Overview
Interesting Facts
- Celtis occidentalis is valued as an ornamental street tree because of its tolerance to drought . [source]
- Native Americans used decoctions prepared from the bark of Celtis occidentalis medicinally as an aid in menses and to treat sore throat (D. E. Moerman 1986). [source]
- This is a highly variable species. Segregates named as varieties follow an east-west geographic gradient and are based primarily on leaf size, shape , and pubescence . [source]
Common Names
Click on the language to view common names.
Common Names in Afrikaans:
Vals Witstinkhout
Common Names in English:
American Hackberry, Common Hackberry, Hackberry, Nettletree, Northern Hackberry, Sugarberry, Western Hackberry
Common Names in French:
Micocoulier Occidental
Common Names in unspecified:
Hackberry, Western Hackberry
Description
Family Cannabaceae
Herbs, annual
or perennial
, erect
or twining
, dioecious or sometimes monoecious, often with cystoliths
(a hard calcium carbonate
structure at base
of a hair) . Stems furrowed
or winged
. Stipules free
. Leaves alternate or opposite, palmately lobed
or compound
, sometimes simple
. Male inflorescences a bracteate
cymose
panicle. Male flowers: pedicellate
; sepals 5, free; petals absent; stamens 5, opposite sepals; filaments
short; anthers
2-loculed, dehiscent
by longitudinal
slits. Female inflorescences a bracteate spicate cyme much reduced in Cannabis, pendent or erect. Female flowers: sessile; calyx appressed
to ovary, membranous; petals absent; ovary 1-loculed; ovule solitary, pendulous from locule apex; style 2-parted, branches filiform
. Fruit an achene, covered by persistent
calyx; endosperm fleshy
; embryo curved
or spirally involute
.
Two genera and four species: N Africa, Asia, Europe, North America; two genera and four species (one endemic) in China.
Because all the Chinese species in this family
are cultivated and are often found naturalized
in disturbed
habitats
, it is difficult to know the true wild distributions.
Cannabaceae has sometimes been included
in Moraceae or Urticaceae but is now usually recognized as a distinct
family. The subfamily
Celtidoideae of Ulmaceae could possibly be included within Cannabaceae (see the discussion after the Ulmaceae family description
) .[1]
Genus Celtis
Trees
or rarely shrubs
, to 30 m
; crowns spreading
. Bark
usually gray, smooth
or often fissured
and conspicuously warty. Branches without or with thorns
, slender, glabrous
or pubescent
. Leaves: stipules falling early. Leaf blade
deltate to ovate
to oblong-lanceolate, base
oblique
or cuneate to rounded
, margins
entire or serrate-dentate; venation
3(-5) -pinnate. Inflorescences: staminate
inflorescences cymes or fascicles; pistillate
solitary or few-flowered clusters
. Flowers usually unisexual
, staminate and pistillate on same plants
, along with a few bisexual
flowers, pedicellate
on branches of current
year, appearing in mid or late spring
. Staminate flowers
: filaments
incurved
in bud, exserted after anthesis
; gynoecium minute, rudimentary
. Pistillate flowers: calyx slightly to deeply 4(-5) -lobed; stamens 4-5, inserted
on pilose
receptacle, included
, often nonfunctional filaments usually shorter than in staminate flowers, rarely absent; anthers
ovate, face
to face in bud, extrorse
; ovaries sessile, ovoid
, 1-locular; styles short, sessile, divided
into 2 divergent, elongate
, reflexed
lobes
, lobes entire or 2-cleft. Fruits fleshy
drupes, ovoid or globose
; outer mesocarp
thick, firm, inner mesocarp thin, fleshy; stones
thick walled, ripening in autumn, persisting after leaves fall
. x
= 10.
Species ca.
60: tropical
and temperate regions
, worldwide.
