Common Names
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Common Names in English:
Iberian Knapweed, Iberian Star Thistle, Iberian Starthistle, Iberian Starthistle Centaurea Iberica, Knapweed
Common Names in Portuguese:
Centáurea-Gigante
Description
Genus Centaurea
Annuals
, biennials, or perennials, 20-300 cm, glabrous
or tomentose
. Stems erect
, ascending
, or spreading
, simple
or branched. Leaves basal and cauline; petiolate
or sessile; proximal
blade
margins
often ± deeply lobed
, (spiny
in C. benedicta ), distal ± smaller, often entire
, faces
glabrous or ± tomentose, sometimes also villous
, strigose
, or puberulent
, often glandular-punctate. Heads discoid
, disciform
, or radiant, borne singly or in corymbiform
arrays. Involucres cylindric
or ovoid
to hemispheric
. Phyllaries many in 6-many series, unequal, proximal part appressed
, body margins entire. distal parts expanded into erect to spreading, usually ± dentate
or fringed
, linear
to ovate
appendages
, spine. tipped or spineless. Receptacles flat, epaleate, bristly
. Florets 10-many; outer usually sterile
, corollas slender and inconspicuous to much expanded, ± bilateral
; inner fertile
, corollas white to blue, pink, purple, or yellow, bilateral or radial
, often bent at junction of tubes
and throats
, lobes
linear-oblong, acute; anther
bases
tailed
, apical appendages oblong
; style
branches: fused portions with minutely hairy
nodes, distinct
portions minute. Cypselae ± barrel-shaped, ± compressed
, smooth
or ribbed
, apices entire (denticulate
in C.
benedicta ), glabrous or with fine, 1-celled hairs
, attachment scar
. lateral
(with or without elaiosomes) ; pappi 0 or ± persistent
, of 1-3 series of smooth or minutely barbed
, stiff bristles
or narrow scales
. x = 8, 9, 10, 11, 12, 13, 15.
Species ca.
500: introduced
; Eurasia
, n Africa, widely introduced worldwide.
Taxonomic
limits
of Centaurea have been controversial. The genus has great morphologic diversity
, and studies have revealed much cytologic (e.g.
, N. Garcia-Jacas et al.
1996) and palynologic (e.g., G. Wagenitz 1955) variation
as well. During the nineteenth and twentieth centuries, various taxonomists attempted, with limited success, to divide Centaurea into smaller genera or workable infrageneric
taxa. The relations of several satellite
genera have been controversial as well.
Recent molecular phylogenetic
studies (A. Susanna et al. 1995; N. Garcia-Jacas et al. 2000, 2001) have begun to clarify relationships
within Centaurea and between Centaurea and other genera. These studies make it clear that Centaurea as traditionally defined is polyphyletic, and that generic
boundaries should be realigned if monophyletic taxa are to be recognized. Some taxa traditionally included
within Centaurea (e.g., the two native
North American species, Centaurea americana and C. rothrockii) fall
outside the redefined generic boundaries and are here treated in Plectocephalus. Others usually placed into segregate
genera (e.g., Cnicus benedictus) are firmly nested within Centaurea. Because the type species of Centaurea (C. centaurium Linnaeus, an African species) falls
outside the main lineage
of the genus, a proposal
has been made to conserve Centaurea with a different type species (W. Greuter et al. 2001), thereby maintaining the nomenclatural
stability
of most of the numerous
species that do fall within the principal Centaurea clade.
Although several Centaurea species are widely established
as members
of the North American flora
, and some of these are widely distributed invasive weeds
, some of the taxa listed by J. T. Kartesz and C. A. Meacham (1999) are apparently waifs
and not permanent members of the flora. These taxa are discussed informally immediately below.
Although Cnicus has usually been recognized as a distinctive monotypic genus, it has been merged into Centaurea by various authors
(e.g., K
. Bremer 1994; G. Wagenitz and F. H. Hellwig 1996) . Recent molecular systematic studies (N. Garcia-Jacas et al. 2000) provide additional evidence that it is nested within Centaurea.David J. Keil, Jörg Ochsmann "Centaurea". in Flora of North America Vol. 19, 20 and 21 Page 52, 57, 58, 67, 83, 84, 96, 171, 172, 176, 177, EFloras.org.
Physical Description
Species Centaurea iberica
Annuals
, biennials, or short-lived perennials, 20-200 cm.
Stems 1-several, divaricately much branched, often forming
rounded
mound, puberulent
to loosely tomentose
. Leaves hispidulous
to loosely tomentose, ± glabrate
, minutely resin-gland-dotted;
proximal
leaves petiolate
, blades
10-20 cm, margins
1-2 times pinnately
lobed
or dissected
, rosette with central cluster
of spines; mid sessile,
not decurrent, blades ± lanceolate, shorter; distal blades
linear
to oblong
, entire
to coarsely dentate
or shallowly lobed.
Heads disciform
, borne singly or in leafy cymiform arrays,
sessile or short-pedunculate. Involucres ovoid
to hemispheric
,
(10-) 13-18 mm.
