Overview
Interesting Facts
Common Names
Common Names in English:
Black Haw, Black Hawthorn, Douglas Hawthorn, Douglas Thorntree, Douglas's Hawthorn, River Hawthorn, Western Thornapple
Description
Family Rosaceae
Trees
, shrubs
, or herbs, deciduous or evergreen
. Stems erect
, scandent
, arching
, prostrate
, or creeping
, armed
or unarmed
. Buds usually with several exposed scales
, sometimes with only 2. Leaves alternate, rarely opposite, simple
or compound
; stipules paired
, free
or adnate
to petiole
, rarely absent, persistent
or deciduous; petiole usually 2-glandular apically; leaf blade
often serrate at margin
, rarely entire. Inflorescences various, from single flowers to umbellate
, corymbose
, racemose or cymose-paniculate. Flowers usually actinomorphic
, bisexual
, rarely unisexual
and then plants
dioecious. Hypanthium (formed from basal parts of sepals, petals, and stamens) free from or adnate to ovary, short or elongate
. Sepals usually 5, rarely fewer or more, imbricate; epicalyx
segments sometimes also present. Petals as many as sepals, inserted
below margin of disk, free, imbricate, sometimes absent. Disk lining hypanthium, usually entire, rarely lobed
. Stamens usually numerous
, rarely few, always in a complete
ring
at margin of or above disk; filaments
usually free, very rarely connate
; anthers
small, didymous
, rarely elongate, 2-locular. Carpels 1 to many, free, or ± connate and then adnate to inner surface of cupular receptacle; ovary inferior, semi-inferior, or superior; ovules usually 2 in each carpel, rarely 1 or several, anatropous
, superposed
. Styles as many as carpels, terminal
, lateral
, or basal, free or sometimes connate. Fruit a follicle, pome, achene, or drupe, rarely a capsule, naked or enclosed in persistent hypanthium and sometimes also by sepals. Seeds erect or pendulous, sometimes winged
, usually exalbuminous
, very rarely with thin endosperm; cotyledons mostly fleshy
and convex
abaxially, rarely folded or convolute.
Between 95 and 125 genera and 2825-3500 species: cosmopolitan
, mostly in N temperate
zone; 55 genera (two endemic) and 950 species (546 endemic) in China.
Many plants of this family
are of economic importance and contribute to people s livelihoods. The Rosaceae contain a great number of fruit trees of temperate regions
. The fruits contain vitamins, acids, and sugars
and can be used both raw and for making preserves, jam, jelly, candy, various drinks, wine, vinegar, etc.
The dried fruits of the genera
Amygdalus and Armeniaca are of high commercial
value. Some plants in the genus Rosa containing essential oils or with a high vitamin content are used in industry
. Rosaceae wood is used for making various articles, stems and roots
are used for making tannin extract, and young leaves are used as a substitute for tea. Numerous species are used for medical purposes or are cultivated as ornamentals
.
The Rosaceae are very well represented in China, with great economic and scientific importance. The Co-chairs of the Editorial Committee (Wu and Raven) here note
that the patterns
of relationship
are complex
and the group is taxonomically difficult. [1]
Genus Crataegus
Shrubs
, subshrubs
, or small trees
, deciduous, rarely evergreen
, armed
, rarely unarmed
; buds ovoid
or subglobose. Leaves simple
, stipulate
, venation
craspedodromous
, margin
serrate and lobed
or partite, rarely entire. Inflorescences corymbose
, sometimes flowers solitary. Hypanthium campanulate
. Sepals 5. Petals 5, white, rarely pinkish. Stamens 5-25; carpels 1-5, connate
, but free
apically. Ovary inferior or semi-inferior, with 2 ovules per locule, but one rudimentary
. Fruit a pome, with persistent
sepals at apex; carpels bony when mature
, each locule with 1 seed; seed erect, cotyledons plano-convex
.
