Interesting Facts
Description
Family Cannabaceae
Herbs, annual
or perennial
, erect
or twining
, dioecious or sometimes monoecious, often with cystoliths
(a hard calcium carbonate
structure at base
of a hair) . Stems furrowed
or winged
. Stipules free
. Leaves alternate or opposite, palmately lobed
or compound
, sometimes simple
. Male inflorescences a bracteate
cymose
panicle. Male flowers: pedicellate
; sepals 5, free; petals absent; stamens 5, opposite sepals; filaments
short; anthers
2-loculed, dehiscent
by longitudinal
slits. Female inflorescences a bracteate spicate cyme much reduced in Cannabis, pendent or erect. Female flowers: sessile; calyx appressed
to ovary, membranous; petals absent; ovary 1-loculed; ovule solitary, pendulous from locule apex; style 2-parted, branches filiform
. Fruit an achene, covered by persistent
calyx; endosperm fleshy
; embryo curved
or spirally involute
.
Two genera and four species: N Africa, Asia, Europe, North America; two genera and four species (one endemic) in China.
Because all the Chinese species in this family
are cultivated and are often found naturalized
in disturbed
habitats
, it is difficult to know the true wild distributions.
Cannabaceae has sometimes been included
in Moraceae or Urticaceae but is now usually recognized as a distinct
family. The subfamily
Celtidoideae of Ulmaceae could possibly be included within Cannabaceae (see the discussion after the Ulmaceae family description
) .[1]
Genus Celtis
Trees
or rarely shrubs
, to 30 m
; crowns spreading
. Bark
usually gray, smooth
or often fissured
and conspicuously warty. Branches without or with thorns
, slender, glabrous
or pubescent
. Leaves: stipules falling early. Leaf blade
deltate to ovate
to oblong-lanceolate, base
oblique
or cuneate to rounded
, margins
entire or serrate-dentate; venation
3(-5) -pinnate. Inflorescences: staminate
inflorescences cymes or fascicles; pistillate
solitary or few-flowered clusters
. Flowers usually unisexual
, staminate and pistillate on same plants
, along with a few bisexual
flowers, pedicellate
on branches of current
year, appearing in mid or late spring
. Staminate flowers
: filaments
incurved
in bud, exserted after anthesis
; gynoecium minute, rudimentary
. Pistillate flowers: calyx slightly to deeply 4(-5) -lobed; stamens 4-5, inserted
on pilose
receptacle, included
, often nonfunctional filaments usually shorter than in staminate flowers, rarely absent; anthers
ovate, face
to face in bud, extrorse
; ovaries sessile, ovoid
, 1-locular; styles short, sessile, divided
into 2 divergent, elongate
, reflexed
lobes
, lobes entire or 2-cleft. Fruits fleshy
drupes, ovoid or globose
; outer mesocarp
thick, firm, inner mesocarp thin, fleshy; stones
thick walled, ripening in autumn, persisting after leaves fall
. x
= 10.
Species ca.
60: tropical
and temperate regions
, worldwide.
The hackberries provide important wildlife habitat
, forming thickets that give shelter
and fleshy drupes that ripen in autumn, persist after leaves fall, and supply winter food for birds and mammals. The treatment presented here is a simplified circumscription of species with no elaboration of infraspecific
variation
or interspecific
hybridization. The group is taxonomically complex
and in need of revision
.[2]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Urticanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Rosales
(
)
- Perleb, 1826
- Family:
Cannabaceae
(
)
- Augier, 1801 ex Martinov, 1820, nom. cons.
- hemp
- Genus:
Celtis
(
)
- C. Linnaeus, 1753
- Hackberry, sugarberry, bois inconnu [Classical Latin, Pliny's name for Celtis australis Linnaeus, the "lotus" of the ancient world]
- Specific epithet:
subdentata
- E.Mey. ex Planch.
- Botanical name: - Celtis subdentata E.Mey. ex Planch.
- Specific epithet:
subdentata
- E.Mey. ex Planch.
- Genus:
Celtis
(
- Family:
Cannabaceae
(
- Order:
Rosales
(
- Superorder:
Urticanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author : E.Mey. ex Planch. Publication : Ann. Sci. Nat., Bot. ser. 3, 10: 341 1848 [Dec 1848]
Similar Species
Members of the genus Celtis
ZipcodeZoo has pages for 31 species, subspecies, varieties, forms, and cultivars in this genus:
C. australis (European Hackberry) · C. bungeana (Hackberry) · C. ehrenbergiana (Spiny Hackberry) · C. iguanaea (Iguana Hackberry) · C. jessoensis (Japanese Hackberry) · C. julianae (Julian Hackberry) · C. laevigata (Net-Leaf Hackberry) · C. laevigata reticulata (Netleaf Hackberry) · C. laevigata var. brevipes (Sugar Hackberry) · C. laevigata var. laevigata (Sugarberry) · C. laevigata var. reticulata (Netleaf Hackberry) · C. laevigata var. texana (Texas Sugarberry) · C. laevigata 'All Seasons' (Sugar Hackberry) · C. lindheimeri (Lindheimer Hackberry) · C. occidentalis (American Hackberry) · C. occidentalis L. var. occidentalis L. (Common Hackberry) · C. occidentalis L. var. pumila (Pursh) A.Gray (Common Hackberry) · C. occidentalis var. canina (Common Hackberry) · C. occidentalis var. occidentalis (Western Hackberry) · C. occidentalis var. pumila (Dwarf Hackberry) · C. occidentalis 'Chicagoland' (American Hackberry) · C. occidentalis 'Delta' (American Hackberry) · C. occidentalis 'Prairie Pride' (American Hackberry) · C. occidentalis 'Windy City' (American Hackberry) · C. pallida (Capul) · C. reticulata var. reticulata (Netleaf Hackberry) · C. sinensis (Chinese Elm) · C. sinensis 'Green Cascade' (Chinese Elm) · C. tenuifolia (Dwarf Hackberry) · C. tournefortii (Oriental Hackberry) · C. trinervia (Almex)
More Info
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- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Catalogue of the African plants London, Printed by order of the Trustees, 1896-1901. url .
- The natural history of plants. By H. Baillon. Tr. by Marcus M. Hartog. London, L. Reeve & co., 1871-88. url p. 192.
- The natural history of plants. London, L. Reeve & Co., 1871-88. url .
- Chang Siushih. 1998. Cannaboideae. In: Chang Siushih & Wu Chengyih, eds., Fl. Reipubl. Popularis Sin. 23(1): 220224.
- Correll, D. S. and M. C. Johnston. 1970. Manual of the Vascular Plants of Texas. Renner, Tex.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Identifiers
- Biodiversity Heritage Library NamebankID: 10813297
- International Plant Names Index (IPNI) ID: 851181-1
- Zipcode Zoo Species Identifier: 3937215
Footnotes
- Zhengyi Wu, Zhe-Kun Zhou & Bruce Bartholomew "Cannabaceae". in Flora of China Vol. 5 Page 74. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Celtis". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