Principal phyllaries: bodies greenish or stramineous
,
ovate
, scarious-margined, appendages
stramineous, spiny-fringed at
base
, each tipped by stout spreading
spine (0.5-) 1-3 cm. Inner
phyllaries: appendages truncate
, spineless. Florets many;
corollas white, pink, or pale
purple, those of sterile
florets
slender,
15-20 mm, those of fertile
florets 15-20 mm. Cypselae white-
or brown-streaked, 3-4 mm, glabrous
; pappi of white bristles
1-2.5(-3) mm. 2n = 16, 20. [source]
Iberian star thistle is considered to be a noxious weed
in several
states of the western United
States. Weed control measures in Oregon
and Washington have apparently eradicated the species in those states.
Centaurea iberica is very similar to C.
calcitrapa,
from which it differs by its pappose cypselae and often more robust
habit. The Kansas and Wyoming plants
were originally reported as
C. calcitrapa (R. L. McGregor 1986). [source]
Habit: Forb/herb
Flowers: Bloom Period: May, June, July, August, September. • Flower Color: lavender, purple, violet
Size/Age/Growth
Size: 36-48" tall.
Habitat
Roadsides, pastures, fields
; 0-1500 m.
Centaurea iberica is
native
to southeastern Europe through central Asia (Ref. 52813).
Biology
Reproduction
Duration: Perennial
Growth
Culture: Space 24-36" apart.
Soil: Minimum pH: 6.1 • Maximum pH: 7.8
Sunlight: Sun Exposure: Full Sun .
Moisture: Drought Tolerance: High
Temperature: Cold Hardiness: Not Applicable (map)
Taxonomy
- Kingdom:
Plantae
(
)
- Plants
- Phylum:
Tracheophyta
(
)
- Vascular Plants
- Class:
Magnoliopsida
(
)
- Dicotyledons
- Order:
Asterales
(
)
- Family:
Noctuoidea
(
)
- Subfamily:
Cichorioideae
(
)
- Tribe:
Cardueae
(
)
- Genus:
Centaurea
(
)
- Linnaeus, Sp. Pl. 2: 909. 1753; Gen. Pl. ed. 5, 389. 1754.
- Knapweed, star thistle, cornflower [Greek kentaurieon, ancient plant name associated with Chiron, a centaur famous for knowledge of medicinal plants]
- Specific epithet:
iberica
- Treviranus ex Sprengel, Syst. Veg. 3: 406. 1826.
- Botanical name: - Centaurea iberica Trev. ex Spreng.
- Specific epithet:
iberica
- Treviranus ex Sprengel, Syst. Veg. 3: 406. 1826.
- Genus:
Centaurea
(
- Tribe:
Cardueae
(
- Subfamily:
Cichorioideae
(
- Family:
Noctuoidea
(
- Order:
Asterales
(
- Class:
Magnoliopsida
(
- Phylum:
Tracheophyta
(
Unambiguous Synonyms
- Centaurea iberica Trev. ex Spreng.
- Centaurea iberica Trev. ex Sprengel
- Centaurea iberica Trevir. ex Spreng.
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: 15-Mar-2000
Place of publication
: Syst. veg. 3:406. 1826
Name verified on 11-Jun-1991 by ARS Systematic Botanists. Last updated: 22-May-1997
Similar Species
Members of the genus Centaurea
There are approximately 2566 species in this genus. Here are just 100 of them:
C. 'Caramia' · C. 'John Coutts' · C. 'Jordy' · C.'Nigra' (Bachelor Buttons) · C. abbreviata · C. abchasica · C. abdelkaderi · C. abdelkaridi · C. abnormis · C. abrotanifolia · C. absinthifolia · C. abyssinica · C. acanthodes · C. acarnanica · C. acaulis · C. acaulis subsp. boissieri · C. achaia · C. achaia achaia · C. achaia subsp. corinthiaca · C. achtarovii · C. achtarovii subsp. sooana · C. acicularis · C. acmophylla · C. acrolepis · C. acromaura · C. acuta · C. acutangula · C. acutifolia · C. acutiloba · C. adami · C. adamovicii · C. adeana · C. adjarica · C. adonidifolia · C. adpressa · C. adulterina · C. adusta · C. aegialophila · C. aegyptiaca · C. aemiliae · C. aemilii · C. aemula · C. aemulans · C. aeolica · C. aeolica subsp. pandataria · C. aetaliae · C. aetolica · C. affinis · C. affinis candida · C. affinis denudata · C. affinis lacerata · C. affinis subsp. balcanica · C. affinis subsp. candida · C. affinis subsp. chalcidicea · C. affinis subsp. denudata · C. affinis subsp. lacerata · C. affinis subsp. peloponesiaca · C. affinis subsp. tartarea · C. africana · C. aggregata · C. ahverdovii · C. aichingeriana · C. ainetensis · C. aintabica · C. aintansis · C. ajtayana · C. akamantis · C. aksamica · C. aladaghensis · C. alaica · C. alata · C. alba · C. alba albanica · C. alba brunnea · C. alba euxina · C. alba maluqueri · C. alba princeps · C. alba subsp. albanica · C. alba subsp. brunnea · C. alba subsp. caliacrae · C. alba subsp. ciliata · C. alba subsp. costae · C. alba subsp. diomedea · C. alba subsp. epapposa · C. alba subsp. euxina · C. alba subsp. formanekii · C. alba subsp. heldreichii · C. alba subsp. ipecensis · C. alba subsp. latronum · C. alba subsp. leucomalla · C. alba subsp. macrocephala · C. alba subsp. maluqueri · C. alba subsp. montsicciana · C. alba subsp. sterilis · C. alba subsp. strepens · C. alba subsp. subciliaris · C. alba subsp. tartesiana · C. alba subsp. tenoreana · C. alba subsp. vandasii · C. alba tartesiana
Bibliography
- Chinese Academy of Sciences. 1959–. Flora reipublicae popularis sinicae. (F China)
- Davis, P. H., ed. 1965–1988. Flora of Turkey and the east Aegean islands. (F Turk)
- Komarov, V. L. et al., eds. 1934–1964. Flora SSSR. (F USSR)
- McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains. (F GPlains) [natzd.].