At least 1000 species: N temperate regions
, especially abundant in North America; 18 species (ten endemic) in China.[2]
Physical Description
Habit: Tree , Shrub • Growth Form: Thicket Forming • Shape and Orientation: Erect
Flowers: Bloom Period: Mid Spring • Flower Color: White • Flower Conspicuous: Yes
Seeds: Seed per Pound: 20033 • Seed Spread Rate: Moderate • Seedling Vigor: High • Fruit/Seed Abundance: Medium • Fruit/Seed Color: Black • Fruit/Seed Conspicuous: Yes • Cold Stratification Required: Yes
Foliage: Foliage Color: Green • Foliage Porosity Summer: Dense • Foliage Porosity Winter: Porous • Foliage Texture: Medium • Fall Conspicuous: Yes • Leaf Retention: No
Size/Age/Growth
Active Growth Period: Spring , Summer, Fall • Growth Rate: Moderate • Mature Height (feet): 25.0 • Maximum Height at 20 Years (feet): 15 • Size: 6-8' tall. • Vegetative Spread Rate: Moderate • Lifespan: Lifespan
Habitat
Typically found at an altitude of 0 to 2,450 meters (0 to 8,038 feet).[3]
Biology
Reproduction
Duration: Perennial • Coppice Potential: Yes • Progagated by Bulbs: No • Propagated by Bare Root: Yes • Propagated by Container: Yes • Propagated by Corms: No • Propagated by Cuttings: Yes • Propagated by Seed: Yes • Propagated by Sod: No • Propagated by Sprigs: No • Propagated by Tubers: No • Fruit/Seed Period Begin: Spring • Fruit/Seed Period End: Fall • Fruit/Seed Persistence: Yes
Growth
Culture: Space 20-30' apart.
Soil: Adapted to Medium Textured: Adapted to Medium Textured Soils • Adapted to Coarse Textured Soils: Yes • Anaerobic Tolerance: Medium • Salinity Tolerance: None • CaCO3 Tolerance: Low • Minimum pH: 6.5 • Maximum pH: 7.5 • Fertility Requirement: Low
Sunlight: Sun Exposure: Sun to Partial Shade. • Shade Tolerance: Intermediate
Moisture: Drought Tolerance: Low • Minimum Precipitation: 16 • Maximum Precipitation: 260 • Moisture Use: High
Temperature: Minimum Temperature (F): -29 • Minimum Frost Free Days: 120 • Cold Hardiness: 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b, 9a, 9b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Rosanae
(
)
- Takhtajan, 1967
- Order:
Rosales
(
)
- Perleb, 1826
- Family:
Rosaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- Rose Family
- Tribe:
Crataegeae
(
)
- Genus:
Crataegus
(
)
- C. Linnaeus, 1753
- Hawthorn
- Specific epithet:
douglasii
- Macoun
- Botanical name: - Crataegus douglasii Macoun
- Specific epithet:
douglasii
- Macoun
- Genus:
Crataegus
(
- Tribe:
Crataegeae
(
- Family:
Rosaceae
(
- Order:
Rosales
(
- Superorder:
Rosanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Anthomeles douglasii (Lindl.) M. Roem. • Crataegus punctata var. brevispina Douglas Ex Hook. • Crataegus sanguinea Nutt. • Crataegus sanguinea var. douglasii (Lindl.) Torr. & A. Gray • Mespilus douglasii (Lindl.) Asch. & Graebn. • Mespilus sanguinea var. douglasii (Lindl.) Wenz.
Notes
Name Status: Accepted Name .
