogenetic information in the evolutionary analysis. Extensive survey of chromosome number and karyotype
indicated two major ploidy groups in Crepis, corresponding to New World and Old World species complexes. Of the 12 species of Crepis native
to North America, 10 are polyploids
with x = 11. The core
diploid populations commonly occupy discrete ecologic zones and are thought to be entirely distinct from one another, yet they are interconnected by a continuous complex
series of intergrading polyploid forms that are partly or completely apomictic (Babcock). The polyploids are of two forms, autopolyploids that are similar to the diploids, and allopolyploids that combine the characteristics of two or more diploid species. The allopolyploid forms of hybrid origin
may exhibit
the characteristics of multiple
parental species and therefore are difficult to classify. Some of the heterogeneous
apomictic populations, or groups of populations, have been grouped together and recognized as subspecies
; those taxa are often difficult to identify and further study is clearly needed. Despite these difficulties, the subspecific taxa of Babcock were tentatively included
in the present study. The Old World species are mostly diploid (n = 3, 4, 5, or 6). Babcock concluded that there was a progressive decrease in the chromosome numbers, from n = 6 to n = 3. Along with the decrease is a corresponding increase in chromosome asymmetry and reduction in chromosome length.[1]
Physical Description
Species Crepis nana
Perennials
, 10-20 cm (taproots
often with creeping
rhizomes,
caudices relatively short). Stems 1-10+, erect
or ascending
(in dense clumps
), simple
or proximally branched, glabrous
. Leaves
basal and cauline; petiolate
(at least basal) ; blades
(often purplish),
orbiculate to spatulate
, less often lyrate or runcinate, 2-9 ×
0.5-2.5 cm, (bases
abruptly 0) margins
entire or pinnately lobed
,
apices obtuse
to acute, faces
glabrous (glaucous). Heads 5-80+
(among or beyond leaves), in cymiform arrays. Calyculi of
5-10 (dark green or blackish), lanceolate, glabrous bractlets 2-3
mm.
Involucres cylindric
, 8-13 × 3-4 mm. Phyllaries
8-10, (dark green or purple medially) oblong
, 10-11 mm, (margins
scarious
, eciliate
) apices acute, faces glabrous. Florets
9-12; corollas yellow, purple-tinged abaxially, 9-12 mm. Cypselae
golden brown, subcylindric
to fusiform
, 4-7 mm, apices sometimes
tapered (not beaked
), ribs
10-13 (broad, smooth
) ; pappi (falling)
bright white, 4-6 mm. 2n = 14. [source]
Crepis nana occurs in North America and northern Asia. It
is recognized by the tufted
, cespitose habit, elongate
roots
and
rhizomes, and occurrence in alpine
habitats
. In the typical form,
the plants
are tufted, the stems are not leafy, and the heads
are
borne among the leaves. Taller specimens with elongated, leafy branches
and heads borne well beyond the basal leaves
are sometimes recognized
as subsp.
ramosa; these characteristics appear to be part
of the normal range
of variation
for the species. [source]
Crepis nana is closely related to C.
elegans, differing
mainly in the shape
of the cypselae. The cypselae of C. nana
are almost always more columnar
, wider at bases, and with broader
ribs, than those of C. elegans. [source]
The name
Crepis nana subsp. clivicola Legge is invalid
.
[source]
Habit: Forb/herb
Flowers: Bloom Period: May, June, July, August, September. • Flower Color: yellow
Size/Age/Growth
Size: under 6" tall.
Habitat
Talus slopes , rocky alpine places, sandy stream banks, gravel bars , exposed sites in shrub communities; 300-4000 m [2].
Typically found at an altitude of 0 to 3,925 meters (0 to 12,877 feet).[3]
Biology
Reproduction
Duration: Perennial
Growth
Sunlight: Sun Exposure: Full sun .
Temperature: Cold Hardiness: 1, 2a, 2b, 3a, 3b, 4a, 4b, 5a, 5b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Campanulanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Asterales
(
)
- Lindley, 1833
- Family:
Compositae
(
)
- Giseke, 1792, nom. cons., nom. alt.
