Overview
Interesting Facts
Common Names
Click on the language to view common names.
Common Names in English:
Iguana Hackberry
Common Names in French:
Croc-á-Chiens, Gratte-Jambes, Liane Crabe
Common Names in Portuguese:
Grapia
Description
Family Cannabaceae
Herbs, annual
or perennial
, erect
or twining
, dioecious or sometimes monoecious, often with cystoliths
(a hard calcium carbonate
structure at base
of a hair) . Stems furrowed
or winged
. Stipules free
. Leaves alternate or opposite, palmately lobed
or compound
, sometimes simple
. Male inflorescences a bracteate
cymose
panicle. Male flowers: pedicellate
; sepals 5, free; petals absent; stamens 5, opposite sepals; filaments
short; anthers
2-loculed, dehiscent
by longitudinal
slits. Female inflorescences a bracteate spicate cyme much reduced in Cannabis, pendent or erect. Female flowers: sessile; calyx appressed
to ovary, membranous; petals absent; ovary 1-loculed; ovule solitary, pendulous from locule apex; style 2-parted, branches filiform
. Fruit an achene, covered by persistent
calyx; endosperm fleshy
; embryo curved
or spirally involute
.
Two genera and four species: N Africa, Asia, Europe, North America; two genera and four species (one endemic) in China.
Because all the Chinese species in this family
are cultivated and are often found naturalized
in disturbed
habitats
, it is difficult to know the true wild distributions.
Cannabaceae has sometimes been included
in Moraceae or Urticaceae but is now usually recognized as a distinct
family. The subfamily
Celtidoideae of Ulmaceae could possibly be included within Cannabaceae (see the discussion after the Ulmaceae family description
) .[1]
Genus Celtis
Trees
or rarely shrubs
, to 30 m
; crowns spreading
. Bark
usually gray, smooth
or often fissured
and conspicuously warty. Branches without or with thorns
, slender, glabrous
or pubescent
. Leaves: stipules falling early. Leaf blade
deltate to ovate
to oblong-lanceolate, base
oblique
or cuneate to rounded
, margins
entire or serrate-dentate; venation
3(-5) -pinnate. Inflorescences: staminate
inflorescences cymes or fascicles; pistillate
solitary or few-flowered clusters
. Flowers usually unisexual
, staminate and pistillate on same plants
, along with a few bisexual
flowers, pedicellate
on branches of current
year, appearing in mid or late spring
. Staminate flowers
: filaments
incurved
in bud, exserted after anthesis
; gynoecium minute, rudimentary
. Pistillate flowers: calyx slightly to deeply 4(-5) -lobed; stamens 4-5, inserted
on pilose
receptacle, included
, often nonfunctional filaments usually shorter than in staminate flowers, rarely absent; anthers
ovate, face
to face in bud, extrorse
; ovaries sessile, ovoid
, 1-locular; styles short, sessile, divided
into 2 divergent, elongate
, reflexed
lobes
, lobes entire or 2-cleft. Fruits fleshy
drupes, ovoid or globose
; outer mesocarp
thick, firm, inner mesocarp thin, fleshy; stones
thick walled, ripening in autumn, persisting after leaves fall
. x
= 10.
Species ca.
60: tropical
and temperate regions
, worldwide.
The hackberries provide important wildlife habitat
, forming thickets that give shelter
and fleshy drupes that ripen in autumn, persist after leaves fall, and supply winter food for birds and mammals. The treatment presented here is a simplified circumscription of species with no elaboration of infraspecific
variation
or interspecific
hybridization. The group is taxonomically complex
and in need of revision
.[2]
Physical Description
Species Celtis iguanaea
Climbing shrub with 2 recurved spines at nodes; many parallel regular branches from main stem ; Leaves trinerved , crenate or serrate beyond the middle ; with tufts of hairs or pits in the axils of the lateral nerves ; base often cordate; 2mm stipules. Frutis fleshy , 6-10mm, truning orange, with 2 styles, each splitting into 2, remaining on top.
