Common Names
Common Names in English:
Cassandra, Dwarf Cassandra, Leatherleaf
Description
Family Ericaceae
Plants
usually woody, or herbs, sometimes lacking chlorophyll. Leaves spiral
, sometimes decussate, margin
often toothed
. Inflorescence racemose; bracteoles paired
, basal. Flowers (4 or) 5-merous. Calyx imbricate. Corolla connate
, imbricate. Stamens 10, sometimes with spurs or awns
, dehiscing by pores
, pollen in tetrads
, rarely single. Ovary superior or inferior, placentation axile
, rarely parietal
, often many ovules per locule. Style ± as long as corolla, slender. Fruit a capsule or berry, rarely a drupe; calyx persistent.
About 125 genera and 4000 species: widely distributed in temperate
and subarctic
regions, also at high elevations
in tropical regions
; 22 genera and 826 species (524 endemic) in China.
The Monotropoideae are here included
in the Ericaceae; previously in FRPS (56: 157 216. 1990), they were treated as the Pyrolaceae. Chiogenes, recorded from China in FRPS (57(3) : 69 71. 1991), is here included in Gaultheria. Over the last half century, the Empetraceae have usually been separated from, but closely associated with, the Ericaceae. In their ecology, leaf morphology and insertion
, rusts, embryology, stamen anatomy, etc.
, they largely agree with that family
. Molecular data place Empetrum and its relatives firmly within the Ericaceae, and in particular within the subfamily
Ericoideae, in agreement with phytochemical and palynological data, and there they are best recognized as a separate tribe
. There are distinctive features of the Empetreae that were responsible for their past familial status, e.g.
, reduced perianth with separate members
, low ovule number, enlarged stigmas, etc. However, these are likely to be derived features associated with wind pollination. See Kron et al.
(Bot. Rev. 68: 335 423. 2002) and the recent treatment of the Ericaceae by Stevens et al. (in Kubitzki, Fam. Gen. Vasc. Pl. 6: 145 194. 2004) .
Several genera and many species are ornamentals
. Some fruits of Vaccinium in N China are sweet and edible, but of no particular value to humans. Some species of Chamaedaphne, Craibiodendron, Leucothoë, Lyonia, Pieris, and Rhododendron contain more or less toxic
diterpenes, which are harmful to humans or domestic animals.[1]
Genus Chamaedaphne
Shrubs
evergreen
, lepidote
. Stems arching
. Leaf blade
leathery, margin
entire or inconspicuously denticulate
. Inflorescences terminal
, secund
racemes
; bracts leaflike. Flowers 5-merous. Corolla urceolate
to tubular
. Stamens included
; filaments
straight, subulate
, lacking appendages
, anthers
dehiscing by terminal pores. Ovary superior, with many ovules per locule; stigma truncate
. Capsule loculicidal; epicarp 5-valved, separating from 10-valved endocarp. Seeds many, small, wingless.
One species: China, N Japan, Mongolia, Russia (Siberia) ; NE Europe, North America.[2]
Physical Description
Habit: Shrub • Growth Form: Rhizomatous • Shape and Orientation: Irregular
Flowers: Bloom Period: Early Spring • Flower Color: White • Flower Conspicuous: No
Seeds: Seed per Pound: 96000 • Seed Spread Rate: Slow • Seedling Vigor: Medium • Fruit/Seed Abundance: Low • Fruit/Seed Color: Brown • Fruit/Seed Conspicuous: No • Cold Stratification Required: No
Foliage: Foliage Color: Yellow-Green • Foliage Porosity Summer: Porous • Foliage Porosity Winter: Porous • Foliage Texture: Fine • Fall Conspicuous: No • Leaf Retention: Yes
Size/Age/Growth
Active Growth Period: Spring and Summer • Growth Rate: Moderate • After Harvest Regrowth Rate: Slow • Mature Height (feet): 4.0 • Maximum Height at 20 Years (feet): 4 • Size: 24-36" tall. • Lifespan: Lifespan
Habitat
Typically found at an altitude of 0 to 2,916 meters (0 to 9,567 feet).[3]
Biology
Reproduction
Duration: Perennial • Coppice Potential: No • Progagated by Bulbs: No • Propagated by Bare Root: Yes • Propagated by Container: Yes • Propagated by Corms: No • Propagated by Cuttings: Yes • Propagated by Seed: Yes • Propagated by Sod: No • Propagated by Sprigs: No • Propagated by Tubers: No • Fruit/Seed Period Begin: Summer • Fruit/Seed Period End: Fall • Fruit/Seed Persistence: No
Growth
Culture: Space 18-24" apart.