- Subfamily:
Cichorioideae
(
)
- Tribe:
Lactuceae
(
)
- Subtribe:
Crepidinae
(
)
- Genus:
Crepis
(
)
- C. Linnaeus, 1753
- Hawksbeard [Greek krepis, slipper or sandal. possibly alluding to shape of cypselae, a name of a plant in writings by Theophrastus]
- Specific epithet:
nana
- Richardson
- Botanical name: - Crepis nana Richardson
- Specific epithet:
nana
- Richardson
- Genus:
Crepis
(
- Subtribe:
Crepidinae
(
- Tribe:
Lactuceae
(
- Subfamily:
Cichorioideae
(
- Family:
Compositae
(
- Order:
Asterales
(
- Superorder:
Campanulanae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
C. nana ramosa Babcock • Crepis nana var. lyratifolia (Turczaninow) Hultén
Notes
Name
Status: Accepted Name
.
Comment: Data Providers: Govaerts World Compositae Checklist
A-G,
IPNI, Tropicos, LCR Editor. GCC LSID: urn
:lsid:compositae.org:names:B046E0DA-5073-42BE-8F4B-EFA65FC2EB0F
Last scrutiny: 02-Nov-09
Similar Species
Members of the genus Crepis
ZipcodeZoo has pages for 44 species, subspecies, varieties, forms, and cultivars in this genus:
C. acuminata (Long-Leaf Hawksbeard) · C. acuminata acuminata (Longleaf Hawksbeard) · C. acuminata pluriflora (Longleaf Hawksbeard) · C. atribarba (Hawksbeard) · C. atribarba atribarba (Hawksbeard) · C. aurea (Golden Hawks Beard) · C. bakeri (Baker's Hawksbeard) · C. bakeri cusickii (Baker's Hawksbeard) · C. bakeri idahoensis (Baker's Hawksbeard) · C. biennis (Rough Hawksbeard) · C. bursifolia (Italian Hawksbeard) · C. capillaris (Smooth Hawk's-Beard) · C. elegans (Elegant Hawksbeard) · C. foetida (Roadside Hawksbeard) · C. foetida foetida (Stinking Hawksbeard) · C. intermedia (Gray Hawksbeard) · C. modocensis rostrata (Common Hawksbeard) · C. monticola (Mountain Hawksbeard) · C. nana (Dwarf Alpine Hawksbeard) · C. nana ramosa (Dwarf Alpine Hawksbeard) · C. nicaeensis (Turkish Hawksbeard) · C. occidentalis conjuncta (Gray Hawksbeard) · C. occidentalis pumila (Large-Flower Hawk's-Beard) · C. pannonica (Pasture Hawksbeard) · C. pleurocarpa (Naked-Stem Hawksbeard) · C. pulchra (Hawksbeard) · C. pulchra pulchra (Smallflower Hawksbeard) · C. pumila (Largeflower Hawksbeard) · C. rostrata (Modoc Hawksbeard) · C. rubra (Hawksbeard) · C. rubra 'Alba' (Hawks Beard) · C. runcinata andersonii (Dandelion Hawksbeard) · C. runcinata barberi (Barber's Hawksbeard) · C. runcinata hallii (Fiddleleaf Hawksbeard) · C. runcinata hispidulosa (Fiddleleaf Hawksbeard) · C. runcinata imbricata (Fiddleleaf Hawksbeard) · C. runcinata runcinata (Fiddle-Leaf Hawk's-Beard) · C. setosa (Bristly Hawksbeard) · C. tectorum (Narrow-Leaf Hawk's-Beard) · C. tectorum tectorum (Narrowleaved Hawk's Beard) · C. vesicaria (Beaked Hawksbeard) · C. vesicaria haenseleri (Beaked Hawksbeard) · C. vesicaria taraxacifolia (Beaked Hawksbeard) · C. zacintha (Striped Hawksbeard)
More Info
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Further Reading
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- A report upon the boreal flora of the Sierra Nevada of California / by Frank Jason Smiley. Berkeley, Calif.: University of California Press, 1921. url p. 405, p. 48.
- Alpine flora of the Canadian Rocky Mountains. Illustrated with water-colour drawings and photographs by Mrs. Charles Schäffer. New York, London, G. P. Putnam's sons, 1907. url p. 280, p. 280.