Similar Species
Members of the genus Crataegus
ZipcodeZoo has pages for 243 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
C. aemula (Rome Hawthorn) · C. aestivalis (Apple Hawthorn) · C. aestivalis var. cerasoides (Eastern Mayhaw) · C. aestivalis var. dormonae (Eastern Mayhaw) · C. aestivalis var. maloides (Eastern Mayhaw) · C. ambigua (Russian Hawthorn) · C. ambitiosa (Grand Rapids Hawthorn) · C. anamesa (Fort Bend Hawthorn) · C. ancisa (Mississippi Hawthorn) · C. annosa (Phoenix City Hawthorn) · C. anomala (Anomalous Hawthorn) · C. apiomorpha (Fort Sheridan Hawthorn) · C. arborea (Montgomery Hawthorn) · C. arcana (Carolina Hawthorn) · C. arnoldiana (Arnold Hawthorn) · C. arrogans (Dixie Hawthorn) · C. ater (Nashville Hawthorn) · C. austromontana (Valley Head Hawthorn) · C. azarolus (Azarole) · C. azarolus var. pontica (Mediterranean-Medlar) · C. beadlei (Beadle's Hawthorn) · C. beata (Dunbar's Hawthorn) · C. berberifolia (Barberry Hawthorn) · C. bona (Berks County Hawthorn) · C. brachyacantha (Blue Haw) · C. brainerdii (Brainerd Hawthorn) · C. brazoria (Brazos Hawthorn) · C. brevipes (Hawthorn) · C. calpodendron (Pear Hawthorn) · C. canadensis (Canadian Hawthorn) · C. carrollensis (Eureka Springs Hawthorn) · C. coccinea (Scarlet Hawthorn) · C. coccinioides (Kansas Haswthorn) · C. coccinioides var. corallicola (Kansas Haswthorn) · C. coccinioides var. pottsii (Kansas Haswthorn) · C. coleae (Cole's Hawthorn) · C. columbiana (Columbian Hawthorn) · C. compacta (Clustered Hawthorn) · C. compta (Adorned Hawthorn) · C. condigna (River Junction Hawthorn) · C. consanguinea (Tallahassee Hawthorn) · C. contrita (Southern Hawthorn) · C. corusca (Shiningbranch Hawthorn) · C. crus-galli (Cockspur Hawthorn) · C. crus-galli var. inermis (Thornless Cockspur Hawthorn) · C. crusgalli (Clarkton Hawthorn) · C. cuneata (Nippon Hawthorn) · C. dallasiana (Dallas Hawthorn) · C. densiflora (Denseflower Hawthorn) · C. desueta (New York Hawthorn) · C. dilatata (A Hawthorn) · C. dispar (Aiken Hawthorn) · C. disperma (Spreading Hawthorn) · C. dispessa (Mink Hawthorn) · C. dissona (Northern Hawthorn) · C. distincta (Distinct Hawthorn) · C. dodgei (Dodge's Hawthorn) · C. douglasii (Black Haw) · C. douglasii var. douglasii (Black Hawthorn) · C. douglasii var. duchesnensis (Duchesne Black Hawthorn) · C. engelmannii (Engelmann's Hawthorn) · C. erythrocarpa (Red Hawthorn) · C. erythropoda (Cerro Hawthorn) · C. exilis (Slender Hawthorn) · C. extraria (Marietta Hawthorn) · C. flabellata (Fanleaf Hawthorn) · C. flava (Summer Haw) · C. flava var. integra (Yellowfruit-Thorn) · C. flava 'Upright' (Upright Yellow Hawthorne) · C. fragilis (Fragile Hawthorn) · C. fulleriana (Fuller's Hawthorn) · C. furtiva (Albany Hawthorn) · C. glareosa (Port Huron Hawthorn) · C. grandis (Grand Hawthorn) · C. greggiana (Gregg Hawthorn) · C. haemacarpa (Hawthorn) · C. harbisonii (Harbison Hawthorn) · C. harveyana (Harvey's Hawthorn) · C. holmesiana (Holmes' Hawthorn) · C. hudsonica (Hudson Hawthorn) · C. ideae (Concord Hawthorn) · C. ignave (Bedford Springs Hawthorn) · C. immanis (Hawthorn) · C. impar (Redclay Hawthorn) · C. inanis (Oldmaid Hawthorn) · C. incaedua (Hawthorn) · C. indicens (Mansfield Hawthorn) · C. insidiosa (Ozark Hawthorn) · C. integra (Lake Ella Hawthorn) · C. intricata (Biltmore Hawthorn) · C. invicta (Fulton Hawthorn) · C. iracunda (Stolonbearing Hawthorn) · C. irrasa (Blanchard Hawthorn) · C. jesupii (Jesup's Hawthorn) · C. jonesiae (Miss Jones Hawthorn) · C. kelloggii (Kellogg Hawthorn) · C. kennedyi (Kennedy's Hawthorn) · C. kingstonensis (Kingston's Hawthorn) · C. knieskerniana (Knieskern's Hawthorn) · C. laciniata (Oriental Hawthorn)
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Further Reading
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- A preliminary classification of the plant communities of northeastern Montana / Robert L. DeVelice [et al.]. Helena, MT: Montana Natural Heritage Program, c1991. url .
