Overview
Interesting Facts
Common Names
Click on the language to view common names.
Common Names in Afrikaans:
Chinese Netelboom
Common Names in Chinese:
Po Shu
Common Names in English:
Chinese Elm, Chinese Hackberry, Chinese Nettle-Tree, Japanese Hackberry
Description
Family Cannabaceae
Herbs, annual
or perennial
, erect
or twining
, dioecious or sometimes monoecious, often with cystoliths
(a hard calcium carbonate
structure at base
of a hair) . Stems furrowed
or winged
. Stipules free
. Leaves alternate or opposite, palmately lobed
or compound
, sometimes simple
. Male inflorescences a bracteate
cymose
panicle. Male flowers: pedicellate
; sepals 5, free; petals absent; stamens 5, opposite sepals; filaments
short; anthers
2-loculed, dehiscent
by longitudinal
slits. Female inflorescences a bracteate spicate cyme much reduced in Cannabis, pendent or erect. Female flowers: sessile; calyx appressed
to ovary, membranous; petals absent; ovary 1-loculed; ovule solitary, pendulous from locule apex; style 2-parted, branches filiform
. Fruit an achene, covered by persistent
calyx; endosperm fleshy
; embryo curved
or spirally involute
.
Two genera and four species: N Africa, Asia, Europe, North America; two genera and four species (one endemic) in China.
Because all the Chinese species in this family
are cultivated and are often found naturalized
in disturbed
habitats
, it is difficult to know the true wild distributions.
Cannabaceae has sometimes been included
in Moraceae or Urticaceae but is now usually recognized as a distinct
family. The subfamily
Celtidoideae of Ulmaceae could possibly be included within Cannabaceae (see the discussion after the Ulmaceae family description
) .[1]
Genus Celtis
Trees
or rarely shrubs
, to 30 m
; crowns spreading
. Bark
usually gray, smooth
or often fissured
and conspicuously warty. Branches without or with thorns
, slender, glabrous
or pubescent
. Leaves: stipules falling early. Leaf blade
deltate to ovate
to oblong-lanceolate, base
oblique
or cuneate to rounded
, margins
entire or serrate-dentate; venation
3(-5) -pinnate. Inflorescences: staminate
inflorescences cymes or fascicles; pistillate
solitary or few-flowered clusters
. Flowers usually unisexual
, staminate and pistillate on same plants
, along with a few bisexual
flowers, pedicellate
on branches of current
year, appearing in mid or late spring
. Staminate flowers
: filaments
incurved
in bud, exserted after anthesis
; gynoecium minute, rudimentary
. Pistillate flowers: calyx slightly to deeply 4(-5) -lobed; stamens 4-5, inserted
on pilose
receptacle, included
, often nonfunctional filaments usually shorter than in staminate flowers, rarely absent; anthers
ovate, face
to face in bud, extrorse
; ovaries sessile, ovoid
, 1-locular; styles short, sessile, divided
into 2 divergent, elongate
, reflexed
lobes
, lobes entire or 2-cleft. Fruits fleshy
drupes, ovoid or globose
; outer mesocarp
thick, firm, inner mesocarp thin, fleshy; stones
thick walled, ripening in autumn, persisting after leaves fall
. x
= 10.
Species ca.
60: tropical
and temperate regions
, worldwide.
The hackberries provide important wildlife habitat
, forming thickets that give shelter
and fleshy drupes that ripen in autumn, persist after leaves fall, and supply winter food for birds and mammals. The treatment presented here is a simplified circumscription of species with no elaboration of infraspecific
variation
or interspecific
hybridization. The group is taxonomically complex
and in need of revision
.[2]
Physical Description
Species Celtis sinensis
Trees , to 20 m tall, deciduous. Bark gray. Branchlets brown, brown pubescent , sometimes glabrescent late in season . Winter buds dark brown, 1-3 mm, glabrous or inconspicuously puberulent . Stipules linear to lanceolate, 3-5 mm, pubescent, fugacious . Petiole brown, 3-10 mm, pubescent, adaxially with a broad and shallow furrow; leaf blade ovate to ovate-elliptic, 3-10 × 3.5-6 cm, thickly papery , abaxially usually inconspicuously yellowish brown puberulent when young, abaxially with hairs scattered on major veins and sometimes tufted in vein axils with age, base rounded , obtuse , or obliquely truncate , ± symmetric to moderately oblique , margin subentire to crenate on apical half, teeth 0-16 on each side, apex acute to shortly acuminate; secondary veins 3 or 4 on each side of midvein . Flowers fascicled in leaf axils and stem bases. Style branches linear, undivided. Infructescences unbranched, 1(-3) per leaf axil, rather stout, pubescent at least proximally, 4-10 mm; fruiting pedicel 1-1.5 × as long as subtending petiole. Drupe ± globose , 5-7(-8) mm in diam. Stone white, ± globose, reticulately foveolate, ribbed . Fl. Mar-Apr, fr. Sep-Oct. [source]
Habit: Deciduous.
Flowers: Bloom Period: March. • Flower Color: inconspicuous, none, orange
Size/Age/Growth
Size: over 40' tall.