Soil: Adapted to Medium Textured: Adapted to Medium Textured Soils • Adapted to Coarse Textured Soils: Yes • Anaerobic Tolerance: Low • Salinity Tolerance: Low • CaCO3 Tolerance: Low • Minimum pH: 5.0 • Maximum pH: 6.0 • Fertility Requirement: Low
Sunlight: Sun Exposure: Sun to Partial Shade. • Shade Tolerance: Intermediate
Moisture: Drought Tolerance: Low • Minimum Precipitation: 35 • Maximum Precipitation: 70 • Moisture Use: High
Temperature: Minimum Temperature (F): -28 • Minimum Frost Free Days: 110 • Cold Hardiness: 3a, 3b, 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Dilleniidae
(
)
- Takhtajan, 1967
- Superorder:
Ericanae
(
)
- Takhtajan, 1967
- Order:
Ericales
(
)
- Dumortier, 1829
- Family:
Ericaceae
(
)
- Durande, 1782, Nom. Cons.
- Heath Family
- Subfamily:
Vaccinioideae
(
)
- Tribe:
Gaultherieae
(
)
- Genus:
Chamaedaphne
(
)
- Moench, Methodus. 457. 1794, nom. cons., not Mitchell (1769).
- Indian Paintbrush
- Specific epithet:
calyculata
- Baill.
- Botanical name: - Chamaedaphne calyculata
- Specific epithet:
calyculata
- Baill.
- Genus:
Chamaedaphne
(
- Tribe:
Gaultherieae
(
- Subfamily:
Vaccinioideae
(
- Family:
Ericaceae
(
- Order:
Ericales
(
- Superorder:
Ericanae
(
- Subclass:
Dilleniidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Andromeda calyculata L.
- Andromeda calyculata Linnaeus
- Cassandra calyculata (L.) D. Don
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: 15-Mar-2000
Place of publication
: Methodus 457. 1794
Name verified on 27-Nov-1998 by ARS Systematic Botanists. Last updated: 27-Nov-1998
Similar Species
Members of the genus Chamaedaphne
ZipcodeZoo has pages for 14 species, subspecies, varieties, forms, and cultivars in this genus:
C. angustifolia · C. caliculata · C. calyculata (Dwarf Cassandra) · C. calyculata nana (Leatherleaf) · C. calyculata 'Tiny Tom' · C. calyculata var. angustifolia (Leatherleaf) · C. calyculata var. latifolia (Leatherleaf) · C. calyculata var. nana (Leatherleaf) · C. calyculata 'Verdant' (Verdant Leatherleaf) · C. crispa · C. cuneata · C. glauca · C. hirsuta · C. latifolia (Leatherleaf)
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Further Reading
- A provisional list of the parasitic fungi of Wisconsin. [Madison, 1914] ENG url p. 859, p. 913, p. 917, p. 932.
- Acta Soc. pro Fauna et Flora Fennica. Helsinki, Societas. ENG url p. 498.
- An ecological characterization of Coastal Maine (north and east of Cape Elizabeth) / Stewart I. Fefer and Patricia S. Schettig: principal investigators. Newton Corner, Mass.: Department of the Interior, U.S. Fish and Wildlife Service, Northeast Region, 1980. ENG url p. 17, p. 23, p. 44.
- An illustrated flora of the northern United States, Canada and the British possessions: from Newfoundland to the parallel of the southern boundary of Virginia, and from the Atlantic Ocean westward to the 102d meridian / by Nathaniel Lord Britton and Hon. New York: C. Scribner's sons, 1913. ENG url p. 687.
- Anglo-russkii biologicheskii slovar' / [avtory, I. N. Afanas'eva et al.; spetsial'nye nauch. redaktory, O. I. Chibisova i L. A. Koziar]. Moskva: Russkii iazyk, 1979. ENG url p. 358.
- Annotated list of the ferns and flowering plants of New York state, by Homer D. House. Albany, The University of the state of New York, 1924. ENG url p. 549.
- Catalog of hymenoptera in America north of Mexico / prepared cooperatively by specialists on the various groups of Hymenoptera under the direction of Karl V. Krombein [et al.]. Washington: Smithsonian Institution Press, 1979- ENG url p. 1755.