The hackberries provide important wildlife habitat
, forming thickets that give shelter
and fleshy drupes that ripen in autumn, persist after leaves fall, and supply winter food for birds and mammals. The treatment presented here is a simplified circumscription of species with no elaboration of infraspecific
variation
or interspecific
hybridization. The group is taxonomically complex
and in need of revision
.[2]
Physical Description
Species Celtis occidentalis
Trees or shrubs , size varying greatly in response to habitat ; crowns rounded . Bark gray, deeply furrowed , warty with age. Wood light yellow, weak. Branches without thorns , spreading , young branches mostly pubescent . Leaves: petiole 0.5-1.2 mm. Leaf blade lance-ovate to broadly ovate or deltate, 5-12 × 3-6(-9) cm (on fertile branches), leathery, base oblique or obliquely somewhat acuminate, margins conspicuously serrate to well below middle , teeth 10-40, apex acuminate; surfaces scabrous . Inflorescences dense pendulous clusters . Drupes dark orange to purple- or blue-black when ripe , orbicular , to 7-11(-20) mm diam., commonly with thick beak ; pedicel to 15 mm. Stones cream colored , 7-9 × 5-8 mm, reticulate . 2 n = 20, 30, and 40. [source]
ID Features: Alternate leaf arrangement. Oblique leaf base. Rough leaf. Corky bark. Small, imbricate lateral buds. Terminal bud absent. Zigzag stem pattern. Serrated leaf tips. Small, red drupe fruit.
Habit: Large deciduous tree . Cylindrical shape when mature . Branches tend to droop. • Growth Form: Single Stem • Shape and Orientation: Erect
Flowers: Blooms late April to early May. Emerge with leaves. Not ornamentally important. • Bloom Period: Mid Spring • Flower Color: Green • Flower Conspicuous: No
Seeds: Seed per Pound: 43000 • Seed Spread Rate: Moderate • Seedling Vigor: Medium • Fruit: Orangy-red fleshy fruit. Drupe. Ripen in September - October. Edible, sweet date taste. Not ornamentally important. Flesh surround hard seed that can break teeth. • Fruit/Seed Abundance: High • Fruit/Seed Color: Purple • Fruit/Seed Conspicuous: No • Cold Stratification Required: Yes
Foliage: Foliage Color: Green • Summer foliage: Alternate leaf arrangement . Leaves are simple and are 3" to 5" long. Oblique leaf base. Serrate margins at leaf tip. Medium green color, pale yellow-green leaf underside. Leaves are glabrous on underside especially around veins. Leaves have rough texture . • Fall foliage: Yellow fall color. Not showy. • Foliage Porosity Summer: Dense • Foliage Porosity Winter: Porous • Foliage Texture: Coarse • Fall Conspicuous: Yes • Leaf Retention: No
Size/Age/Growth
Active Growth Period: Spring and Summer • Growth Rate: Moderate-fast. • Mature Height (feet): 50.0 • Maximum Height at 20 Years (feet): 25 • Size: 40' to 60' tall and has an almost equal spread . • Vegetative Spread Rate: None • Lifespan: Lifespan
Landscaping
Landscape Uses: Good tree for poor conditions. Park tree. Screen . For edible fruit. • Liabilities: Leaf spot, powdery mildew , hackberry nipple gall, scale. Witches' broom caused by insect. Problems don't kill tree , but make tree very unattractive.
Habitat
Flowering late winter-spring (Mar-May). In rich moist soil along
streams
, on flood plains
, on rock, on wooded hillsides, and in woodlands;
0-1800 m
[3].
Zone 3.
Typically found at an altitude of 0 to 2,370 meters (0 to 7,776 feet).[4]
Biology
Reproduction
Duration: Perennial • Coppice Potential: Yes • Progagated by Bulbs: No • Propagated by Bare Root: Yes • Propagated by Container: Yes • Propagated by Corms: No • Propagated by Cuttings: No • Propagated by Seed: Yes • Propagated by Sod: No • Propagated by Sprigs: No • Propagated by Tubers: No • Fruit/Seed Period Begin: Summer • Fruit/Seed Period End: Fall • Fruit/Seed Persistence: Yes
Growth
Culture: Easily transplant form B&B container or bare root . Prefers rich soil. Very soil adaptable. Tolerant of most conditions. Wind tolerant. Full sun . Urban tolerant.