Habitat
Roadsides, slopes ; 100-1500 m [3].
Biology
Growth
Culture: Space 30-40' apart.
Soil: Minimum pH: 4.5 • Maximum pH: 7.8
Sunlight: Sun Exposure: Full Sun .
Moisture: Drought Tolerance: High
Temperature: Cold Hardiness: 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b, 9a, 9b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Urticanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Rosales
(
)
- Perleb, 1826
- Family:
Cannabaceae
(
)
- Augier, 1801 ex Martinov, 1820, nom. cons.
- hemp
- Genus:
Celtis
(
)
- C. Linnaeus, 1753
- Hackberry, sugarberry, bois inconnu [Classical Latin, Pliny's name for Celtis australis Linnaeus, the "lotus" of the ancient world]
- Specific epithet:
sinensis
- Pers.
- Botanical name: - Celtis sinensis Pers.
- Specific epithet:
sinensis
- Pers.
- Genus:
Celtis
(
- Family:
Cannabaceae
(
- Order:
Rosales
(
- Superorder:
Urticanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
C. bungeana var. pubipedicella G. H. Wang • C. cercidifolia C. K. Schneider • C. hunanensis Handel-Mazzetti; C. Labilis C. K. Schneider • C. nervosa Hemsley • C. tetrandra Roxburgh sinensis (Persoon) Y. C. Tang. • Celtis bodinieri H. Léveillé
Notes
Publishing author : Pers. Publication : Syn. Pl. (Persoon) 1: 292 1805 [1 Apr-15 Jun 1805]
Similar Species
Members of the genus Celtis
ZipcodeZoo has pages for 31 species, subspecies, varieties, forms, and cultivars in this genus:
C. australis (European Hackberry) · C. bungeana (Hackberry) · C. ehrenbergiana (Spiny Hackberry) · C. iguanaea (Iguana Hackberry) · C. jessoensis (Japanese Hackberry) · C. julianae (Julian Hackberry) · C. laevigata (Net-Leaf Hackberry) · C. laevigata reticulata (Netleaf Hackberry) · C. laevigata var. brevipes (Sugar Hackberry) · C. laevigata var. laevigata (Sugarberry) · C. laevigata var. reticulata (Netleaf Hackberry) · C. laevigata var. texana (Texas Sugarberry) · C. laevigata 'All Seasons' (Sugar Hackberry) · C. lindheimeri (Lindheimer Hackberry) · C. occidentalis (American Hackberry) · C. occidentalis L. var. occidentalis L. (Common Hackberry) · C. occidentalis L. var. pumila (Pursh) A.Gray (Common Hackberry) · C. occidentalis var. canina (Common Hackberry) · C. occidentalis var. occidentalis (Western Hackberry) · C. occidentalis var. pumila (Dwarf Hackberry) · C. occidentalis 'Chicagoland' (American Hackberry) · C. occidentalis 'Delta' (American Hackberry) · C. occidentalis 'Prairie Pride' (American Hackberry) · C. occidentalis 'Windy City' (American Hackberry) · C. pallida (Capul) · C. reticulata var. reticulata (Netleaf Hackberry) · C. sinensis (Chinese Elm) · C. sinensis 'Green Cascade' (Chinese Elm) · C. tenuifolia (Dwarf Hackberry) · C. tournefortii (Oriental Hackberry) · C. trinervia (Almex)
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- A naturalist in western China, with vasculum, camera, and gun; being some account of eleven years' travel, exploration, and observation in the more remote parts of the Flowery kingdom, by Ernest Henry Wilson...with an introduction by Charles Sprague Sargent...With one hundred and one full-page illustrations and a map. London, Methuen & co., ltd.[1913] url p. 20, p. 213.
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Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
- "Celtis sinensis". in Flora of China Vol. 5 Page 18. Published by Missouri Botanical Garden Press. Online at EFloras.org.
- Global Biodiversity Information Facility. Accessed November 19, 2007. http://www.gbif.org Mediated distribution data from 10 providers.
- The International Plant Names Index. Accessed Dec 27, 2011.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 27, 2008)
Data Sources
Accessed through GBIF Data Portal November 19, 2007:
- Australian National Herbarium
- , Australian National Herbarium
- International Plant Genetic Resources Institute(IPGRI), The System-wide Information Network for Genetic Resources
- National Herbarium of New South Wales, NSW herbarium collection
- National Science Museum of Korea, National Science Museum of Korea Plant
- Taiwan Biodiversity Information Facility, Magnoliophyta
Identifiers
- Biodiversity Heritage Library NamebankID: 5829235
- Global Biodiversity Information Facility Taxonkey: 15886654
- Globally Unique Identifier: urn:lsid:ipni.org:names:851173-1
- GRIN Nomen Number: 9781
- International Plant Names Index (IPNI) ID: 851173-1
- Zipcode Zoo Species Identifier: 707205
Footnotes
- Zhengyi Wu, Zhe-Kun Zhou & Bruce Bartholomew "Cannabaceae". in Flora of China Vol. 5 Page 74. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Celtis". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- "Celtis sinensis". in Flora of China Vol. 5 Page 18. Missouri Botanical Garden Press. Online at EFloras.org. [back]