- Chinese Academy of Sciences. 1959–. Flora reipublicae popularis sinicae. (F China)
- Classification of wetlands and deepwater habitats of the United States / by Lewis M. Cowardin [et al.]. Washington, D.C.: Fish and Wildlife Service, U.S. Dept. of the Interior, 1979. ENG url p. 22, p. 37, p. 67, p. 84, p. 98, p. 99.
- Deam, C. C. 1940. Flora of Indiana. (F Indiana)
- Entomological news, and proceedings of the Entomological Section of the Academy of Natural Sciences of Philadelphia. Philadelphia[Entomological Rooms of the Academy of Natural Sciences] ENG url p. 66.
- Entomological news. [Philadelphia]American Entomological Society, 1925- ENG url p. 202.
- FWS/0BS. [Washington]Fish and Wildlife Service, U.S. Dept. of the Interior. ENG url p. 100, p. 123, p. 125, p. 129, p. 14, p. 20, p. 75, p. 94, p. 97.
- Fieldbook of native Illinois shrubs, by Leo R. Tehon Urbana, Natural History Survey Division, 1942. ENG url p. 224, p. 225.
- Fishery circular / U.S. Department of Commerce, Bureau of Fisheries. [Washington]: The Bureau, 1931-1939. ENG url p. 60.
- Flora Rossica; sive, Enumeratio plantarum in totius Imperii Rossici provinciis Europaeis, Asiaticis et Americanis hucusque observatarum, auctore Carolo Friderico a Ledebour. Stuttgartiae, Sumptibus Librariae E. Schweizerbart, 1842-1853. LAT url p. 911.
- Flora of Berkshire County, Massachusetts. Boston, 1922. ENG url p. 188, p. 310.
- Flora of Delaware and the Eastern Shore: an annotated list of the ferns and flowering plants of the peninsula of Delaware, Maryland and Virginia. [Wilmington]: Society of Natural History of Delaware, 1946. ENG url p. 198.
- Flora of Illinois, containing keys for identification of flowering plants and ferns. Notre Dame, Ind., University of Notre Dame Press, 1963. ENG url p. 177, p. 5.
- Flora of Indiana, by Charles C. Deam. Indianapolis, Wm. B. Burford printing co., contractor for state printing and binding, 1940. ENG url p. 738.
- Flora of Pennsylvania. Edited with the addition of analytical keys by John Kunkel Small. Boston, Ginn & Co., 1903. ENG url p. 240.
- Flora of the Indiana dunes, a handbook of the flowering plants and ferns of the lake Michigan Coast of Indiana and of the Calumet District, by Donald Culross Peattie. Chicago, Field Museum of Natural History, 1930. ENG url p. 294.
- Flora of the southeastern United States; being descriptions of the seed-plants, ferns and fern-allies growing naturally in North Carolina, South Carolina, Georgia, Florida, Tennessee, Alabama, Mississippi, Arkansas, Louisiana, and in Oklahom by John Kunke New York, The author, 1913. ENG url p. 887.
- Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2. (Glea Cron ed2)
- Handbook of the flora of Philadelphia and vicinity, containing data relating to the plants within the following radius: eastern Pennsylvania; all of New Jersey except the northern counties; and New Castle County, Delaware, with keys for identification co Philadelphia, Philadelphia Botanical Club, 1905. ENG url p. 247.
- Inland wetlands of the United States: evaluated as potential registered natural landmarks / Richard H. Goodwin, William A. Niering. [Washington]: National Park Service: may be purchased from the Supt. of Docs., Govt. Print. Off., 1975. ENG url p. 19.
- Jahresbericht über das Gebiet der Pflanzenkrankheiten. Berlin: P. Parey, 1907-1917. GER url p. 170, p. 301, p. 374.
- Jones, G. N. & G. D. Fuller. 1955. Vascular plants of Illinois. (F IllJones)
- Journal of the Elisha Mitchell Scientific Society. Chapel Hill, N.C. [etc.]:Elisha Mitchell Scientific Society [etc.], 1884-2001. ENG url p. 108, p. 141.
- Just's botanischer jahresbericht. Systematisch geordnetes repertorium der botanischen literatur aller länder. Berlin, Gebr. Borntraeger, 1874-98; GER url p. 1101, p. 125, p. 1392, p. 1752, p. 204, p. 36, p. 498, p. 52, p. 560, p. 679, p. 718, p. 849, p. 927, p. 930.
- Komarov, V. L. et al., eds. 1934–1964. Flora SSSR. (F USSR)
- Meddelanden af Societatis pro Fauna et Flora Fennica. Helsingfors[Societas pro Fauna et Flora Fennica] SWE url p. 101, p. 136, p. 97.