Soil: Adapted to Medium Textured: Adapted to Medium Textured Soils • Adapted to Coarse Textured Soils: Yes • Anaerobic Tolerance: Medium • Salinity Tolerance: None • CaCO3 Tolerance: Medium • Minimum pH: 6.0 • Maximum pH: 7.8 • Fertility Requirement: Medium
Sunlight: Sun Exposure: Full Sun . • Shade Tolerance: Intermediate
Moisture: Drought Tolerance: High • Minimum Precipitation: 14 • Maximum Precipitation: 32 • Moisture Use: Low
Temperature: Minimum Temperature (F): -38 • Minimum Frost Free Days: 90 • Cold Hardiness: 3a, 3b, 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Urticanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Rosales
(
)
- Perleb, 1826
- Family:
Cannabaceae
(
)
- Augier, 1801 ex Martinov, 1820, nom. cons.
- hemp
- Genus:
Celtis
(
)
- C. Linnaeus, 1753
- Hackberry, sugarberry, bois inconnu [Classical Latin, Pliny's name for Celtis australis Linnaeus, the "lotus" of the ancient world]
- Specific epithet:
occidentalis
- L.
- Botanical name: - Celtis occidentalis L.
- Specific epithet:
occidentalis
- L.
- Genus:
Celtis
(
- Family:
Cannabaceae
(
- Order:
Rosales
(
- Superorder:
Urticanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author : L. Publication : Sp. Pl. 2: 1044 1753 [1 May 1753]
Similar Species
Members of the genus Celtis
ZipcodeZoo has pages for 31 species, subspecies, varieties, forms, and cultivars in this genus:
C. australis (European Hackberry) · C. bungeana (Hackberry) · C. ehrenbergiana (Spiny Hackberry) · C. iguanaea (Iguana Hackberry) · C. jessoensis (Japanese Hackberry) · C. julianae (Julian Hackberry) · C. laevigata (Net-Leaf Hackberry) · C. laevigata reticulata (Netleaf Hackberry) · C. laevigata var. brevipes (Sugar Hackberry) · C. laevigata var. laevigata (Sugarberry) · C. laevigata var. reticulata (Netleaf Hackberry) · C. laevigata var. texana (Texas Sugarberry) · C. laevigata 'All Seasons' (Sugar Hackberry) · C. lindheimeri (Lindheimer Hackberry) · C. occidentalis (American Hackberry) · C. occidentalis L. var. occidentalis L. (Common Hackberry) · C. occidentalis L. var. pumila (Pursh) A.Gray (Common Hackberry) · C. occidentalis var. canina (Common Hackberry) · C. occidentalis var. occidentalis (Western Hackberry) · C. occidentalis var. pumila (Dwarf Hackberry) · C. occidentalis 'Chicagoland' (American Hackberry) · C. occidentalis 'Delta' (American Hackberry) · C. occidentalis 'Prairie Pride' (American Hackberry) · C. occidentalis 'Windy City' (American Hackberry) · C. pallida (Capul) · C. reticulata var. reticulata (Netleaf Hackberry) · C. sinensis (Chinese Elm) · C. sinensis 'Green Cascade' (Chinese Elm) · C. tenuifolia (Dwarf Hackberry) · C. tournefortii (Oriental Hackberry) · C. trinervia (Almex)
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Further Reading
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- A botanical survey of the Sugar Grove region / by Robert F. Griggs. Columbus: Ohio State University, 1914. url p. 325.
- A catalogue of the Coccidæ of the world. [By] Mrs. Maria E. Fernald, A.M. Amherst, Mass., Carpenter & Morehouse, 1903. url p. 197, p. 198, p. 257, p. 54.
- A dictionary of American plant names, compiled by Willard N. Clute. Joliet, Ill., W. N. Clute, 1923. url p. 164.
- A faunal investigation of Long Point and vicinity, Norfolk County, Ontario / [Toronto: Royal Ontario Museum of Zoology], 1931. url p. 122.
- A faunal investigation of Prince Edward County, Ontario / Toronto: University of Toronto Press, 1941. url p. 7.
- A field and laboratory guide in biological nature-study, by Elliot R. Downing. Chicago, Ill., The University of Chicago Press[c1918] url .
