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Coldenia grandiflora

Interesting Facts

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Description

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Family Boraginaceae

Herbs perennial , biennial, or annual , less often lianas, shrubs , or trees , usually bristly or scabrous-pubescent. Leaves simple , exstipulate , alternate, rarely opposite, entire or serrate at margin . Inflorescences often double scorpioid cymes, rarely solitary; bracts present or absent. Flowers bisexual , actinomorphic , rarely zygomorphic. Calyx usually 5-parted or lobed , mostly persistent . Corolla tubular , campanulate , rotate, funnelform , or salverform ; tube appendages 5, rarely more, mostly trapeziform, rarely absent, sometimes a ring of hairs present; limb usually 5-parted; lobes overlapping, rarely twisted in bud. Stamens 5, inserted on corolla tube or rarely at throat , included or rarely exserted; anthers introrse , 2-loculed, usually dorsifixed at base , less often medifixed , dehiscence longitudinal . Nectaries at base of corolla tube or on disc below ovary. Ovary superior, 2-carpellate; locules 2 and each with 2 ovules, or 4 and each with 1 ovule; ovules nearly atropous , semianatropous, or anatropous . Style terminal or gynobasic , branched or not. Gynobase flat, fastigiate , or subulate . Fruit 1-4-seeded drupes or nutlets (mericarps) ; nutlets mostly dry, often ornamented with wings , prickles and/or glochids (stiff bristles with barbed or anchorlike tips ) . Seeds vertical or oblique , coat membranous; embryo straight, less often curved ; cotyledons flat, fleshy .

About 156 genera and 2500 species: temperate and tropical regions , centered in the Mediterranean region; 47 genera and 294 species in China, of which four genera and 156 species are endemic.[1]

Genus Coldenia

Herbs annual . Leaves alternate, margin lobed ; secondary veins ending in sinuses. Inflorescences generally extra-axillary on leafy twigs or in forks of branches, sometimes glomerate . Flowers sessile or on short pedicels, 4-merous. Calyx 4-lobed. Corolla throat glabrous , naked or with 4 scaly appendages ; lobes spreading , overlapping in bud. Ovary ovoid , somewhat 4-lobed, 2-loculed and with 2 ovules per locule, or falsely 4-loculed and with 1 ovule per locule; ovules anatropous . Styles 2, terminal , divided or united to middle ; stigmas 2, usually divided. Fruit fleshy or dry, fastigiate or semiglobose, endocarp bony, divided into 4 1-seeded achenelike mericarps; mericarps ± united ventrally or joined to a central extension of receptacle. Seeds with little or no endosperm or not; embryo straight or curved .

For a discussion on the generic limits of Coldenia, see Richardson (Rhodora 79: 476. 1977).

One species: Cambodia, China, India, Indonesia, Malaysia, Pakistan, Sri Lanka, Thailand, Vietnam; Africa, Australia, North America, South America.[2]

Taxonomy

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Notes

Publishing author : Phil. Publication : in Anal . Mus. nac. Chile, (1892) 55.

Similar Species

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Members of the genus Coldenia

ZipcodeZoo has pages for 0 species, subspecies, varieties, forms, and cultivars in this genus:

More Info

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Further Reading

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Notes

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Contributors

Identifiers

Footnotes

  1. Gelin Zhu, Harald Riedl & Rudolf V. Kamelin "Boraginaceae". in Flora of China Vol. 16 Page 329. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
  2. "Coldenia". in Flora of China Vol. 16 Page 337. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
Last Revised: 7/20/2012