- Meikle, R. D. 1977–1985. Flora of Cyprus. (F Cyprus)
- Mouterde, P. 1966–. Nouvelle flore du Liban et de la Syrie. (F Liban)
- Munz, P. A. & D. D. Keck. 1959. A California flora. (F CalifMunz) [natzd.].
- Ochsmann, J. 2001. Knapweeds (Centaurea L. and some related genera) in North America. In: Ochsmann, J., Welcome to the Centaurea pages. (Centaurea Pages)
- Rechinger, K. H., ed. 1963–. Flora iranica. (F Iran)
- Rollins, R. & I. A. Al-Shehbaz. 1986. Proc. Roy. Soc. Edinburgh. 89:296.
- Seed Regulatory and Testing Branch, Agricultural Marketing Service, U.S.D.A. 1999. State noxious-weed seed requirements recognized in the administration of the Federal Seed Act. (State Noxweed Seed)
- Sharma, B. D. et al., eds. 1993–. Flora of India. (F India)
- Tutin, T. G. et al., eds. 1964–1980. Flora europaea. (F Eur)
- Zohary, M. & N. Feinbrun-Dothan. 1966–. Flora palaestina. (F Palest)
- Garcia-Jacas, N., A. Susanna, V. Mozaffarian, and R. Ilarslan. 2000. The natural delimitation of Centaurea (Asteraceae: Cardueae): ITS sequence analysis of the Centaurea jacea group. Pl. Syst. Evol. 223: 185199.
- Moore, R. J. 1972. Distribution of native and introduced knapweeds (Centaurea) in Canada and the United States. Rhodora 74: 331346.
- Roché, B. F. and C. T. Roché. 1991. Identification, introduction, distribution, ecology, and economics of Centaurea species. In: L. F. James et al., eds. 1991. Noxious Range Weeds. Boulder, San Francisco, and Oxford. Pp. 274291.
- Wagenitz, G. 1955. Pollenmorphologie und Systematik in der Gattung Centaurea L. s.l. Flora 142: 213279.
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Notes
Contributors
- "Centaurea iberica". in Flora of North America Vol. 19, 20 and 21 Page 183, 192. Published by Oxford University Press. Online at EFloras.org.
- Bisby FA, Roskov YR, Orrell TM, Nicolson D, Paglinawan LE, Bailly N, Kirk PM, Bourgoin T, van Hertum J, eds (2008). Species 2000 & ITIS Catalogue of Life: 2008 Annual Checklist Taxonomic Classification. CD-ROM; Species 2000: Reading, U.K.
- Global Biodiversity Information Facility. Accessed February 16, 2008. http://www.gbif.org Mediated distribution data from 10 providers.
- MBLWHOI Library: Universal Biological Index and Organizer. uBio.org accessed July 17, 2008.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 26, 2008)
Data Sources
Accessed through GBIF Data Portal February 16, 2008:
- Australian National Herbarium
- , Australian National Herbarium
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- Israel Nature and Parks Authority, Israel Nature and Parks Authority
- Oregon State University, Vascular Plant Collection
- The Swedish Museum of Natural History
- , Lund Botanical Museum
- UK National Biodiversity Network, Botanical Society of the British Isles - Vascular Plants Database
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2658260
- Catalogue of Life Accepted Name Code: ITS-36961
- Global Biodiversity Information Facility Taxonkey: 13749284
- Globally Unique Identifier: urn:lsid:ipni.org:names:190696-1
- GRIN Nomen Number: 316084
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 36961
- Natural Heritage Network Species Identifier: PDAST1Y090
- U.S.D.A. Plant Symbol: CEIB
- Zipcode Zoo Species Identifier: 27158