- A first report on the relations between climates and crops. By Cleveland Abbe. Prepared under the direction of Willis L. Moore, chief United States Weather Bureau. Washington, Govt. Print. Off., 1905. url p. 195.
- A flora of northwest America: containing brief descriptions of all the known indigenous and naturalized plants growing without cultivation north of California, west of Utah, and south of British Columbia / by Thomas Howell. Vol. 1, Phanerogamae. Portland, Or.: [s.n.], 1903. url p. 601, p. 601.
- A general introduction to forestry in the United States, with special reference to recent forest conservation policies, by Nelson Courtlandt Brown. New York, Chapman & Hall, limited, 1935. url p. 288.
- A manual for the study of insects / by John Henry Comstock and Anna Botsford Comstock. Ithaca, N.Y.: The Comstock Pub. Co., 1920. url p. 397, p. 682.
- A manual for the study of insects, by John Henry Comstock. .. and Anna Botsford Comstock. .. Ithaca, N.Y., Comstock Publishing Company, 1916 [c1895] url p. 682.
- A manual of poisonous plants, chiefly of eastern North America, with brief notes on economic and medicinal plants, and numerous illustrations, by L.H. Pammel Cedar Rapids, Ia., The Torch press, 1911. url p. 930.
- A manual of poisonous plants: chiefly of eastern North America, with brief notes on economic and medicinal plants, and numerous illustrations / by L. H. Pammel. 1911 Cedar Rapids, Ia.: The Torch Press, 1910-11. url p. 930.
- A narrative history of the town of Cohasset, Massachusetts. [Cohasset]Committee on Town History, 1898-1956. url p. 552.
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- Identification of the economic woods of the United States, including a discussion of the structural and physical properties of wood, New York, John Wiley & sons, inc.; [etc., etc.]1919. url , , , p. 51, p. 89.
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- Standardized plant names; a catalogue of approved scientific and common names of plants in American commerce. Salem, Mass., 1923. url p. 194.
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Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
- "Celtis occidentalis". in Flora of North America Vol. 3. Published by Oxford University Press. Online at EFloras.org.
- Global Biodiversity Information Facility. Accessed November 21, 2007. http://www.gbif.org Mediated distribution data from 20 providers.
- Ruggiero M., Gordon D., Bailly N., Kirk P., Nicolson D. (2011). The Catalogue of Life Taxonomic Classification, Edition 2, Part A. In: Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist (Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds). DVD; Species 2000: Reading, UK.
- The International Plant Names Index. Accessed Dec 27, 2011.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 24, 2008)
Data Sources
Accessed through GBIF Data Portal November 21, 2007:
- Forest Research Institute, Department of Natural Forests, Herbarium
- Missouri Botanical Garden, Missouri Botanical Garden
- The New York Botanical Garden, Bronx River Bioblitz
- The Swedish Museum of Natural History
- , Herbarium of Oskarshamn
- USDA PLANTS, USDA PLANTS Database
- University of Alabama Biodiversity and Systematics, Herbarium
- Utah Valley State College
- , Utah Valley State College Herbarium
- Yale University Peabody Museum, Yale Peabody Museum - Invertebrate Paleontology DiGIR provider service
Identifiers
- Biodiversity Heritage Library NamebankID: 2645770
- Catalogue of Life Accepted Name Code: ITS-19040
- Global Biodiversity Information Facility Taxonkey: 13727278
- Globally Unique Identifier: urn:lsid:ipni.org:names:851127-1
- GRIN Nomen Number: 9774
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19040
- International Plant Names Index (IPNI) ID: 851127-1
- Natural Heritage Network Species Identifier: PDULM01040
- U.S.D.A. Plant Symbol: CEOC
- Zipcode Zoo Species Identifier: 11528
Footnotes
- Zhengyi Wu, Zhe-Kun Zhou & Bruce Bartholomew "Cannabaceae". in Flora of China Vol. 5 Page 74. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Celtis". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- "Celtis occidentalis". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- Mean = 302.900 meters (993.766 feet), Standard Deviation = 245.450 based on 2,268 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