- Michigan flora: a list of the fern and seed plants growing without cultivation / prepared by W.J. Beal. [Lansing, Mich.]: State Board of Agriculture: 1904. ENG url p. 31.
- Miscellaneous publication - University of Kansas, Museum of Natural History. Lawrence, University of Kansas, 1946-1996. ENG url p. 166, p. 172, p. 24, p. 25, p. 7, p. 91.
- Mitchell, R. S., ed. 1986. A checklist of New York state plants. New York State Mus. Bull. 458. (L New York)
- National list of scientific plant names. [Washington, D.C.]: U.S. Dept. of Agriculture, Soil Conservation Service, 1982- ENG url p. 83.
- Naturalist's guide to the Americas, prepared by the Committee on the Preservation of Natural Conditions of the Ecological Society of America, with assistance from numerous organizations and individuals, assembled and edited by chairman, Baltimore, Williams & Wilkins, 1926. ENG url p. 117, p. 269, p. 311.
- Ohio Flora Committee (E. L. Braun, T. S. Cooperrider, T. R. Fisher, J. J. Furlow). 1967–. The vascular flora of Ohio. (F Ohio)
- Ohwi, J. 1965. Flora of Japan (Engl. ed.). (F JapanOhwi)
- Ownbey, G. B. & T. Morley. 1991. Vascular plants of Minnesota: a checklist and atlas. (L Minn)
- Proceedings and transactions of the Royal Society of Canada. Délibérations et mémoires de la Société royale du Canada. Ottawa [etc.]Société royale du Canada. FRE url p. 136, p. 139.
- Proceedings of the Academy of Natural Sciences of Philadelphia. [Philadelphia, Academy of Natural Sciences of Philadelphia] ENG url p. 102, p. 109, p. 404.
- Proceedings of the United States National Museum. Washington: Smithsonian Institution Press, [etc.] ENG url p. 419.
- Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas. (F Carolin)
- Report of the Michigan Academy of Science. Lansing, Mich.: The Academy, 1904-1916. ENG url p. 127, p. 41.
- Rhoads, A. F. & W. M. Klein, Jr. 1993. The vascular flora of Pennsylvannia. Annotated checklist and atlas. (L Penn)
- Sand dunes and salt marshes / by Charles Wendell Townsend. Boston: L.C. Page & company, c1925. ENG url p. 306.
- Scoggan, H. J. 1978–1979. The flora of Canada, 4 vol. (F Canada)
- Seymour, F. 1969. The flora of New England. (F NEng)
- Stone, W. 1973. The plants of southern New Jersey. (F SNJersey)
- The Canadian field-naturalist. Ottawa, The Ottawa-Field-Naturalists Club. ENG url p. 27.
- The Journal of the College of Science, Imperial University of Tokyo, Japan = Tokyo Teikoku Daigaku kiyo. Rika. Tokyo, Japan: The University, 1898-1925. ENG url p. 99.
- The Nautilus. [Melbourne, Fla., etc., American Malacologists, inc., etc.] ENG url p. 23.
- The North American cup-fungi (inoperculates) New York, 1951. ENG url p. 294, p. 388.
- The Ohio journal of science. [Columbus, Ohio, The Ohio State University and the Ohio Academy of Science] ENG url p. 17, p. 208, p. 329, p. 416, p. 471, p. 52, p. 56.
- The Plant world. Baltimore [etc.]Plant World Association [etc.] ENG url p. 207, p. 33.
- The University of Kansas science bulletin. [Lawrence]: University of Kansas, 1902-1996. ENG url p. 400, p. 975.
- The plants of Southern New Jersey; with especial reference to the flora of the Pine Barrens and the geographic distribution of the species. Trenton, 1911. ENG url p. 78.
- The plants of southern New Jersey; with especial reference to the flora of the pine barrens and the geographic distribution of the species. Trenton, 1911. ENG url p. 619.
- The study of plant communities: an introduction to plant ecology. San Francisco: W.H. Freeman, 1948. ENG url p. 373.
- The vascular flora of the eastern Penobscot Bay region, Maine. [Portland, Maine, 1919. ENG url p. 257.
- Transactions - The Connecticut Academy of Arts and Sciences. New Haven: Connecticut Academy of Arts and Sciences; distributed by Archon Books, Hamden, Conn. [etc.], 1866- ENG url p. 358, p. 364, p. 402, p. 417, p. 446, p. 458.