- Annual report. United States Geological and Geographical Survey of the Territories. 1st-12th, 1867-1878. Washington, Govt. Print. Off., 1867-83. url p. 757, p. 772.
- Appendix [to] Captain Parry's journal [of a] second voyage [for] the discovery of a North West passage from the Atlantic to the Pacific [performed in] His Majesty's ships Fury and Hecla, [in] the years 1821-22-23. London: J. Murray, 1825. url .
- Appendix to Captain Parry's journal of a second voyage for the discovery of a north-west passage from the Atlantic to the Pacific, performed in His Majesty's ships Fury and Hecla in the years 1821-22-23. London: J. Murray, 1825. url .
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- Budd's flora of the Canadian Prairie Provinces / [Ottawa]: Research Branch, Agriculture Canada, 1987. url p. 691, p. 692.
- Bulletin of the Southern California Academy of Sciences. Los Angeles, Calif.: The Academy, 1902-1971. url , , p. 13.
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- Cyclopedia of American horticulture, comprising suggestions for cultivation of horticultural plants, descriptions of the species of fruits, vegetables, flowers and ornamental plants sold in the United States and Canada, together with geo by L.H. Bailey assisted by Wilhelm Miller, PH. D., associate editor, and many expert cultivators and botanists. Illustrated with nearly three thousand engravings and one hundred and forty-five full-pa New York, Doubleday, Page & Company, 1906. url p. 1539.
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- Flora of southern British Columbia and Vancouver Island: with many references to Alaska and northern species / by Joseph Kaye Henry. Toronto: W.J. Gage, c1915. url p. 329.
- Flora of the Rocky Mountains and adjacent plains, Colorado, Utah, Wyoming, Idaho, Montana, Saskatchewan, Alberta, and neighboring parts of Nebraska, South Dakota, North Dakota, and British Columbia / by P.A. Rydberg. New York: The author, 1922. url p. 1021, p. 1069.
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- Flora of the northwest coast, including the area west of the summit of the Cascade Mountains, from the forty-ninth parallel south to the Calapooia Mountains on the south border of Lane County, Oregon. Lancaster, Pa., Press of the New Era Printing Company, 1915. url .
- Flora of the northwest coast: including the area west of the summit of the Cascade Mountains, from the forty-ninth parallel south to the Calapooia Mountains on the south border of Lane County, Oregon / by Charles V. Piper and R. Kent Beattie. Lancaster, Pa.: Press of the New era printing company, 1915. url p. 358.
- Flora of the state of Washington / by Charles V. Piper. Washington: G.P.O., 1906 url p. 545.
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- Historical view of the progress of discovery on the more northern coasts of America, from the earliest period to the present time. New York, Harper & brothers, 1839. url p. 311.
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- Science studies, v.1, no. 1-3. Bozeman, Mont.1905. url p. 105, p. 34.
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Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Data Sources
Accessed through GBIF Data Portal February 16, 2008:
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- Canadian Museum of Nature, Canadian Museum of Nature Herbarium
- USDA PLANTS, USDA PLANTS Database
- University of Alaska Museum of the North, University of Alaska Museum of the North Herbarium
- University of Colorado Museum, Zoological specimens
- Utah State University, USU-UTC Specimen Database
- Utah Valley State College
- , Utah Valley State College Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2658424
- Catalogue of Life Accepted Name Code: Ast-1823
- Global Biodiversity Information Facility Taxonkey: 13749460
- Globally Unique Identifier: urn:lsid:ipni.org:names:200085-1
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 37191
- Natural Heritage Network Species Identifier: PDAST2R0C0
- U.S.D.A. Plant Symbol: CRNAL CRNA
- Zipcode Zoo Species Identifier: 33019
Footnotes
- David J. Bogler "Crepis". in Flora of North America Vol. 19, 20 and 21 Page 214, 216, 217, 219, 222, 223, 228. Oxford University Press. Online at EFloras.org. [back]
- "Crepis nana". in Flora of North America Vol. 19, 20 and 21 Page 224, 228, 231. Oxford University Press. Online at EFloras.org. [back]
- Mean = 1,226.150 meters (4,022.802 feet), Standard Deviation = 858.160 based on 147 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