Habit: Tree , Vine , Shrub
Habitat
Typically found at an altitude of 0 to 4,936 meters (0 to 16,194 feet).[3]
Biology
Reproduction
Duration: Perennial
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Urticanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Rosales
(
)
- Perleb, 1826
- Family:
Cannabaceae
(
)
- Augier, 1801 ex Martinov, 1820, nom. cons.
- hemp
- Genus:
Celtis
(
)
- C. Linnaeus, 1753
- Hackberry, sugarberry, bois inconnu [Classical Latin, Pliny's name for Celtis australis Linnaeus, the "lotus" of the ancient world]
- Specific epithet:
iguanaea
- (Jacq.) Sarg.
- Botanical name: - Celtis iguanaea (Jacq.) Sarg.
- Specific epithet:
iguanaea
- (Jacq.) Sarg.
- Genus:
Celtis
(
- Family:
Cannabaceae
(
- Order:
Rosales
(
- Superorder:
Urticanae
(
- Subclass:
Rosidae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Synonyms
Mertensia iguanaea (Jacq.) Schult. • Momisia iguanaea (Jacq.) Rose & Standl. • Momisia iguanaea (Jacq.) Rose and Standl. • Rhamnus Iguanaea • Rhamnus iguanaea Jacq. • Ziziphus Iguanaea • Ziziphus iguanaea (Jacq.) Lam. • Zizyphus iguanaea (Jacq.) Lam.
Notes
Publishing author
: Sarg.
Basionym
author: (Jacq.)
Name
Status: Accepted Name
.
Last scrutiny: 15-Mar-2000
Similar Species
Members of the genus Celtis
ZipcodeZoo has pages for 31 species, subspecies, varieties, forms, and cultivars in this genus:
C. australis (European Hackberry) · C. bungeana (Hackberry) · C. ehrenbergiana (Spiny Hackberry) · C. iguanaea (Iguana Hackberry) · C. jessoensis (Japanese Hackberry) · C. julianae (Julian Hackberry) · C. laevigata (Net-Leaf Hackberry) · C. laevigata reticulata (Netleaf Hackberry) · C. laevigata var. brevipes (Sugar Hackberry) · C. laevigata var. laevigata (Sugarberry) · C. laevigata var. reticulata (Netleaf Hackberry) · C. laevigata var. texana (Texas Sugarberry) · C. laevigata 'All Seasons' (Sugar Hackberry) · C. lindheimeri (Lindheimer Hackberry) · C. occidentalis (American Hackberry) · C. occidentalis L. var. occidentalis L. (Common Hackberry) · C. occidentalis L. var. pumila (Pursh) A.Gray (Common Hackberry) · C. occidentalis var. canina (Common Hackberry) · C. occidentalis var. occidentalis (Western Hackberry) · C. occidentalis var. pumila (Dwarf Hackberry) · C. occidentalis 'Chicagoland' (American Hackberry) · C. occidentalis 'Delta' (American Hackberry) · C. occidentalis 'Prairie Pride' (American Hackberry) · C. occidentalis 'Windy City' (American Hackberry) · C. pallida (Capul) · C. reticulata var. reticulata (Netleaf Hackberry) · C. sinensis (Chinese Elm) · C. sinensis 'Green Cascade' (Chinese Elm) · C. tenuifolia (Dwarf Hackberry) · C. tournefortii (Oriental Hackberry) · C. trinervia (Almex)
More Info
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Further Reading
- Bulletin / Smithsonian Institution, Bureau of American Ethnology. Washington: G.P.O., 1901-1971. url p. 202, p. 220, p. 363.
- Contributions from the United States National Herbarium 27 1928 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 157, p. 200, p. 201, p. 21, p. 387, p. 462, p. 8.
- Contributions from the University of Michigan Herbarium. 9 1966 Ann Arbor: University Herbarium, University of Michigan, 1939- url p. 93.
- Flora Costaricensis / William Burger, editor. 40 1977 Chicago, Ill.: Field Museum of Natural History, 1977. url p. 87, p. 88, p. 89.