- Trees and shrubs of Nantucket; Descriptions, identification keys, list of trees and shrubs [by] Mabel Agnes Rice. [Ann Arbor, Michigan., Lithoprinted by Edwards brothers, inc.]1946. ENG url p. 60, p. 75.
- Voss, E. 1972–. Michigan flora. (F Mich)
- Welsh, S. L. 1974. Anderson's flora of Alaska and adjacent parts of Canada. (F AlaskaAnd)
- Wetlands of the United States: current status and recent trends / by Ralph W. Tiner, Jr. [et al.]. Washington, D.C.: National Wetlands Inventory, Fish and Wildlife Service, U.S. Dept. of the Interior: 1984. ENG url p. 58.
- Woody plants in winter; a manual of common trees and shrubs in winter in the Northeastern United States and southeastern Canada, by Earl L. Core and Nelle P. Ammons. Pittsburgh, Boxwood Press[1958] ENG url p. 171.
- Woody plants of the western national parks; containing keys for the identification of trees and shrubs, by Virginia Long Bailey and Harold Edwards Bailey. Notre Dame, Ind., University Press, 1949. ENG url p. 261.
- Chou Yiliang & Zhou Ruichang. 1990. Pyrolaceae. In: Fang Wenpei & Hu Wenkuang, eds., Fl. Reipubl. Popularis Sin. 56: 157216
- Fang Rhuicheng, Yang Hanbi & Chin Tsenli. 1999. Ericaceae (1). In: Fang Rhuicheng, ed., Fl. Reipubl. Popularis Sin. 57(1): 1213
- Hsu Tingzhi, Gao Baochun, Fang Rhuicheng & Huang Shuhua. 1991. Ericaceae (3). In: Fang Rhuicheng, ed., Fl. Reipubl. Popularis Sin. 57(3): 1207
- Hu Wenkuang, Hu Lincheng, Fang Mingyuan & He Mingyou. 1994. Ericaceae (2). In: Hu Lincheng, Fang Mingyuan, ed., Fl. Reipubl. Popularis Sin. 57(2): 1438
- Ming Tien lu. 1980. Empetraceae. In: Cheng Mien & Ming Tien lu, eds., Fl. Reipubl. Popularis Sin. 45(1): 6062.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-2005. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. [http://sn2000.taxonomy.nl/Taxonomicon/]. Access date: Nov 23, 2005
- Global Biodiversity Information Facility. Accessed December 02, 2007. http://www.gbif.org Mediated distribution data from 2 providers.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 26, 2008)
Data Sources
Accessed through GBIF Data Portal November 12, 2007:
- Bundesamt für Naturschutz / Zentralstelle für Phytodiversität Deutschland, Bundesamt fuer Naturschutz / Zentralstelle fuer Phytodiversitaet Deutschland
- Canadian Museum of Nature, Canadian Museum of Nature Herbarium
- European Environment Agency, EUNIS
- Institute of Nature Conservation PAS, National System of Proetcted Areas
- Jyväskylä University Museum - The Section of Natural Sciences, Vascular plant collection of Jyvaskyla University Museum
- Marine Science Institute, UCSB, Paleobiology Database
- Missouri Botanical Garden, Missouri Botanical Garden
- The Swedish Museum of Natural History
- , Herbarium of Oskarshamn
- USDA PLANTS, USDA PLANTS Database
- University of Alabama Biodiversity and Systematics, Herbarium
- University of Alaska Museum of the North, University of Alaska Museum of the North Herbarium
- University of Washington Burke Museum, Vascular Plant Collection - University of Washington Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2649155
- Catalogue of Life Accepted Name Code: ITS-23637
- Global Biodiversity Information Facility Taxonkey: 13742531
- Globally Unique Identifier: urn:lsid:ipni.org:names:327542-1
- GRIN Nomen Number: 10105
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 23637
- International Plant Names Index (IPNI) ID: 327542-1
- Natural Heritage Network Species Identifier: PDERI08010
- U.S.D.A. Plant Symbol: CACAL6 CHC
- Zipcode Zoo Species Identifier: 30003
Footnotes
- Mingyuan Fang, Ruizheng Fang, Mingyou He, Linzheng Hu, Hanbi Yang, Haining Qin, Tianlu Min, David F. Chamberlain, Peter Stevens, Gary D. Wallace & Arne Anderberg "Ericaceae". in Flora of China Vol. 14 Page 242. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Fang Rhui-cheng, Peter F. Stevens "Chamaedaphne". in Flora of China Vol. 14 Page 464. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = 278.540 meters (913.845 feet), Standard Deviation = 248.300 based on 466 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
