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Chamaedaphne calyculata

(Dwarf Cassandra)

Common Names

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Common Names in English:

Cassandra, Dwarf Cassandra, Leatherleaf

Description

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Family Ericaceae

Plants usually woody, or herbs, sometimes lacking chlorophyll. Leaves spiral , sometimes decussate, margin often toothed . Inflorescence racemose; bracteoles paired , basal. Flowers (4 or) 5-merous. Calyx imbricate. Corolla connate , imbricate. Stamens 10, sometimes with spurs or awns , dehiscing by pores , pollen in tetrads , rarely single. Ovary superior or inferior, placentation axile , rarely parietal , often many ovules per locule. Style ± as long as corolla, slender. Fruit a capsule or berry, rarely a drupe; calyx persistent.

About 125 genera and 4000 species: widely distributed in temperate and subarctic regions, also at high elevations in tropical regions ; 22 genera and 826 species (524 endemic) in China.

The Monotropoideae are here included in the Ericaceae; previously in FRPS (56: 157 216. 1990), they were treated as the Pyrolaceae. Chiogenes, recorded from China in FRPS (57(3) : 69 71. 1991), is here included in Gaultheria. Over the last half century, the Empetraceae have usually been separated from, but closely associated with, the Ericaceae. In their ecology, leaf morphology and insertion , rusts, embryology, stamen anatomy, etc. , they largely agree with that family . Molecular data place Empetrum and its relatives firmly within the Ericaceae, and in particular within the subfamily Ericoideae, in agreement with phytochemical and palynological data, and there they are best recognized as a separate tribe . There are distinctive features of the Empetreae that were responsible for their past familial status, e.g. , reduced perianth with separate members , low ovule number, enlarged stigmas, etc. However, these are likely to be derived features associated with wind pollination. See Kron et al. (Bot. Rev. 68: 335 423. 2002) and the recent treatment of the Ericaceae by Stevens et al. (in Kubitzki, Fam. Gen. Vasc. Pl. 6: 145 194. 2004) .

Several genera and many species are ornamentals . Some fruits of Vaccinium in N China are sweet and edible, but of no particular value to humans. Some species of Chamaedaphne, Craibiodendron, Leucothoë, Lyonia, Pieris, and Rhododendron contain more or less toxic diterpenes, which are harmful to humans or domestic animals.[1]

Genus Chamaedaphne

Shrubs evergreen , lepidote . Stems arching . Leaf blade leathery, margin entire or inconspicuously denticulate . Inflorescences terminal , secund racemes ; bracts leaflike. Flowers 5-merous. Corolla urceolate to tubular . Stamens included ; filaments straight, subulate , lacking appendages , anthers dehiscing by terminal pores. Ovary superior, with many ovules per locule; stigma truncate . Capsule loculicidal; epicarp 5-valved, separating from 10-valved endocarp. Seeds many, small, wingless.

One species: China, N Japan, Mongolia, Russia (Siberia) ; NE Europe, North America.[2]

Physical Description

Habit: ShrubGrowth Form: RhizomatousShape and Orientation: Irregular

Flowers: Bloom Period: Early SpringFlower Color: White • Flower Conspicuous: No

Seeds: Seed per Pound: 96000 • Seed Spread Rate: Slow • Seedling Vigor: Medium • Fruit/Seed Abundance: Low • Fruit/Seed Color: Brown • Fruit/Seed Conspicuous: No • Cold Stratification Required: No

Foliage: Foliage Color: Yellow-Green • Foliage Porosity Summer: Porous • Foliage Porosity Winter: Porous • Foliage Texture: Fine • Fall Conspicuous: No • Leaf Retention: Yes

Size/Age/Growth

Active Growth Period: Spring and Summer • Growth Rate: Moderate • After Harvest Regrowth Rate: Slow • Mature Height (feet): 4.0 • Maximum Height at 20 Years (feet): 4 • Size: 24-36" tall. • Lifespan: Lifespan

Habitat

Typically found at an altitude of 0 to 2,916 meters (0 to 9,567 feet).[3]

Biology

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Reproduction

Duration: PerennialCoppice Potential: No • Progagated by Bulbs: No • Propagated by Bare Root: Yes • Propagated by Container: Yes • Propagated by Corms: No • Propagated by Cuttings: Yes • Propagated by Seed: Yes • Propagated by Sod: No • Propagated by Sprigs: No • Propagated by Tubers: No • Fruit/Seed Period Begin: Summer • Fruit/Seed Period End: FallFruit/Seed Persistence: No

Growth

Culture: Space 18-24" apart.

Soil: Adapted to Medium Textured: Adapted to Medium Textured Soils • Adapted to Coarse Textured Soils: Yes • Anaerobic Tolerance: Low • Salinity Tolerance: Low • CaCO3 Tolerance: Low • Minimum pH: 5.0 • Maximum pH: 6.0 • Fertility Requirement: Low

Sunlight: Sun Exposure: Sun to Partial Shade. • Shade Tolerance: Intermediate

Moisture: Drought Tolerance: Low • Minimum Precipitation: 35 • Maximum Precipitation: 70 • Moisture Use: High

Temperature: Minimum Temperature (F): -28 • Minimum Frost Free Days: 110 • Cold Hardiness: 3a, 3b, 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b. (map)

Taxonomy

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Unambiguous Synonyms

  1. Andromeda calyculata L.
  2. Andromeda calyculata Linnaeus
  3. Cassandra calyculata (L.) D. Don

Notes

Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000

Place of publication : Methodus 457. 1794

Name verified on 27-Nov-1998 by ARS Systematic Botanists. Last updated: 27-Nov-1998

Similar Species

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Members of the genus Chamaedaphne

ZipcodeZoo has pages for 14 species, subspecies, varieties, forms, and cultivars in this genus:

C. angustifolia · C. caliculata · C. calyculata (Dwarf Cassandra) · C. calyculata nana (Leatherleaf) · C. calyculata 'Tiny Tom' · C. calyculata var. angustifolia (Leatherleaf) · C. calyculata var. latifolia (Leatherleaf) · C. calyculata var. nana (Leatherleaf) · C. calyculata 'Verdant' (Verdant Leatherleaf) · C. crispa · C. cuneata · C. glauca · C. hirsuta · C. latifolia (Leatherleaf)

More Info

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Further Reading

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Notes

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Contributors

Data Sources

Accessed through GBIF Data Portal November 12, 2007:

Identifiers

Footnotes

  1. Mingyuan Fang, Ruizheng Fang, Mingyou He, Linzheng Hu, Hanbi Yang, Haining Qin, Tianlu Min, David F. Chamberlain, Peter Stevens, Gary D. Wallace & Arne Anderberg "Ericaceae". in Flora of China Vol. 14 Page 242. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
  2. Fang Rhui-cheng, Peter F. Stevens "Chamaedaphne". in Flora of China Vol. 14 Page 464. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
  3. Mean = 278.540 meters (913.845 feet), Standard Deviation = 248.300 based on 466 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
Last Revised: 7/1/2009