- Flora of Costa Rica. .. by Paul C. Standley. .. 18 1937 Chicago, 1937. url p. 377.
- Flora of Guatemala, by Paul C. Standley and J.A. Steyermark. 24 1946 Chicago, 1946- url p. 3, p. 4.
- Flora of Yucatan, by Paul C. Standley. 3 1930 Chicago, Field Museum of Natural History, 1930. url p. 242.
- Flora of the Aguan valley and the coastal regions near La Ceiba, Honduras, by T. G. Yuncker. 9 1940 [Chicago]1940. url p. 282.
- Gardens, wildlife densities, and subsistence hunting by Maya Indians in Quintana Roo, Mexico / 1993. url p. 278, p. 291, p. 299.
- Journal of ethnobiology. 24 2004 Flagstaff, Ariz.: Center for Western Studies, 1981- url p. 289, p. 295.
- National list of scientific plant names. [Washington, D.C.]: U.S. Dept. of Agriculture, Soil Conservation Service, 1982- url p. 267.
- Native woody plants of the United States, their erosion-control and wildlife values. Washington, U. S. Govt. print. off., 1938. url p. 171, p. 92, p. 93.
- Phytologia memoirs. Plainfield, N.J.: H.N. Moldenke and A.L. Moldenke, 1980- url p. 120.
- The Bradley bibliography; a guide to the literature of the woody plants of the world published before the beginning of the twentieth century; Cambridge, Riverside Press, 1911-18. url p. 141.
- The flora of Barro Colorado Island, Panama, by Paul C. Standley. City of Washington, The Smithsonian Institution, 1927. url p. 16.
- The forests and flora of British Honduras / by Paul C. Standley and Samuel J. Record; in cooperation with the Conservator of Forests and the Agricultural Officer of the Colony. 12 1936 Chicago: [Field Museum of Natural History], 1936. url p. 107.
- Torreya. Burlington, Vt., Torrey Botanical Club, 1901-1945. url p. 191.
- Trees and shrubs of Mexico / By Paul C. Standley. Washington, Govt. Print. Off., 1920-1926. url p. 200, p. 201.
- University of Kansas publications, Museum of Natural History. 13 1960 Lawrence, University of Kansas. url p. 32, plate 36, p. 614, p. 747.
- Wrightia. 7 1981-1984 Richardson, Tex. [etc.]University of Texas at Dallas. url p. 128, p. 200.
- Chang Siushih. 1998. Cannaboideae. In: Chang Siushih & Wu Chengyih, eds., Fl. Reipubl. Popularis Sin. 23(1): 220224.
- Correll, D. S. and M. C. Johnston. 1970. Manual of the Vascular Plants of Texas. Renner, Tex.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 10, 2012.
Data Sources
Accessed through GBIF Data Portal November 19, 2007:
- Botanical Research Institute of Texas, Andes to Amazon Biodiversity Program
- Herbarium of the University of Aarhus, The AAU Herbarium Database
- Herbier de la Guyane, Herbier de la Guyane
- Instituto Nacional de Biodiversidad
- , Biodiversidad de Costa Rica
- Missouri Botanical Garden, Missouri Botanical Garden
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2645771
- Catalogue of Life Accepted Name Code: ITS-19041
- Global Biodiversity Information Facility Taxonkey: 13736425
- Globally Unique Identifier: urn:lsid:ipni.org:names:1213898-2
- GRIN Nomen Number: 101687
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19041
- International Plant Names Index (IPNI) ID: 1213898-1
- Natural Heritage Network Species Identifier: PDULM01010
- U.S.D.A. Plant Symbol: CEIG
- Zipcode Zoo Species Identifier: 26691
Footnotes
- Zhengyi Wu, Zhe-Kun Zhou & Bruce Bartholomew "Cannabaceae". in Flora of China Vol. 5 Page 74. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Celtis". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- Mean = 440.550 meters (1,445.374 feet), Standard Deviation = 737.820 based on 987 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