- American forest trees, by Henry H. Gibson; ed. by Hu Maxwell. Chicago, Hardwood record, 1913. url , p. 12.
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- Circle West vegetation monitoring study. Helena, Mont.: Energy Division, Montana Dept. of Natural Resources and Conservation, -[1982] url p. 29, p. 37.
- Conservation strategy for Silene spaldingii (Spalding's catchfly) in Montana / Helena, Mont.: Montana Natural Heritage Program, 2005. url , .
- Contributions from the United States National Herbarium 25 1925 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 284, p. 347, p. 366.
- Ecological inventory of wetland sites in the Thompson-Fisher conservation easement / Helena, MT: Montana Natural Heritage Program, c2002. url , .
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- Flora of the northwest coast: including the area west of the summit of the Cascade Mountains, from the forty-ninth parallel south to the Calapooia Mountains on the south border of Lane County, Oregon / by Charles V. Piper and R. Kent Beattie. Lancaster, Pa.: Press of the New era printing company, 1915. url p. 200.
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- Forest trails and highways of the Mount Hood region. Wash., 1920 url p. 31.
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- Yü Te-tsun, Lu Ling-ti, Ku Tsue-chih, Li Chao-luan, Kuan Ke-chien & Chiang Wan-fu. 1974, 1985, 1986. Rosaceae. In: Yü Te-tsun, ed., Fl. Reipubl. Popularis Sin. 36: 1443; 37: 1516; 38: 1133.
- Yü Te-tsun, Lu Ling-ti, Ku Tsue-chih, Li Chao-luan, Kuan Ke-chien & Chiang Wan-fu. 1974, 1985, 1986. Rosaceae. In: Yü Te-tsun, ed., Fl. Reipubl. Popularis Sin. 36: 1443; 37: 1516; 38: 1133.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
- Global Biodiversity Information Facility. Accessed September 15, 2007. http://www.gbif.org Mediated distribution data from provider.
- IOPI Global Plant Checklist 2005.
- IOPI Global Plant Checklist. Release date: August 1, 2007
- IOPI-GPC
- Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
- The International Plant Names Index. Accessed Jan 19, 2007.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 27, 2008)
Data Sources
Accessed through GBIF Data Portal November 19, 2007:
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- Canadian Museum of Nature, Canadian Museum of Nature Herbarium
- Missouri Botanical Garden, Missouri Botanical Garden
- Oregon State University, Vascular Plant Collection
- USDA PLANTS, USDA PLANTS Database
- University of Washington Burke Museum, Vascular Plant Collection - University of Washington Herbarium
- Utah State University, USU-UTC Specimen Database
- Utah Valley State College
- , Utah Valley State College Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2649831
- Catalogue of Life Accepted Name Code: Ros-5448
- Global Biodiversity Information Facility Taxonkey: 13688706
- Globally Unique Identifier: urn:lsid:ipni.org:names:723298-1
- GRIN Nomen Number: 12096
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 24557
- International Plant Names Index (IPNI) ID: 68215-2
- Natural Heritage Network Species Identifier: PDROS0H860
- U.S.D.A. Plant Symbol: CRDO2
- Zipcode Zoo Species Identifier: 32912
Footnotes
- Cuizhi Gu, Chaoluan Li, Lingdi Lu, Shunyuan Jiang, Crinan Alexander, Bruce Bartholomew, Anthony R. Brach, David E. Boufford, Hiroshi Ikeda, Hideaki Ohba, Kenneth R. Robertson & Steven A. Spongberg "Rosaceae". in Flora of China Vol. 9 Page 46. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Ku Tsue-chih, Stephen A. Spongberg "Crataegus". in Flora of China Vol. 9 Page 111. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = 969.820 meters (3,181.824 feet), Standard Deviation = 582.150 based on 279 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
